Konsep indik komposisi komunitas entikan sekadi alas sané entikané madon ageng sampun magentos saking pamargin suksesi ekologi sané kalimbakang olih Henry Chandler Cowles, Arthur Tansley miwah Frederic Clements.
The concept that the composition of plant communities such as temperate broadleaf forest changes by a process of ecological succession was developed by Henry Chandler Cowles, Arthur Tansley and Frederic Clements.