Definition of GRAVAMEN: The burden or gist of a charge; the grievance or injury specially complained of.In English ecclesiastical law. A grievance complained of by the clergy befo The Law Dictionary Featuring Black's Law Dictionary Free Online Legal Dictionary 2nd Ed.
Phosphate and sugar make up the sides of a DNA ladder.
  The twisted ladder metaphorA DNA molecule is often likened to a spiral (or twisted) ladder. In this analogy, the rungs of the ladder are the base pairs (A-T and G-C). The sides of the ladder are the sugar-phosphate backbones of the two strands. This refers to B form DNA.wo hydrogen bonds form between adenine and thymine; three hydrogen bonds form between guanine and cytosine. There are three forms of DNA that differ significantly. The most common, B form, is the structure most people have heard of.