The Great Plains is the broad expanse of flat land (a plain), much of it covered in prairie, steppe and grassland, that lies west of the Mississippi River tallgrass prairie states and east of the Rocky Mountains in the United States and Canada. This area covers parts, but not all, of the states of Colorado, Kansas, Montana, Nebraska, New Mexico, North Dakota, Oklahoma, South Dakota, Texas, and Wyoming, and the Canadian provinces of Alberta, Manitoba and Saskatchewan. The region is known for supporting extensive cattle ranching and dry farming.


The North American Environmental Atlas, produced by the Commission for Environmental Cooperation, a NAFTA agency composed of the geographical agencies of the Mexican, American, and Canadian governments uses the "Great Plains" as an ecoregion synonymous with predominant prairies and grasslands rather than as physiographic region defined by topography. The Great Plains ecoregion includes five sub-regions: Temperate Prairies, West-Central Semi-Arid Prairies, South-Central Semi-Arid Prairies, Texas Louisiana Coastal Plains, and Tamaulipus-Texas Semi-Arid Plain, which overlap or expand upon other Great Plains designations.


The railroads opened up the Great Plains for settlement, for now it was possible to ship wheat and other crops at low cost to the urban markets in the East, and Europe. Homestead land was free for American settlers. Railroads sold their land at cheap rates to immigrants in expectation they would generate traffic as soon as farms were established. Immigrants poured in, especially from Germany and Scandinavia. On the plains, very few single men attempted to operate a farm or ranch by themselves; they clearly understood the need for a hard-working wife, and numerous children, to handle the many chores, including child-rearing, feeding and clothing the family, managing the housework, feeding the hired hands, and, especially after the 1930s, handling paperwork and financial details. During the early years of settlement, farm women played an integral role in assuring family survival by working outdoors. After approximately one generation, women increasingly left the fields, thus redefining their roles within the family. New technology including sewing and washing machines encouraged women to turn to domestic roles. The scientific housekeeping movement, promoted across the land by the media and government extension agents, as well as county fairs which featured achievements in home cookery and canning, advice columns for women regarding farm bookkeeping, and home economics courses in the schools.


During the Cenozoic era, specifically about 25 million years ago during the Miocene and Pliocene epochs, the continental climate became favorable to the evolution of grasslands. Existing forest biomes declined and grasslands became much more widespread. The grasslands provided a new niche for mammals, including many ungulates and glires, that switched from browsing diets to grazing diets. Traditionally, the spread of grasslands and the development of grazers have been strongly linked. However, an examination of mammalian teeth suggests that it is the open, gritty habitat and not the grass itself which is linked to diet changes in mammals, giving rise to the "grit, not grass" hypothesis.


To allow for agricultural development of the Great Plains and house a growing population, the US passed the Homestead Acts of 1862: it allowed a settler to claim up to 160 acres (65 ha) of land, provided that he lived on it for a period of five years and cultivated it. The provisions were expanded under the Kinkaid Act of 1904 to include a homestead of an entire section. Hundreds of thousands of people claimed such homesteads, sometimes building sod houses out of the very turf of their land. Many of them were not skilled dryland farmers and failures were frequent. Much of the Plains were settled during relatively wet years. Government experts did not understand how farmers should cultivate the prairies and gave advice counter to what would have worked[citation needed]. Germans from Russia who had previously farmed, under similar circumstances, in what is now Ukraine were marginally more successful than other homesteaders. The Dominion Lands Act of 1871 served a similar function for establishing homesteads on the prairies in Canada.


Infrared radiation is used in industrial, scientific, and medical applications. Night-vision devices using active near-infrared illumination allow people or animals to be observed without the observer being detected. Infrared astronomy uses sensor-equipped telescopes to penetrate dusty regions of space, such as molecular clouds; detect objects such as planets, and to view highly red-shifted objects from the early days of the universe. Infrared thermal-imaging cameras are used to detect heat loss in insulated systems, to observe changing blood flow in the skin, and to detect overheating of electrical apparatus.


The onset of infrared is defined (according to different standards) at various values typically between 700 nm and 800 nm, but the boundary between visible and infrared light is not precisely defined. The human eye is markedly less sensitive to light above 700 nm wavelength, so longer wavelengths make insignificant contributions to scenes illuminated by common light sources. However, particularly intense near-IR light (e.g., from IR lasers, IR LED sources, or from bright daylight with the visible light removed by colored gels) can be detected up to approximately 780 nm, and will be perceived as red light. Sources providing wavelengths as long as 1050 nm can be seen as a dull red glow in intense sources, causing some difficulty in near-IR illumination of scenes in the dark (usually this practical problem is solved by indirect illumination). Leaves are particularly bright in the near IR, and if all visible light leaks from around an IR-filter are blocked, and the eye is given a moment to adjust to the extremely dim image coming through a visually opaque IR-passing photographic filter, it is possible to see the Wood effect that consists of IR-glowing foliage.


The concept of emissivity is important in understanding the infrared emissions of objects. This is a property of a surface that describes how its thermal emissions deviate from the ideal of a black body. To further explain, two objects at the same physical temperature will not show the same infrared image if they have differing emissivity. For example, for any pre-set emissivity value, objects with higher emissivity will appear hotter, and those with a lower emissivity will appear cooler. For that reason, incorrect selection of emissivity will give inaccurate results when using infrared cameras and pyrometers.


Infrared vibrational spectroscopy (see also near-infrared spectroscopy) is a technique that can be used to identify molecules by analysis of their constituent bonds. Each chemical bond in a molecule vibrates at a frequency characteristic of that bond. A group of atoms in a molecule (e.g., CH2) may have multiple modes of oscillation caused by the stretching and bending motions of the group as a whole. If an oscillation leads to a change in dipole in the molecule then it will absorb a photon that has the same frequency. The vibrational frequencies of most molecules correspond to the frequencies of infrared light. Typically, the technique is used to study organic compounds using light radiation from 4000–400 cm−1, the mid-infrared. A spectrum of all the frequencies of absorption in a sample is recorded. This can be used to gain information about the sample composition in terms of chemical groups present and also its purity (for example, a wet sample will show a broad O-H absorption around 3200 cm−1).


In infrared photography, infrared filters are used to capture the near-infrared spectrum. Digital cameras often use infrared blockers. Cheaper digital cameras and camera phones have less effective filters and can "see" intense near-infrared, appearing as a bright purple-white color. This is especially pronounced when taking pictures of subjects near IR-bright areas (such as near a lamp), where the resulting infrared interference can wash out the image. There is also a technique called 'T-ray' imaging, which is imaging using far-infrared or terahertz radiation. Lack of bright sources can make terahertz photography more challenging than most other infrared imaging techniques. Recently T-ray imaging has been of considerable interest due to a number of new developments such as terahertz time-domain spectroscopy.


Infrared reflectography (fr; it; es), as called by art conservators, can be applied to paintings to reveal underlying layers in a completely non-destructive manner, in particular the underdrawing or outline drawn by the artist as a guide. This often reveals the artist's use of carbon black, which shows up well in reflectograms, as long as it has not also been used in the ground underlying the whole painting. Art conservators are looking to see whether the visible layers of paint differ from the underdrawing or layers in between – such alterations are called pentimenti when made by the original artist. This is very useful information in deciding whether a painting is the prime version by the original artist or a copy, and whether it has been altered by over-enthusiastic restoration work. In general, the more pentimenti the more likely a painting is to be the prime version. It also gives useful insights into working practices.


The discovery of infrared radiation is ascribed to William Herschel, the astronomer, in the early 19th century. Herschel published his results in 1800 before the Royal Society of London. Herschel used a prism to refract light from the sun and detected the infrared, beyond the red part of the spectrum, through an increase in the temperature recorded on a thermometer. He was surprised at the result and called them "Calorific Rays". The term 'Infrared' did not appear until late in the 19th century.


Infrared radiation is popularly known as "heat radiation"[citation needed], but light and electromagnetic waves of any frequency will heat surfaces that absorb them. Infrared light from the Sun accounts for 49% of the heating of Earth, with the rest being caused by visible light that is absorbed then re-radiated at longer wavelengths. Visible light or ultraviolet-emitting lasers can char paper and incandescently hot objects emit visible radiation. Objects at room temperature will emit radiation concentrated mostly in the 8 to 25 µm band, but this is not distinct from the emission of visible light by incandescent objects and ultraviolet by even hotter objects (see black body and Wien's displacement law).


Infrared tracking, also known as infrared homing, refers to a passive missile guidance system, which uses the emission from a target of electromagnetic radiation in the infrared part of the spectrum to track it. Missiles that use infrared seeking are often referred to as "heat-seekers", since infrared (IR) is just below the visible spectrum of light in frequency and is radiated strongly by hot bodies. Many objects such as people, vehicle engines, and aircraft generate and retain heat, and as such, are especially visible in the infrared wavelengths of light compared to objects in the background.


High, cold ice clouds such as Cirrus or Cumulonimbus show up bright white, lower warmer clouds such as Stratus or Stratocumulus show up as grey with intermediate clouds shaded accordingly. Hot land surfaces will show up as dark-grey or black. One disadvantage of infrared imagery is that low cloud such as stratus or fog can be a similar temperature to the surrounding land or sea surface and does not show up. However, using the difference in brightness of the IR4 channel (10.3–11.5 µm) and the near-infrared channel (1.58–1.64 µm), low cloud can be distinguished, producing a fog satellite picture. The main advantage of infrared is that images can be produced at night, allowing a continuous sequence of weather to be studied.


The sensitivity of Earth-based infrared telescopes is significantly limited by water vapor in the atmosphere, which absorbs a portion of the infrared radiation arriving from space outside of selected atmospheric windows. This limitation can be partially alleviated by placing the telescope observatory at a high altitude, or by carrying the telescope aloft with a balloon or an aircraft. Space telescopes do not suffer from this handicap, and so outer space is considered the ideal location for infrared astronomy.


