Definition of GRAVAMEN: The burden or gist of a charge; the grievance or injury specially complained of.In English ecclesiastical law. A grievance complained of by the clergy befo The Law Dictionary Featuring Black's Law Dictionary Free Online Legal Dictionary 2nd Ed.
Phosphate and sugar make up the sides of a DNA ladder.
The steps, or rungs, of the DNA ladder are made up of two bases joined together with either two or three weak hydrogen bonds. The rails of the ladder are made up of alternating sugar and phosphate molecules. The rungs are comprised of two bases between sugar molecules.