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Plymouth was colder, whereas Jamestown was warmer and more humid, and therefore more prone to harbor diseases. I'm sure that there were a lot more differences, but those are the big ones that leapt into my mind.
Location and Transportation. Colonial Williamsburg is roughly 50 miles from Richmond and 150 miles from Washington, DC. As previously mentioned, Jamestown Settlement is only four miles from Williamsburg. The Colonial Williamsburg Foundation and the National Park Service cooperate to run a free Historic Triangle Shuttle from late spring to late autumn, with service between Colonial Williamsburg, Jamestown Island and Yorktown Battlefield.