Near-infrared is the region closest in wavelength to the radiation detectable by the human eye, mid- and far-infrared are progressively further from the visible spectrum. Other definitions follow different physical mechanisms (emission peaks, vs. bands, water absorption) and the newest follow technical reasons (the common silicon detectors are sensitive to about 1,050 nm, while InGaAs's sensitivity starts around 950 nm and ends between 1,700 and 2,600 nm, depending on the specific configuration). Unfortunately, international standards for these specifications are not currently available.


Heat is energy in transit that flows due to temperature difference. Unlike heat transmitted by thermal conduction or thermal convection, thermal radiation can propagate through a vacuum. Thermal radiation is characterized by a particular spectrum of many wavelengths that is associated with emission from an object, due to the vibration of its molecules at a given temperature. Thermal radiation can be emitted from objects at any wavelength, and at very high temperatures such radiations are associated with spectra far above the infrared, extending into visible, ultraviolet, and even X-ray regions (i.e., the solar corona). Thus, the popular association of infrared radiation with thermal radiation is only a coincidence based on typical (comparatively low) temperatures often found near the surface of planet Earth.


Thermographic cameras detect radiation in the infrared range of the electromagnetic spectrum (roughly 900–14,000 nanometers or 0.9–14 μm) and produce images of that radiation. Since infrared radiation is emitted by all objects based on their temperatures, according to the black body radiation law, thermography makes it possible to "see" one's environment with or without visible illumination. The amount of radiation emitted by an object increases with temperature, therefore thermography allows one to see variations in temperature (hence the name).


The infrared portion of the spectrum has several useful benefits for astronomers. Cold, dark molecular clouds of gas and dust in our galaxy will glow with radiated heat as they are irradiated by imbedded stars. Infrared can also be used to detect protostars before they begin to emit visible light. Stars emit a smaller portion of their energy in the infrared spectrum, so nearby cool objects such as planets can be more readily detected. (In the visible light spectrum, the glare from the star will drown out the reflected light from a planet.)


Infrared is used in night vision equipment when there is insufficient visible light to see. Night vision devices operate through a process involving the conversion of ambient light photons into electrons that are then amplified by a chemical and electrical process and then converted back into visible light. Infrared light sources can be used to augment the available ambient light for conversion by night vision devices, increasing in-the-dark visibility without actually using a visible light source.


IR data transmission is also employed in short-range communication among computer peripherals and personal digital assistants. These devices usually conform to standards published by IrDA, the Infrared Data Association. Remote controls and IrDA devices use infrared light-emitting diodes (LEDs) to emit infrared radiation that is focused by a plastic lens into a narrow beam. The beam is modulated, i.e. switched on and off, to encode the data. The receiver uses a silicon photodiode to convert the infrared radiation to an electric current. It responds only to the rapidly pulsing signal created by the transmitter, and filters out slowly changing infrared radiation from ambient light. Infrared communications are useful for indoor use in areas of high population density. IR does not penetrate walls and so does not interfere with other devices in adjoining rooms. Infrared is the most common way for remote controls to command appliances. Infrared remote control protocols like RC-5, SIRC, are used to communicate with infrared.


In the semiconductor industry, infrared light can be used to characterize materials such as thin films and periodic trench structures. By measuring the reflectance of light from the surface of a semiconductor wafer, the index of refraction (n) and the extinction Coefficient (k) can be determined via the Forouhi-Bloomer dispersion equations. The reflectance from the infrared light can also be used to determine the critical dimension, depth, and sidewall angle of high aspect ratio trench structures.


Infrared cleaning is a technique used by some Motion picture film scanner, film scanners and flatbed scanners to reduce or remove the effect of dust and scratches upon the finished scan. It works by collecting an additional infrared channel from the scan at the same position and resolution as the three visible color channels (red, green, and blue). The infrared channel, in combination with the other channels, is used to detect the location of scratches and dust. Once located, those defects can be corrected by scaling or replaced by inpainting.


Earth's surface and the clouds absorb visible and invisible radiation from the sun and re-emit much of the energy as infrared back to atmosphere. Certain substances in the atmosphere, chiefly cloud droplets and water vapor, but also carbon dioxide, methane, nitrous oxide, sulfur hexafluoride, and chlorofluorocarbons, absorb this infrared, and re-radiate it in all directions including back to Earth. Thus, the greenhouse effect keeps the atmosphere and surface much warmer than if the infrared absorbers were absent from the atmosphere.


Biodiversity, a contraction of "biological diversity," generally refers to the variety and variability of life on Earth. One of the most widely used definitions defines it in terms of the variability within species, between species, and between ecosystems. It is a measure of the variety of organisms present in different ecosystems. This can refer to genetic variation, ecosystem variation, or species variation (number of species) within an area, biome, or planet. Terrestrial biodiversity tends to be greater near the equator, which seems to be the result of the warm climate and high primary productivity. Biodiversity is not distributed evenly on Earth. It is richest in the tropics. Marine biodiversity tends to be highest along coasts in the Western Pacific, where sea surface temperature is highest and in the mid-latitudinal band in all oceans. There are latitudinal gradients in species diversity. Biodiversity generally tends to cluster in hotspots, and has been increasing through time, but will be likely to slow in the future.


This multilevel construct is consistent with Dasmann and Lovejoy. An explicit definition consistent with this interpretation was first given in a paper by Bruce A. Wilcox commissioned by the International Union for the Conservation of Nature and Natural Resources (IUCN) for the 1982 World National Parks Conference. Wilcox's definition was "Biological diversity is the variety of life forms...at all levels of biological systems (i.e., molecular, organismic, population, species and ecosystem)...". The 1992 United Nations Earth Summit defined "biological diversity" as "the variability among living organisms from all sources, including, 'inter alia', terrestrial, marine, and other aquatic ecosystems, and the ecological complexes of which they are part: this includes diversity within species, between species and of ecosystems". This definition is used in the United Nations Convention on Biological Diversity.


On the other hand, changes through the Phanerozoic correlate much better with the hyperbolic model (widely used in population biology, demography and macrosociology, as well as fossil biodiversity) than with exponential and logistic models. The latter models imply that changes in diversity are guided by a first-order positive feedback (more ancestors, more descendants) and/or a negative feedback arising from resource limitation. Hyperbolic model implies a second-order positive feedback. The hyperbolic pattern of the world population growth arises from a second-order positive feedback between the population size and the rate of technological growth. The hyperbolic character of biodiversity growth can be similarly accounted for by a feedback between diversity and community structure complexity. The similarity between the curves of biodiversity and human population probably comes from the fact that both are derived from the interference of the hyperbolic trend with cyclical and stochastic dynamics.


Interspecific crop diversity is, in part, responsible for offering variety in what we eat. Intraspecific diversity, the variety of alleles within a single species, also offers us choice in our diets. If a crop fails in a monoculture, we rely on agricultural diversity to replant the land with something new. If a wheat crop is destroyed by a pest we may plant a hardier variety of wheat the next year, relying on intraspecific diversity. We may forgo wheat production in that area and plant a different species altogether, relying on interspecific diversity. Even an agricultural society which primarily grows monocultures, relies on biodiversity at some point.


In absolute terms, the planet has lost 52% of its biodiversity since 1970 according to a 2014 study by the World Wildlife Fund. The Living Planet Report 2014 claims that "the number of mammals, birds, reptiles, amphibians and fish across the globe is, on average, about half the size it was 40 years ago". Of that number, 39% accounts for the terrestrial wildlife gone, 39% for the marine wildlife gone, and 76% for the freshwater wildlife gone. Biodiversity took the biggest hit in Latin America, plummeting 83 percent. High-income countries showed a 10% increase in biodiversity, which was canceled out by a loss in low-income countries. This is despite the fact that high-income countries use five times the ecological resources of low-income countries, which was explained as a result of process whereby wealthy nations are outsourcing resource depletion to poorer nations, which are suffering the greatest ecosystem losses.


A 2007 study conducted by the National Science Foundation found that biodiversity and genetic diversity are codependent—that diversity among species requires diversity within a species, and vice versa. "If any one type is removed from the system, the cycle can break down, and the community becomes dominated by a single species." At present, the most threatened ecosystems are found in fresh water, according to the Millennium Ecosystem Assessment 2005, which was confirmed by the "Freshwater Animal Diversity Assessment", organised by the biodiversity platform, and the French Institut de recherche pour le développement (MNHNP).


Finally, an introduced species may unintentionally injure a species that depends on the species it replaces. In Belgium, Prunus spinosa from Eastern Europe leafs much sooner than its West European counterparts, disrupting the feeding habits of the Thecla betulae butterfly (which feeds on the leaves). Introducing new species often leaves endemic and other local species unable to compete with the exotic species and unable to survive. The exotic organisms may be predators, parasites, or may simply outcompete indigenous species for nutrients, water and light.


The forests play a vital role in harbouring more than 45,000 floral and 81,000 faunal species of which 5150 floral and 1837 faunal species are endemic. Plant and animal species confined to a specific geographical area are called endemic species. In reserved forests, rights to activities like hunting and grazing are sometimes given to communities living on the fringes of the forest, who sustain their livelihood partially or wholly from forest resources or products. The unclassed forests covers 6.4 percent of the total forest area and they are marked by the following characteristics:


Global agreements such as the Convention on Biological Diversity, give "sovereign national rights over biological resources" (not property). The agreements commit countries to "conserve biodiversity", "develop resources for sustainability" and "share the benefits" resulting from their use. Biodiverse countries that allow bioprospecting or collection of natural products, expect a share of the benefits rather than allowing the individual or institution that discovers/exploits the resource to capture them privately. Bioprospecting can become a type of biopiracy when such principles are not respected.[citation needed]


Rapid environmental changes typically cause mass extinctions. More than 99 percent of all species, amounting to over five billion species, that ever lived on Earth are estimated to be extinct. Estimates on the number of Earth's current species range from 10 million to 14 million, of which about 1.2 million have been documented and over 86 percent have not yet been described. The total amount of related DNA base pairs on Earth is estimated at 5.0 x 1037, and weighs 50 billion tonnes. In comparison, the total mass of the biosphere has been estimated to be as much as 4 TtC (trillion tons of carbon).


