Sakéwanten, kahanan korespondensi sederana pantaraning alel miwah sifat maguna ring makudang-kudang kasus , katahan sané madué sifat kompleks miwah kaénterang olih lokus sifat kuantitatif (makudang gen sané magubugan).
However, while this simple correspondence between an allele and a trait works in some cases, most traits are more complex and are controlled by quantitative trait loci (multiple interacting genes).