Nincapang kawruhan indik anatomi entik-entikan, morfologi miwah pamargin urip ngawinang kauningin akéh pisanafinitas alami pantaraning entikan yéning bandingang sistem seksual pakaryan Linnaeus.
Increasing knowledge of plant anatomy, morphology and life cycles led to the realisation that there were more natural affinities between plants than the artificial sexual system of Linnaeus.