The history of biodiversity during the Phanerozoic (the last 540 million years), starts with rapid growth during the Cambrian explosion—a period during which nearly every phylum of multicellular organisms first appeared. Over the next 400 million years or so, invertebrate diversity showed little overall trend, and vertebrate diversity shows an overall exponential trend. This dramatic rise in diversity was marked by periodic, massive losses of diversity classified as mass extinction events. A significant loss occurred when rainforests collapsed in the carboniferous. The worst was the Permian-Triassic extinction event, 251 million years ago. Vertebrates took 30 million years to recover from this event.


Jared Diamond describes an "Evil Quartet" of habitat destruction, overkill, introduced species, and secondary extinctions. Edward O. Wilson prefers the acronym HIPPO, standing for Habitat destruction, Invasive species, Pollution, human over-Population, and Over-harvesting. The most authoritative classification in use today is IUCN's Classification of Direct Threats which has been adopted by major international conservation organizations such as the US Nature Conservancy, the World Wildlife Fund, Conservation International, and BirdLife International.


Endemic species can be threatened with extinction through the process of genetic pollution, i.e. uncontrolled hybridization, introgression and genetic swamping. Genetic pollution leads to homogenization or replacement of local genomes as a result of either a numerical and/or fitness advantage of an introduced species. Hybridization and introgression are side-effects of introduction and invasion. These phenomena can be especially detrimental to rare species that come into contact with more abundant ones. The abundant species can interbreed with the rare species, swamping its gene pool. This problem is not always apparent from morphological (outward appearance) observations alone. Some degree of gene flow is normal adaptation, and not all gene and genotype constellations can be preserved. However, hybridization with or without introgression may, nevertheless, threaten a rare species' existence.


The age of the Earth is about 4.54 billion years old. The earliest undisputed evidence of life on Earth dates at least from 3.5 billion years ago, during the Eoarchean Era after a geological crust started to solidify following the earlier molten Hadean Eon. There are microbial mat fossils found in 3.48 billion-year-old sandstone discovered in Western Australia. Other early physical evidence of a biogenic substance is graphite in 3.7 billion-year-old metasedimentary rocks discovered in Western Greenland. More recently, in 2015, "remains of biotic life" were found in 4.1 billion-year-old rocks in Western Australia. According to one of the researchers, "If life arose relatively quickly on Earth ... then it could be common in the universe."


The fossil record suggests that the last few million years featured the greatest biodiversity in history. However, not all scientists support this view, since there is uncertainty as to how strongly the fossil record is biased by the greater availability and preservation of recent geologic sections. Some scientists believe that corrected for sampling artifacts, modern biodiversity may not be much different from biodiversity 300 million years ago., whereas others consider the fossil record reasonably reflective of the diversification of life. Estimates of the present global macroscopic species diversity vary from 2 million to 100 million, with a best estimate of somewhere near 9 million, the vast majority arthropods. Diversity appears to increase continually in the absence of natural selection.


Agricultural diversity can also be divided by whether it is ‘planned’ diversity or ‘associated’ diversity. This is a functional classification that we impose and not an intrinsic feature of life or diversity. Planned diversity includes the crops which a farmer has encouraged, planted or raised (e.g.: crops, covers, symbionts and livestock, among others), which can be contrasted with the associated diversity that arrives among the crops, uninvited (e.g.: herbivores, weed species and pathogens, among others).


Biodiversity's relevance to human health is becoming an international political issue, as scientific evidence builds on the global health implications of biodiversity loss. This issue is closely linked with the issue of climate change, as many of the anticipated health risks of climate change are associated with changes in biodiversity (e.g. changes in populations and distribution of disease vectors, scarcity of fresh water, impacts on agricultural biodiversity and food resources etc.) This is because the species most likely to disappear are those that buffer against infectious disease transmission, while surviving species tend to be the ones that increase disease transmission, such as that of West Nile Virus, Lyme disease and Hantavirus, according to a study done co-authored by Felicia Keesing, an ecologist at Bard College, and Drew Harvell, associate director for Environment of the Atkinson Center for a Sustainable Future (ACSF) at Cornell University.


Since life began on Earth, five major mass extinctions and several minor events have led to large and sudden drops in biodiversity. The Phanerozoic eon (the last 540 million years) marked a rapid growth in biodiversity via the Cambrian explosion—a period during which the majority of multicellular phyla first appeared. The next 400 million years included repeated, massive biodiversity losses classified as mass extinction events. In the Carboniferous, rainforest collapse led to a great loss of plant and animal life. The Permian–Triassic extinction event, 251 million years ago, was the worst; vertebrate recovery took 30 million years. The most recent, the Cretaceous–Paleogene extinction event, occurred 65 million years ago and has often attracted more attention than others because it resulted in the extinction of the dinosaurs.


The number of species invasions has been on the rise at least since the beginning of the 1900s. Species are increasingly being moved by humans (on purpose and accidentally). In some cases the invaders are causing drastic changes and damage to their new habitats (e.g.: zebra mussels and the emerald ash borer in the Great Lakes region and the lion fish along the North American Atlantic coast). Some evidence suggests that invasive species are competitive in their new habitats because they are subject to less pathogen disturbance. Others report confounding evidence that occasionally suggest that species-rich communities harbor many native and exotic species simultaneously while some say that diverse ecosystems are more resilient and resist invasive plants and animals. An important question is, "do invasive species cause extinctions?" Many studies cite effects of invasive species on natives, but not extinctions. Invasive species seem to increase local (i.e.: alpha diversity) diversity, which decreases turnover of diversity (i.e.: beta diversity). Overall gamma diversity may be lowered because species are going extinct because of other causes, but even some of the most insidious invaders (e.g.: Dutch elm disease, emerald ash borer, chestnut blight in North America) have not caused their host species to become extinct. Extirpation, population decline, and homogenization of regional biodiversity are much more common. Human activities have frequently been the cause of invasive species circumventing their barriers, by introducing them for food and other purposes. Human activities therefore allow species to migrate to new areas (and thus become invasive) occurred on time scales much shorter than historically have been required for a species to extend its range.


Brazil's Atlantic Forest is considered one such hotspot, containing roughly 20,000 plant species, 1,350 vertebrates, and millions of insects, about half of which occur nowhere else.[citation needed] The island of Madagascar and India are also particularly notable. Colombia is characterized by high biodiversity, with the highest rate of species by area unit worldwide and it has the largest number of endemics (species that are not found naturally anywhere else) of any country. About 10% of the species of the Earth can be found in Colombia, including over 1,900 species of bird, more than in Europe and North America combined, Colombia has 10% of the world’s mammals species, 14% of the amphibian species, and 18% of the bird species of the world. Madagascar dry deciduous forests and lowland rainforests possess a high ratio of endemism.[citation needed] Since the island separated from mainland Africa 66 million years ago, many species and ecosystems have evolved independently.[citation needed] Indonesia's 17,000 islands cover 735,355 square miles (1,904,560 km2) and contain 10% of the world's flowering plants, 12% of mammals, and 17% of reptiles, amphibians and birds—along with nearly 240 million people. Many regions of high biodiversity and/or endemism arise from specialized habitats which require unusual adaptations, for example, alpine environments in high mountains, or Northern European peat bogs.[citation needed]


The existence of a "global carrying capacity", limiting the amount of life that can live at once, is debated, as is the question of whether such a limit would also cap the number of species. While records of life in the sea shows a logistic pattern of growth, life on land (insects, plants and tetrapods)shows an exponential rise in diversity. As one author states, "Tetrapods have not yet invaded 64 per cent of potentially habitable modes, and it could be that without human influence the ecological and taxonomic diversity of tetrapods would continue to increase in an exponential fashion until most or all of the available ecospace is filled."


From 1950 to 2011, world population increased from 2.5 billion to 7 billion and is forecast to reach a plateau of more than 9 billion during the 21st century. Sir David King, former chief scientific adviser to the UK government, told a parliamentary inquiry: "It is self-evident that the massive growth in the human population through the 20th century has had more impact on biodiversity than any other single factor." At least until the middle of the 21st century, worldwide losses of pristine biodiverse land will probably depend much on the worldwide human birth rate.


The control of associated biodiversity is one of the great agricultural challenges that farmers face. On monoculture farms, the approach is generally to eradicate associated diversity using a suite of biologically destructive pesticides, mechanized tools and transgenic engineering techniques, then to rotate crops. Although some polyculture farmers use the same techniques, they also employ integrated pest management strategies as well as strategies that are more labor-intensive, but generally less dependent on capital, biotechnology and energy.


National park and nature reserve is the area selected by governments or private organizations for special protection against damage or degradation with the objective of biodiversity and landscape conservation. National parks are usually owned and managed by national or state governments. A limit is placed on the number of visitors permitted to enter certain fragile areas. Designated trails or roads are created. The visitors are allowed to enter only for study, cultural and recreation purposes. Forestry operations, grazing of animals and hunting of animals are prohibited. Exploitation of habitat or wildlife is banned.


During the last century, decreases in biodiversity have been increasingly observed. In 2007, German Federal Environment Minister Sigmar Gabriel cited estimates that up to 30% of all species will be extinct by 2050. Of these, about one eighth of known plant species are threatened with extinction. Estimates reach as high as 140,000 species per year (based on Species-area theory). This figure indicates unsustainable ecological practices, because few species emerge each year.[citation needed] Almost all scientists acknowledge that the rate of species loss is greater now than at any time in human history, with extinctions occurring at rates hundreds of times higher than background extinction rates. As of 2012, some studies suggest that 25% of all mammal species could be extinct in 20 years.


