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Plymouth was colder, whereas Jamestown was warmer and more humid, and therefore more prone to harbor diseases. I'm sure that there were a lot more differences, but those are the big ones that leapt into my mind.
Main Difference -Town vs City. Both city and town refer to an urban area that has a name, defined boundaries and governance structure. The main difference between town and city is that a city is larger than a town. However, the definition of town and city varies considerably in different countries. What is a Town