Habitat size and numbers of species are systematically related. Physically larger species and those living at lower latitudes or in forests or oceans are more sensitive to reduction in habitat area. Conversion to "trivial" standardized ecosystems (e.g., monoculture following deforestation) effectively destroys habitat for the more diverse species that preceded the conversion. In some countries lack of property rights or lax law/regulatory enforcement necessarily leads to biodiversity loss (degradation costs having to be supported by the community).[citation needed]


Not all introduced species are invasive, nor all invasive species deliberately introduced. In cases such as the zebra mussel, invasion of US waterways was unintentional. In other cases, such as mongooses in Hawaii, the introduction is deliberate but ineffective (nocturnal rats were not vulnerable to the diurnal mongoose). In other cases, such as oil palms in Indonesia and Malaysia, the introduction produces substantial economic benefits, but the benefits are accompanied by costly unintended consequences.


Less than 1% of all species that have been described have been studied beyond simply noting their existence. The vast majority of Earth's species are microbial. Contemporary biodiversity physics is "firmly fixated on the visible [macroscopic] world". For example, microbial life is metabolically and environmentally more diverse than multicellular life (see e.g., extremophile). "On the tree of life, based on analyses of small-subunit ribosomal RNA, visible life consists of barely noticeable twigs. The inverse relationship of size and population recurs higher on the evolutionary ladder—"to a first approximation, all multicellular species on Earth are insects". Insect extinction rates are high—supporting the Holocene extinction hypothesis.


The number and variety of plants, animals and other organisms that exist is known as biodiversity. It is an essential component of nature and it ensures the survival of human species by providing food, fuel, shelter, medicines and other resources to mankind. The richness of biodiversity depends on the climatic conditions and area of the region. All species of plants taken together are known as flora and about 70,000 species of plants are known till date. All species of animals taken together are known as fauna which includes birds, mammals, fish, reptiles, insects, crustaceans, molluscs, etc.


The term biological diversity was used first by wildlife scientist and conservationist Raymond F. Dasmann in the year 1968 lay book A Different Kind of Country advocating conservation. The term was widely adopted only after more than a decade, when in the 1980s it came into common usage in science and environmental policy. Thomas Lovejoy, in the foreword to the book Conservation Biology, introduced the term to the scientific community. Until then the term "natural diversity" was common, introduced by The Science Division of The Nature Conservancy in an important 1975 study, "The Preservation of Natural Diversity." By the early 1980s TNC's Science program and its head, Robert E. Jenkins, Lovejoy and other leading conservation scientists at the time in America advocated the use of the term "biological diversity".


Biodiversity provides critical support for drug discovery and the availability of medicinal resources. A significant proportion of drugs are derived, directly or indirectly, from biological sources: at least 50% of the pharmaceutical compounds on the US market are derived from plants, animals, and micro-organisms, while about 80% of the world population depends on medicines from nature (used in either modern or traditional medical practice) for primary healthcare. Only a tiny fraction of wild species has been investigated for medical potential. Biodiversity has been critical to advances throughout the field of bionics. Evidence from market analysis and biodiversity science indicates that the decline in output from the pharmaceutical sector since the mid-1980s can be attributed to a move away from natural product exploration ("bioprospecting") in favor of genomics and synthetic chemistry, indeed claims about the value of undiscovered pharmaceuticals may not provide enough incentive for companies in free markets to search for them because of the high cost of development; meanwhile, natural products have a long history of supporting significant economic and health innovation. Marine ecosystems are particularly important, although inappropriate bioprospecting can increase biodiversity loss, as well as violating the laws of the communities and states from which the resources are taken.


In agriculture and animal husbandry, the Green Revolution popularized the use of conventional hybridization to increase yield. Often hybridized breeds originated in developed countries and were further hybridized with local varieties in the developing world to create high yield strains resistant to local climate and diseases. Local governments and industry have been pushing hybridization. Formerly huge gene pools of various wild and indigenous breeds have collapsed causing widespread genetic erosion and genetic pollution. This has resulted in loss of genetic diversity and biodiversity as a whole.


Originally based on the English alphabet, ASCII encodes 128 specified characters into seven-bit integers as shown by the ASCII chart on the right. The characters encoded are numbers 0 to 9, lowercase letters a to z, uppercase letters A to Z, basic punctuation symbols, control codes that originated with Teletype machines, and a space. For example, lowercase j would become binary 1101010 and decimal 106. ASCII includes definitions for 128 characters: 33 are non-printing control characters (many now obsolete) that affect how text and space are processed and 95 printable characters, including the space (which is considered an invisible graphic:223).


The code itself was patterned so that most control codes were together, and all graphic codes were together, for ease of identification. The first two columns (32 positions) were reserved for control characters.:220, 236 § 8,9) The "space" character had to come before graphics to make sorting easier, so it became position 20hex;:237 § 10 for the same reason, many special signs commonly used as separators were placed before digits. The committee decided it was important to support uppercase 64-character alphabets, and chose to pattern ASCII so it could be reduced easily to a usable 64-character set of graphic codes,:228, 237 § 14 as was done in the DEC SIXBIT code. Lowercase letters were therefore not interleaved with uppercase. To keep options available for lowercase letters and other graphics, the special and numeric codes were arranged before the letters, and the letter A was placed in position 41hex to match the draft of the corresponding British standard.:238 § 18 The digits 0–9 were arranged so they correspond to values in binary prefixed with 011, making conversion with binary-coded decimal straightforward.


ASCII was incorporated into the Unicode character set as the first 128 symbols, so the 7-bit ASCII characters have the same numeric codes in both sets. This allows UTF-8 to be backward compatible with 7-bit ASCII, as a UTF-8 file containing only ASCII characters is identical to an ASCII file containing the same sequence of characters. Even more importantly, forward compatibility is ensured as software that recognizes only 7-bit ASCII characters as special and does not alter bytes with the highest bit set (as is often done to support 8-bit ASCII extensions such as ISO-8859-1) will preserve UTF-8 data unchanged.


When a Teletype 33 ASR equipped with the automatic paper tape reader received a Control-S (XOFF, an abbreviation for transmit off), it caused the tape reader to stop; receiving Control-Q (XON, "transmit on") caused the tape reader to resume. This technique became adopted by several early computer operating systems as a "handshaking" signal warning a sender to stop transmission because of impending overflow; it persists to this day in many systems as a manual output control technique. On some systems Control-S retains its meaning but Control-Q is replaced by a second Control-S to resume output. The 33 ASR also could be configured to employ Control-R (DC2) and Control-T (DC4) to start and stop the tape punch; on some units equipped with this function, the corresponding control character lettering on the keycap above the letter was TAPE and TAPE respectively.


DEC operating systems (OS/8, RT-11, RSX-11, RSTS, TOPS-10, etc.) used both characters to mark the end of a line so that the console device (originally Teletype machines) would work. By the time so-called "glass TTYs" (later called CRTs or terminals) came along, the convention was so well established that backward compatibility necessitated continuing the convention. When Gary Kildall cloned RT-11 to create CP/M he followed established DEC convention. Until the introduction of PC DOS in 1981, IBM had no hand in this because their 1970s operating systems used EBCDIC instead of ASCII and they were oriented toward punch-card input and line printer output on which the concept of carriage return was meaningless. IBM's PC DOS (also marketed as MS-DOS by Microsoft) inherited the convention by virtue of being a clone of CP/M, and Windows inherited it from MS-DOS.


C trigraphs were created to solve this problem for ANSI C, although their late introduction and inconsistent implementation in compilers limited their use. Many programmers kept their computers on US-ASCII, so plain-text in Swedish, German etc. (for example, in e-mail or Usenet) contained "{, }" and similar variants in the middle of words, something those programmers got used to. For example, a Swedish programmer mailing another programmer asking if they should go for lunch, could get "N{ jag har sm|rg}sar." as the answer, which should be "Nä jag har smörgåsar." meaning "No I've got sandwiches."


The X3.2 subcommittee designed ASCII based on the earlier teleprinter encoding systems. Like other character encodings, ASCII specifies a correspondence between digital bit patterns and character symbols (i.e. graphemes and control characters). This allows digital devices to communicate with each other and to process, store, and communicate character-oriented information such as written language. Before ASCII was developed, the encodings in use included 26 alphabetic characters, 10 numerical digits, and from 11 to 25 special graphic symbols. To include all these, and control characters compatible with the Comité Consultatif International Téléphonique et Télégraphique (CCITT) International Telegraph Alphabet No. 2 (ITA2) standard, Fieldata, and early EBCDIC, more than 64 codes were required for ASCII.


ASCII itself was first used commercially during 1963 as a seven-bit teleprinter code for American Telephone & Telegraph's TWX (TeletypeWriter eXchange) network. TWX originally used the earlier five-bit ITA2, which was also used by the competing Telex teleprinter system. Bob Bemer introduced features such as the escape sequence. His British colleague Hugh McGregor Ross helped to popularize this work –  according to Bemer, "so much so that the code that was to become ASCII was first called the Bemer-Ross Code in Europe". Because of his extensive work on ASCII, Bemer has been called "the father of ASCII."


For example, character 10 represents the "line feed" function (which causes a printer to advance its paper), and character 8 represents "backspace". RFC 2822 refers to control characters that do not include carriage return, line feed or white space as non-whitespace control characters. Except for the control characters that prescribe elementary line-oriented formatting, ASCII does not define any mechanism for describing the structure or appearance of text within a document. Other schemes, such as markup languages, address page and document layout and formatting.


Some software assigned special meanings to ASCII characters sent to the software from the terminal. Operating systems from Digital Equipment Corporation, for example, interpreted DEL as an input character as meaning "remove previously-typed input character", and this interpretation also became common in Unix systems. Most other systems used BS for that meaning and used DEL to mean "remove the character at the cursor".[citation needed] That latter interpretation is the most common now.[citation needed]


Computers attached to the ARPANET included machines running operating systems such as TOPS-10 and TENEX using CR-LF line endings, machines running operating systems such as Multics using LF line endings, and machines running operating systems such as OS/360 that represented lines as a character count followed by the characters of the line and that used EBCDIC rather than ASCII. The Telnet protocol defined an ASCII "Network Virtual Terminal" (NVT), so that connections between hosts with different line-ending conventions and character sets could be supported by transmitting a standard text format over the network. Telnet used ASCII along with CR-LF line endings, and software using other conventions would translate between the local conventions and the NVT. The File Transfer Protocol adopted the Telnet protocol, including use of the Network Virtual Terminal, for use when transmitting commands and transferring data in the default ASCII mode. This adds complexity to implementations of those protocols, and to other network protocols, such as those used for E-mail and the World Wide Web, on systems not using the NVT's CR-LF line-ending convention.


From early in its development, ASCII was intended to be just one of several national variants of an international character code standard, ultimately published as ISO/IEC 646 (1972), which would share most characters in common but assign other locally useful characters to several code points reserved for "national use." However, the four years that elapsed between the publication of ASCII-1963 and ISO's first acceptance of an international recommendation during 1967 caused ASCII's choices for the national use characters to seem to be de facto standards for the world, causing confusion and incompatibility once other countries did begin to make their own assignments to these code points.


Most early home computer systems developed their own 8-bit character sets containing line-drawing and game glyphs, and often filled in some or all of the control characters from 0–31 with more graphics. Kaypro CP/M computers used the "upper" 128 characters for the Greek alphabet. The IBM PC defined code page 437, which replaced the control-characters with graphic symbols such as smiley faces, and mapped additional graphic characters to the upper 128 positions. Operating systems such as DOS supported these code pages, and manufacturers of IBM PCs supported them in hardware. Digital Equipment Corporation developed the Multinational Character Set (DEC-MCS) for use in the popular VT220 terminal as one of the first extensions designed more for international languages than for block graphics. The Macintosh defined Mac OS Roman and Postscript also defined a set, both of these contained both international letters and typographic punctuation marks instead of graphics, more like modern character sets.


ASCII (i/ˈæski/ ASS-kee), abbreviated from American Standard Code for Information Interchange, is a character-encoding scheme (the IANA prefers the name US-ASCII). ASCII codes represent text in computers, communications equipment, and other devices that use text. Most modern character-encoding schemes are based on ASCII, though they support many additional characters. ASCII was the most common character encoding on the World Wide Web until December 2007, when it was surpassed by UTF-8, which is fully backward compatibe to ASCII.


The committee debated the possibility of a shift function (like in ITA2), which would allow more than 64 codes to be represented by a six-bit code. In a shifted code, some character codes determine choices between options for the following character codes. It allows compact encoding, but is less reliable for data transmission as an error in transmitting the shift code typically makes a long part of the transmission unreadable. The standards committee decided against shifting, and so ASCII required at least a seven-bit code.:215, 236 § 4


Many more of the control codes have been given meanings quite different from their original ones. The "escape" character (ESC, code 27), for example, was intended originally to allow sending other control characters as literals instead of invoking their meaning. This is the same meaning of "escape" encountered in URL encodings, C language strings, and other systems where certain characters have a reserved meaning. Over time this meaning has been co-opted and has eventually been changed. In modern use, an ESC sent to the terminal usually indicates the start of a command sequence, usually in the form of a so-called "ANSI escape code" (or, more properly, a "Control Sequence Introducer") beginning with ESC followed by a "[" (left-bracket) character. An ESC sent from the terminal is most often used as an out-of-band character used to terminate an operation, as in the TECO and vi text editors. In graphical user interface (GUI) and windowing systems, ESC generally causes an application to abort its current operation or to exit (terminate) altogether.


Older operating systems such as TOPS-10, along with CP/M, tracked file length only in units of disk blocks and used Control-Z (SUB) to mark the end of the actual text in the file. For this reason, EOF, or end-of-file, was used colloquially and conventionally as a three-letter acronym for Control-Z instead of SUBstitute. The end-of-text code (ETX), also known as Control-C, was inappropriate for a variety of reasons, while using Z as the control code to end a file is analogous to it ending the alphabet and serves as a very convenient mnemonic aid. A historically common and still prevalent convention uses the ETX code convention to interrupt and halt a program via an input data stream, usually from a keyboard.


ASCII developed from telegraphic codes. Its first commercial use was as a seven-bit teleprinter code promoted by Bell data services. Work on the ASCII standard began on October 6, 1960, with the first meeting of the American Standards Association's (ASA) X3.2 subcommittee. The first edition of the standard was published during 1963, underwent a major revision during 1967, and experienced its most recent update during 1986. Compared to earlier telegraph codes, the proposed Bell code and ASCII were both ordered for more convenient sorting (i.e., alphabetization) of lists, and added features for devices other than teleprinters.


The committee considered an eight-bit code, since eight bits (octets) would allow two four-bit patterns to efficiently encode two digits with binary-coded decimal. However, it would require all data transmission to send eight bits when seven could suffice. The committee voted to use a seven-bit code to minimize costs associated with data transmission. Since perforated tape at the time could record eight bits in one position, it also allowed for a parity bit for error checking if desired.:217, 236 § 5 Eight-bit machines (with octets as the native data type) that did not use parity checking typically set the eighth bit to 0.


With the other special characters and control codes filled in, ASCII was published as ASA X3.4-1963, leaving 28 code positions without any assigned meaning, reserved for future standardization, and one unassigned control code.:66, 245 There was some debate at the time whether there should be more control characters rather than the lowercase alphabet.:435 The indecision did not last long: during May 1963 the CCITT Working Party on the New Telegraph Alphabet proposed to assign lowercase characters to columns 6 and 7, and International Organization for Standardization TC 97 SC 2 voted during October to incorporate the change into its draft standard. The X3.2.4 task group voted its approval for the change to ASCII at its May 1963 meeting. Locating the lowercase letters in columns 6 and 7 caused the characters to differ in bit pattern from the upper case by a single bit, which simplified case-insensitive character matching and the construction of keyboards and printers.


Other international standards bodies have ratified character encodings such as ISO/IEC 646 that are identical or nearly identical to ASCII, with extensions for characters outside the English alphabet and symbols used outside the United States, such as the symbol for the United Kingdom's pound sterling (£). Almost every country needed an adapted version of ASCII, since ASCII suited the needs of only the USA and a few other countries. For example, Canada had its own version that supported French characters. Other adapted encodings include ISCII (India), VISCII (Vietnam), and YUSCII (Yugoslavia). Although these encodings are sometimes referred to as ASCII, true ASCII is defined strictly only by the ANSI standard.


Probably the most influential single device on the interpretation of these characters was the Teletype Model 33 ASR, which was a printing terminal with an available paper tape reader/punch option. Paper tape was a very popular medium for long-term program storage until the 1980s, less costly and in some ways less fragile than magnetic tape. In particular, the Teletype Model 33 machine assignments for codes 17 (Control-Q, DC1, also known as XON), 19 (Control-S, DC3, also known as XOFF), and 127 (Delete) became de facto standards. The Model 33 was also notable for taking the description of Control-G (BEL, meaning audibly alert the operator) literally as the unit contained an actual bell which it rang when it received a BEL character. Because the keytop for the O key also showed a left-arrow symbol (from ASCII-1963, which had this character instead of underscore), a noncompliant use of code 15 (Control-O, Shift In) interpreted as "delete previous character" was also adopted by many early timesharing systems but eventually became neglected.


The inherent ambiguity of many control characters, combined with their historical usage, created problems when transferring "plain text" files between systems. The best example of this is the newline problem on various operating systems. Teletype machines required that a line of text be terminated with both "Carriage Return" (which moves the printhead to the beginning of the line) and "Line Feed" (which advances the paper one line without moving the printhead). The name "Carriage Return" comes from the fact that on a manual typewriter the carriage holding the paper moved while the position where the typebars struck the ribbon remained stationary. The entire carriage had to be pushed (returned) to the right in order to position the left margin of the paper for the next line.


Many of the non-alphanumeric characters were positioned to correspond to their shifted position on typewriters; an important subtlety is that these were based on mechanical typewriters, not electric typewriters. Mechanical typewriters followed the standard set by the Remington No. 2 (1878), the first typewriter with a shift key, and the shifted values of 23456789- were "#$%_&'() –  early typewriters omitted 0 and 1, using O (capital letter o) and l (lowercase letter L) instead, but 1! and 0) pairs became standard once 0 and 1 became common. Thus, in ASCII !"#$% were placed in second column, rows 1–5, corresponding to the digits 1–5 in the adjacent column. The parentheses could not correspond to 9 and 0, however, because the place corresponding to 0 was taken by the space character. This was accommodated by removing _ (underscore) from 6 and shifting the remaining characters left, which corresponded to many European typewriters that placed the parentheses with 8 and 9. This discrepancy from typewriters led to bit-paired keyboards, notably the Teletype Model 33, which used the left-shifted layout corresponding to ASCII, not to traditional mechanical typewriters. Electric typewriters, notably the more recently introduced IBM Selectric (1961), used a somewhat different layout that has become standard on computers—following the IBM PC (1981), especially Model M (1984)—and thus shift values for symbols on modern keyboards do not correspond as closely to the ASCII table as earlier keyboards did. The /? pair also dates to the No. 2, and the ,< .> pairs were used on some keyboards (others, including the No. 2, did not shift , (comma) or . (full stop) so they could be used in uppercase without unshifting). However, ASCII split the ;: pair (dating to No. 2), and rearranged mathematical symbols (varied conventions, commonly -* =+) to :* ;+ -=.


Code 127 is officially named "delete" but the Teletype label was "rubout". Since the original standard did not give detailed interpretation for most control codes, interpretations of this code varied. The original Teletype meaning, and the intent of the standard, was to make it an ignored character, the same as NUL (all zeroes). This was useful specifically for paper tape, because punching the all-ones bit pattern on top of an existing mark would obliterate it. Tapes designed to be "hand edited" could even be produced with spaces of extra NULs (blank tape) so that a block of characters could be "rubbed out" and then replacements put into the empty space.


Unfortunately, requiring two characters to mark the end of a line introduces unnecessary complexity and questions as to how to interpret each character when encountered alone. To simplify matters plain text data streams, including files, on Multics used line feed (LF) alone as a line terminator. Unix and Unix-like systems, and Amiga systems, adopted this convention from Multics. The original Macintosh OS, Apple DOS, and ProDOS, on the other hand, used carriage return (CR) alone as a line terminator; however, since Apple replaced these operating systems with the Unix-based OS X operating system, they now use line feed (LF) as well.


In the human digestive system, food enters the mouth and mechanical digestion of the food starts by the action of mastication (chewing), a form of mechanical digestion, and the wetting contact of saliva. Saliva, a liquid secreted by the salivary glands, contains salivary amylase, an enzyme which starts the digestion of starch in the food; the saliva also contains mucus, which lubricates the food, and hydrogen carbonate, which provides the ideal conditions of pH (alkaline) for amylase to work. After undergoing mastication and starch digestion, the food will be in the form of a small, round slurry mass called a bolus. It will then travel down the esophagus and into the stomach by the action of peristalsis. Gastric juice in the stomach starts protein digestion. Gastric juice mainly contains hydrochloric acid and pepsin. As these two chemicals may damage the stomach wall, mucus is secreted by the stomach, providing a slimy layer that acts as a shield against the damaging effects of the chemicals. At the same time protein digestion is occurring, mechanical mixing occurs by peristalsis, which is waves of muscular contractions that move along the stomach wall. This allows the mass of food to further mix with the digestive enzymes.


Other animals, such as rabbits and rodents, practise coprophagia behaviours - eating specialised faeces in order to re-digest food, especially in the case of roughage. Capybara, rabbits, hamsters and other related species do not have a complex digestive system as do, for example, ruminants. Instead they extract more nutrition from grass by giving their food a second pass through the gut. Soft faecal pellets of partially digested food are excreted and generally consumed immediately. They also produce normal droppings, which are not eaten.


Digestive systems take many forms. There is a fundamental distinction between internal and external digestion. External digestion developed earlier in evolutionary history, and most fungi still rely on it. In this process, enzymes are secreted into the environment surrounding the organism, where they break down an organic material, and some of the products diffuse back to the organism. Animals have a tube (gastrointestinal tract) in which internal digestion occurs, which is more efficient because more of the broken down products can be captured, and the internal chemical environment can be more efficiently controlled.


The nitrogen fixing Rhizobia are an interesting case, wherein conjugative elements naturally engage in inter-kingdom conjugation. Such elements as the Agrobacterium Ti or Ri plasmids contain elements that can transfer to plant cells. Transferred genes enter the plant cell nucleus and effectively transform the plant cells into factories for the production of opines, which the bacteria use as carbon and energy sources. Infected plant cells form crown gall or root tumors. The Ti and Ri plasmids are thus endosymbionts of the bacteria, which are in turn endosymbionts (or parasites) of the infected plant.


Teeth (singular tooth) are small whitish structures found in the jaws (or mouths) of many vertebrates that are used to tear, scrape, milk and chew food. Teeth are not made of bone, but rather of tissues of varying density and hardness, such as enamel, dentine and cementum. Human teeth have a blood and nerve supply which enables proprioception. This is the ability of sensation when chewing, for example if we were to bite into something too hard for our teeth, such as a chipped plate mixed in food, our teeth send a message to our brain and we realise that it cannot be chewed, so we stop trying.


The abomasum is the fourth and final stomach compartment in ruminants. It is a close equivalent of a monogastric stomach (e.g., those in humans or pigs), and digesta is processed here in much the same way. It serves primarily as a site for acid hydrolysis of microbial and dietary protein, preparing these protein sources for further digestion and absorption in the small intestine. Digesta is finally moved into the small intestine, where the digestion and absorption of nutrients occurs. Microbes produced in the reticulo-rumen are also digested in the small intestine.


An earthworm's digestive system consists of a mouth, pharynx, esophagus, crop, gizzard, and intestine. The mouth is surrounded by strong lips, which act like a hand to grab pieces of dead grass, leaves, and weeds, with bits of soil to help chew. The lips break the food down into smaller pieces. In the pharynx, the food is lubricated by mucus secretions for easier passage. The esophagus adds calcium carbonate to neutralize the acids formed by food matter decay. Temporary storage occurs in the crop where food and calcium carbonate are mixed. The powerful muscles of the gizzard churn and mix the mass of food and dirt. When the churning is complete, the glands in the walls of the gizzard add enzymes to the thick paste, which helps chemically breakdown the organic matter. By peristalsis, the mixture is sent to the intestine where friendly bacteria continue chemical breakdown. This releases carbohydrates, protein, fat, and various vitamins and minerals for absorption into the body.


Digestion of some fats can begin in the mouth where lingual lipase breaks down some short chain lipids into diglycerides. However fats are mainly digested in the small intestine. The presence of fat in the small intestine produces hormones that stimulate the release of pancreatic lipase from the pancreas and bile from the liver which helps in the emulsification of fats for absorption of fatty acids. Complete digestion of one molecule of fat (a triglyceride) results a mixture of fatty acids, mono- and di-glycerides, as well as some undigested triglycerides, but no free glycerol molecules.


Digestion is the breakdown of large insoluble food molecules into small water-soluble food molecules so that they can be absorbed into the watery blood plasma. In certain organisms, these smaller substances are absorbed through the small intestine into the blood stream. Digestion is a form of catabolism that is often divided into two processes based on how food is broken down: mechanical and chemical digestion. The term mechanical digestion refers to the physical breakdown of large pieces of food into smaller pieces which can subsequently be accessed by digestive enzymes. In chemical digestion, enzymes break down food into the small molecules the body can use.


Different phases of digestion take place including: the cephalic phase , gastric phase, and intestinal phase. The cephalic phase occurs at the sight, thought and smell of food, which stimulate the cerebral cortex. Taste and smell stimuli are sent to the hypothalamus and medulla oblongata. After this it is routed through the vagus nerve and release of acetylcholine. Gastric secretion at this phase rises to 40% of maximum rate. Acidity in the stomach is not buffered by food at this point and thus acts to inhibit parietal (secretes acid) and G cell (secretes gastrin) activity via D cell secretion of somatostatin. The gastric phase takes 3 to 4 hours. It is stimulated by distension of the stomach, presence of food in stomach and decrease in pH. Distention activates long and myenteric reflexes. This activates the release of acetylcholine, which stimulates the release of more gastric juices. As protein enters the stomach, it binds to hydrogen ions, which raises the pH of the stomach. Inhibition of gastrin and gastric acid secretion is lifted. This triggers G cells to release gastrin, which in turn stimulates parietal cells to secrete gastric acid. Gastric acid is about 0.5% hydrochloric acid (HCl), which lowers the pH to the desired pH of 1-3. Acid release is also triggered by acetylcholine and histamine. The intestinal phase has two parts, the excitatory and the inhibitory. Partially digested food fills the duodenum. This triggers intestinal gastrin to be released. Enterogastric reflex inhibits vagal nuclei, activating sympathetic fibers causing the pyloric sphincter to tighten to prevent more food from entering, and inhibits local reflexes.


In a channel transupport system, several proteins form a contiguous channel traversing the inner and outer membranes of the bacteria. It is a simple system, which consists of only three protein subunits: the ABC protein, membrane fusion protein (MFP), and outer membrane protein (OMP)[specify]. This secretion system transports various molecules, from ions, drugs, to proteins of various sizes (20 - 900 kDa). The molecules secreted vary in size from the small Escherichia coli peptide colicin V, (10 kDa) to the Pseudomonas fluorescens cell adhesion protein LapA of 900 kDa.


In addition to the use of the multiprotein complexes listed above, Gram-negative bacteria possess another method for release of material: the formation of outer membrane vesicles. Portions of the outer membrane pinch off, forming spherical structures made of a lipid bilayer enclosing periplasmic materials. Vesicles from a number of bacterial species have been found to contain virulence factors, some have immunomodulatory effects, and some can directly adhere to and intoxicate host cells. While release of vesicles has been demonstrated as a general response to stress conditions, the process of loading cargo proteins seems to be selective.


Underlying the process is muscle movement throughout the system through swallowing and peristalsis. Each step in digestion requires energy, and thus imposes an "overhead charge" on the energy made available from absorbed substances. Differences in that overhead cost are important influences on lifestyle, behavior, and even physical structures. Examples may be seen in humans, who differ considerably from other hominids (lack of hair, smaller jaws and musculature, different dentition, length of intestines, cooking, etc.).


Digestion begins in the mouth with the secretion of saliva and its digestive enzymes. Food is formed into a bolus by the mechanical mastication and swallowed into the esophagus from where it enters the stomach through the action of peristalsis. Gastric juice contains hydrochloric acid and pepsin which would damage the walls of the stomach and mucus is secreted for protection. In the stomach further release of enzymes break down the food further and this is combined with the churning action of the stomach. The partially digested food enters the duodenum as a thick semi-liquid chyme. In the small intestine, the larger part of digestion takes place and this is helped by the secretions of bile, pancreatic juice and intestinal juice. The intestinal walls are lined with villi, and their epithelial cells is covered with numerous microvilli to improve the absorption of nutrients by increasing the surface area of the intestine.


Lactase is an enzyme that breaks down the disaccharide lactose to its component parts, glucose and galactose. Glucose and galactose can be absorbed by the small intestine. Approximately 65 percent of the adult population produce only small amounts of lactase and are unable to eat unfermented milk-based foods. This is commonly known as lactose intolerance. Lactose intolerance varies widely by ethnic heritage; more than 90 percent of peoples of east Asian descent are lactose intolerant, in contrast to about 5 percent of people of northern European descent.


After some time (typically 1–2 hours in humans, 4–6 hours in dogs, 3–4 hours in house cats),[citation needed] the resulting thick liquid is called chyme. When the pyloric sphincter valve opens, chyme enters the duodenum where it mixes with digestive enzymes from the pancreas and bile juice from the liver and then passes through the small intestine, in which digestion continues. When the chyme is fully digested, it is absorbed into the blood. 95% of absorption of nutrients occurs in the small intestine. Water and minerals are reabsorbed back into the blood in the colon (large intestine) where the pH is slightly acidic about 5.6 ~ 6.9. Some vitamins, such as biotin and vitamin K (K2MK7) produced by bacteria in the colon are also absorbed into the blood in the colon. Waste material is eliminated from the rectum during defecation.


In mammals, preparation for digestion begins with the cephalic phase in which saliva is produced in the mouth and digestive enzymes are produced in the stomach. Mechanical and chemical digestion begin in the mouth where food is chewed, and mixed with saliva to begin enzymatic processing of starches. The stomach continues to break food down mechanically and chemically through churning and mixing with both acids and enzymes. Absorption occurs in the stomach and gastrointestinal tract, and the process finishes with defecation.


Protein digestion occurs in the stomach and duodenum in which 3 main enzymes, pepsin secreted by the stomach and trypsin and chymotrypsin secreted by the pancreas, break down food proteins into polypeptides that are then broken down by various exopeptidases and dipeptidases into amino acids. The digestive enzymes however are mostly secreted as their inactive precursors, the zymogens. For example, trypsin is secreted by pancreas in the form of trypsinogen, which is activated in the duodenum by enterokinase to form trypsin. Trypsin then cleaves proteins to smaller polypeptides.


Gymnasts sprint down a runway, which is a maximum of 25 meters in length, before hurdling onto a spring board. The gymnast is allowed to choose where they start on the runway. The body position is maintained while "punching" (blocking using only a shoulder movement) the vaulting platform. The gymnast then rotates to a standing position. In advanced gymnastics, multiple twists and somersaults may be added before landing. Successful vaults depend on the speed of the run, the length of the hurdle, the power the gymnast generates from the legs and shoulder girdle, the kinesthetic awareness in the air, and the speed of rotation in the case of more difficult and complex vaults.


According to FIG rules, only women compete in rhythmic gymnastics. This is a sport that combines elements of ballet, gymnastics, dance, and apparatus manipulation. The sport involves the performance of five separate routines with the use of five apparatus; ball, ribbon, hoop, clubs, rope—on a floor area, with a much greater emphasis on the aesthetic rather than the acrobatic. There are also group routines consisting of 5 gymnasts and 5 apparatuses of their choice. Rhythmic routines are scored out of a possible 30 points; the score for artistry (choreography and music) is averaged with the score for difficulty of the moves and then added to the score for execution.


Aesthetic Group Gymnastics (AGG) was developed from the Finnish "naisvoimistelu". It differs from Rhythmic Gymnastics in that body movement is large and continuous and teams are larger' Athletes do not use apparatus in international AGG competitions compared to Rhythmic Gymnastics where ball, ribbon, hoop and clubs are used on the floor area. The sport requires physical qualities such as flexibility, balance, speed, strength, coordination and sense of rhythm where movements of the body are emphasized through the flow, expression and aesthetic appeal. A good performance is characterized by uniformity and simultaneity. The competition program consists of versatile and varied body movements, such as body waves, swings, balances, pivots, jumps and leaps, dance steps, and lifts. The International Federation of Aesthetic Group Gymnastics (IFAGG) was established in 2003.


This apparatus may be made of hemp or a synthetic material which retains the qualities of lightness and suppleness. Its length is in proportion to the size of the gymnast. The rope should, when held down by the feet, reach both of the gymnasts' armpits. One or two knots at each end are for keeping hold of the rope while doing the routine. At the ends (to the exclusion of all other parts of the rope) an anti-slip material, either coloured or neutral may cover a maximum of 10 cm (3.94 in). The rope must be coloured, either all or partially and may either be of a uniform diameter or be progressively thicker in the center provided that this thickening is of the same material as the rope. The fundamental requirements of a rope routine include leaps and skipping. Other elements include swings, throws, circles, rotations and figures of eight. In 2011, the FIG decided to nullify the use of rope in rhythmic gymnastic competitions.


The Federation of International Gymnastics (FIG) was founded in Liege in 1881. By the end of the nineteenth century, men's gymnastics competition was popular enough to be included in the first "modern" Olympic Games in 1896. From then on until the early 1950s, both national and international competitions involved a changing variety of exercises gathered under the rubric, gymnastics, that would seem strange to today's audiences and that included for example, synchronized team floor calisthenics, rope climbing, high jumping, running, and horizontal ladder. During the 1920s, women organized and participated in gymnastics events. The first women's Olympic competition was primitive, only involving synchronized calisthenics and track and field. These games were held in 1928, in Amsterdam.


In the vaulting events, gymnasts sprint down a 25 metres (82 ft) runway, jump onto a spring filled board or perform a roundoff, or handspring entry onto a springboard (run/ take-off segment), land momentarily, inverted on the hands on the vaulting horse, or vaulting table (pre flight segment), then propel themselves forward or backward, off of this platform to a two footed landing (post flight segment). Every gymnast starts at a different point on the vault runway depending on their height and strength. The post flight segment may include one or more multiple saltos or somersaults, and/or twisting movements. A round-off entry vault, called a Yurchenko, is the most common vault in elite level gymnastics. When performing a yurchenko, gymnasts "round-off" so hands are on the runway while the feet land on the springboard (beatboard). From the roundoff position the gymnast travels backwards and executes a backhandspring so that the hands land on the vaulting table. The gymnast then blocks off the vaulting platform into various twisting and/or somersaulting combinations. The post flight segment brings the gymnast to her feet.


A gymnast's score comes from deductions taken from their start value. The start value of a routine is based on the difficulty of the elements the gymnast attempts and whether or not the gymnast meets composition requirements. The composition requirements are different for each apparatus; this score is called the D score. Deductions in execution and artistry are taken from 10.0. This score is called the E score. The final score is calculated by taking deductions from the E score, and adding the result to the D score. Since 2007, the scoring system has changed by adding bonus plus the execution and then adding those two together to get the final score.


The technical rules for the Japanese version of men's rhythmic gymnastics came around the 1970s. For individuals, only four types of apparatus are used: the double rings, the stick, the rope, and the clubs. Groups do not use any apparatus. The Japanese version includes tumbling performed on a spring floor. Points are awarded based a 10-point scale that measures the level of difficulty of the tumbling and apparatus handling. On November 27–29, 2003, Japan hosted first edition of the Men's Rhythmic Gymnastics World Championship.


The word gymnastics derives from the common Greek adjective γυμνός (gymnos) meaning "naked", by way of the related verb γυμνάζω (gymnazo), whose meaning is to train naked", "train in gymnastic exercise", generally "to train, to exercise". The verb had this meaning, because athletes in ancient times exercised and competed without clothing. It came into use in the 1570s, from Latin gymnasticus, from Greek gymnastikos "fond of or skilled in bodily exercise," from gymnazein "to exercise or train" (see gymnasium).


By 1954, Olympic Games apparatus and events for both men and women had been standardized in modern format, and uniform grading structures (including a point system from 1 to 15) had been agreed upon. At this time, Soviet gymnasts astounded the world with highly disciplined and difficult performances, setting a precedent that continues. The new medium of television has helped publicize and initiate a modern age of gymnastics. Both men's and women's gymnastics now attract considerable international interest, and excellent gymnasts can be found on every continent. Nadia Comăneci received the first perfect score, at the 1976 Summer Olympics held in Montreal, Canada. She was coached in Romania by coach, (Hungarian ethnicity), Béla Károlyi. Comaneci scored four of her perfect tens on the uneven bars, two on the balance beam and one in the floor exercise. Even with Nadia's perfect scores, the Romanians lost the gold medal to the Soviet Union. Nevertheless, Comaneci became an Olympic icon.


A typical pommel horse exercise involves both single leg and double leg work. Single leg skills are generally found in the form of scissors, an element often done on the pommels. Double leg work however, is the main staple of this event. The gymnast swings both legs in a circular motion (clockwise or counterclockwise depending on preference) and performs such skills on all parts of the apparatus. To make the exercise more challenging, gymnasts will often include variations on a typical circling skill by turning (moores and spindles) or by straddling their legs (Flares). Routines end when the gymnast performs a dismount, either by swinging his body over the horse, or landing after a handstand. This requires back muscles to do any sort of skill. From handstands being easy to back or front flips being a little difficult.


In a tumbling pass, dismount or vault, landing is the final phase, following take off and flight  This is a critical skill in terms of execution in competition scores, general performance, and injury occurrence. Without the necessary magnitude of energy dissipation during impact, the risk of sustaining injuries during somersaulting increases. These injuries commonly occur at the lower extremities such as: cartilage lesions, ligament tears, and bone bruises/fractures. To avoid such injuries, and to receive a high performance score, proper technique must be used by the gymnast. "The subsequent ground contact or impact landing phase must be achieved using a safe, aesthetic and well-executed double foot landing."  A successful landing in gymnastics is classified as soft, meaning the knee and hip joints are at greater than 63 degrees of flexion.


Individual routines in trampolining involve a build-up phase during which the gymnast jumps repeatedly to achieve height, followed by a sequence of ten bounces without pause during which the gymnast performs a sequence of aerial skills. Routines are marked out of a maximum score of 10 points. Additional points (with no maximum at the highest levels of competition) can be earned depending on the difficulty of the moves and the length of time taken to complete the ten skills which is an indication of the average height of the jumps. In high level competitions, there are two preliminary routines, one which has only two moves scored for difficulty and one where the athlete is free to perform any routine. This is followed by a final routine which is optional. Some competitions restart the score from zero for the finals, other add the final score to the preliminary results.


On the uneven bars, the gymnast performs a routine on two horizontal bars set at different heights. These bars are made of fiberglass covered in wood laminate, to prevent them from breaking. In the past, bars were made of wood, but the bars were prone to breaking, providing an incentive to switch to newer technologies. The width and height of the bars may be adjusted. In the past, the uneven parallel bars were closer together. They've been moved increasingly further apart, allowing gymnasts to perform swinging, circling, transitional, and release moves, that may pass over, under, and between the two bars. At the Elite level, movements must pass through the handstand. Gymnasts often mount the Uneven Bars using a springboard, or a small mat. Chalk and grips (a leather strip with holes for fingers to protect hands and improve performance) may be used while doing this event. The chalk helps take the moisture out of gymnast's hands to decrease friction and prevent rips (tears to the skin of the hands), dowel grips help gymnasts grip the bar.


A higher flight phase results in a higher vertical ground reaction force. Vertical ground reaction force represents external force which the gymnasts have to overcome with their muscle force and has an impact on the gymnasts linear and angular momentum. Another important variable that affects linear and angular momentum is time the landing takes Gymnasts can alter the shape of the area by increasing the time taken to perform the landing. Gymnasts can achieve this by increasing hip, knee and ankle amplitude. With the increase of height, the amplitude in ankles knees and hips rise the bars.


Aerobic gymnastics (formally Sport Aerobics) involves the performance of routines by individuals, pairs, trios or groups up to 6 people, emphasizing strength, flexibility, and aerobic fitness rather than acrobatic or balance skills. Routines are performed for all individuals on a 7x7m floor and also for 12–14 and 15-17 trios and mixed pairs. From 2009, all senior trios and mixed pairs were required to be on the larger floor (10x10m), all groups also perform on this floor. Routines generally last 60–90 seconds depending on age of participant and routine category.


General gymnastics enables people of all ages and abilities to participate in performance groups of 6 to more than 150 athletes. They perform synchronized, choreographed routines. Troupes may consist of both genders and are not separated into age divisions. The largest general gymnastics exhibition is the quadrennial World Gymnaestrada which was first held in 1939. In 1984 Gymnastics for All was officially recognized first as a Sport Program by the FIG (International Gymnastic Federation), and subsequently by national gymnastic federations worldwide with participants that now number 30 million.


Gymnastics is a sport involving the performance of exercises requiring strength, flexibility, balance and control. Internationally, all events are governed by the Fédération Internationale de Gymnastique (FIG). Each country has its own national governing body (BIW) affiliated to FIG. Competitive artistic gymnastics is the best known of the gymnastic events. It typically involves the women's events of vault, uneven bars, balance beam, and floor exercise. Men's events are floor exercise, pommel horse, still rings, vault, parallel bars, and the high bar. Gymnastics evolved from exercises used by the ancient Greeks that included skills for mounting and dismounting a horse, and from circus performance skills.


In the late eighteenth- and early nineteenth-century Germany, three pioneer physical educators – Johann Friedrich GutsMuths (1759–1839) and Friedrich Ludwig Jahn (1778–1852) – created exercises for boys and young men on apparatus they had designed that ultimately led to what is considered modern gymnastics. Don Francisco Amorós y Ondeano, was born on February 19, 1770 in Valence and died on August 8, 1848 in Paris. He was a Spanish colonel, and the first person to introduce educative gymnastic in France. Jahn promoted the use of parallel bars, rings and high bar in international competition.


In 2006, FIG introduced a new points system for Artistic gymnastics in which scores are no longer limited to 10 points. The system is used in the US for elite level competition. Unlike the old code of points, there are two separate scores, an execution score and a difficulty score. In the previous system, the "execution score" was the only score. It was and still is out of 10.00. During the gymnast's performance, the judges deduct this score only. A fall, on or off the event, is a 1.00 deduction, in elite level gymnastics. The introduction of the difficulty score is a significant change. The gymnast's difficulty score is based on what elements they perform and is subject to change if they do not perform or complete all the skills, or they do not connect a skill meant to be connected to another. Connection bonuses are the most common deduction from a difficulty score, as it can be difficult to connect multiple flight elements. It is very hard to connect skills if the first skill is not performed correctly. The new code of points allows the gymnasts to gain higher scores based on the difficulty of the skills they perform as well as their execution. There is no maximum score for difficulty, as it can keep increasing as the difficulty of the skills increase.


In the past, the floor exercise event was executed on the bare floor or mats such as wrestling mats. Today, the floor event occurs on a carpeted 12m × 12m square, usually consisting of hard foam over a layer of plywood, which is supported by springs or foam blocks generally called a "spring" floor. This provides a firm surface that provides extra bounce or spring when compressed, allowing gymnasts to achieve greater height and a softer landing after the composed skill. Gymnasts perform a choreographed routine up to 90 seconds in the floor exercise event; Depending on the level, they may choose their own, or, if known as a "compulsory gymnast," default music must be played. In some gymnastic associations such as United States Association of Gymnastic Clubs (USAIGC), gymnasts are allowed to have vocals in their music but at USA Gymnastics competitions a large deduction is taken from the score for having vocals in the music. The routine should consist of tumbling lines, series of jumps, leaps, dance elements, acrobatic skills, and turns, or piviots, on one foot. A gymnast can perform up to four tumbling lines that usually includes at least one flight element without hand support. Each level of gymnastics requires the athlete to perform a different number of tumbling passes. In level 7 in the United States, a gymnast is required to do 2–3, and in levels 8–10, at least 3–4 tumbling passes are required.


In tumbling, athletes perform an explosive series of flips and twists down a sprung tumbling track. Scoring is similar to trampolining. Tumbling was originally contested as one of the events in Men's Artistic Gymnastics at the 1932 Summer Olympics, and in 1955 and 1959 at the Pan American Games. From 1974 to 1998 it was included as an event for both genders at the Acrobatic Gymnastics World Championships. The event has also been contested since 1976 at the Trampoline World Championships. Since the recognition of Trampoline and Acrobatic Gymnastics as FIG disciplines in 1999, official Tumbling competitions are only allowed as an event in Trampoline gymnastics meets.


Men's rhythmic gymnastics is related to both Men's artistic gymnastics and wushu martial arts. It emerged in Japan from stick gymnastics. Stick gymnastics has been taught and performed for many years with the aim of improving physical strength and health. Male athletes are judged on some of the same physical abilities and skills as their female counterparts, such as hand/body-eye co-ordination, but tumbling, strength, power, and martial arts skills are the main focus, as opposed to flexibility and dance in women's rhythmic gymnastics. There are a growing number of participants, competing alone and on a team; it is most popular in Asia, especially in Japan where high school and university teams compete fiercely. As of 2002[update], there were 1000 men's rhythmic gymnasts in Japan.[citation needed]


Domestically, Barcelona has won 23 La Liga, 27 Copa del Rey, 11 Supercopa de España, 3 Copa Eva Duarte and 2 Copa de la Liga trophies, as well as being the record holder for the latter four competitions. In international club football, Barcelona has won five UEFA Champions League titles, a record four UEFA Cup Winners' Cup, a shared record five UEFA Super Cup, a record three Inter-Cities Fairs Cup and a record three FIFA Club World Cup trophies. Barcelona was ranked first in the IFFHS Club World Ranking for 1997, 2009, 2011, 2012 and 2015 and currently occupies the second position on the UEFA club rankings. The club has a long-standing rivalry with Real Madrid; matches between the two teams are referred to as El Clásico.


On 14 June 1925, in a spontaneous reaction against Primo de Rivera's dictatorship, the crowd in the stadium jeered the Royal March. As a reprisal, the ground was closed for six months and Gamper was forced to relinquish the presidency of the club. This coincided with the transition to professional football, and, in 1926, the directors of Barcelona publicly claimed, for the first time, to operate a professional football club. On 3 July 1927, the club held a second testimonial match for Paulino Alcántara, against the Spanish national team. To kick off the match, local journalist and pilot Josep Canudas dropped the ball onto the pitch from his airplane. In 1928, victory in the Spanish Cup was celebrated with a poem titled "Oda a Platko", which was written by a member of the Generation of '27, Rafael Alberti, inspired by the heroic performance of the Barcelona goalkeeper, Franz Platko. On 23 June 1929, Barcelona won the inaugural Spanish League. A year after winning the championship, on 30 July 1930, Gamper committed suicide after a period of depression brought on by personal and financial problems.


The 1973–74 season saw the arrival of Johan Cruyff, who was bought for a world record £920,000 from Ajax. Already an established player with Ajax, Cruyff quickly won over the Barcelona fans when he told the European press that he chose Barcelona over Real Madrid because he could not play for a club associated with Francisco Franco. He further endeared himself when he named his son Jordi, after the local Catalan Saint George. Next to champions like Juan Manuel Asensi, Carles Rexach and Hugo Sotil, he helped the club win the 1973–74 season for the first time since 1960, defeating Real Madrid 5–0 at the Bernabéu along the way. He was crowned European Footballer of the Year in 1973 during his first season with Barcelona (his second Ballon d'Or win; he won his first while playing for Ajax in 1971). Cruyff received this prestigious award a third time (the first player to do so) in 1974, while he was still with Barcelona.


Like Maradona, Ronaldo only stayed a short time before he left for Internazionale. However, new heroes emerged, such as Luís Figo, Patrick Kluivert, Luis Enrique and Rivaldo, and the team won a Copa del Rey and La Liga double in 1998. In 1999, the club celebrated its centenari, winning the Primera División title, and Rivaldo became the fourth Barcelona player to be awarded European Footballer of the Year. Despite this domestic success, the failure to emulate Real Madrid in the Champions League led to van Gaal and Núñez resigning in 2000.