Horn of Africa
A study by Hassan et al. (2008) analysed the Y-DNA of populations in the Sudan region, with various local Nilotic groups included for comparison. The researchers found the signature Nilotic A and B clades to be the most common paternal lineages amongst the Nilo-Saharan speakers, except those inhabiting western Sudan. There, a prominent North African influence was noted. Haplogroup A was observed amongst 62% of Dinka, 53.3% of Shilluk, 46.4% of Nuba, 33.3% of Nuer, 31.3% of Fur and 18.8% of Masalit. Haplogroup B was found in 50% of Nuer, 26.7% of Shilluk, 23% of Dinka, 14.3% of Nuba, 3.1% of Fur and 3.1% of Masalit. The E1b1b clade was also observed in 71.9% of the Masalit, 59.4% of the Fur, 39.3% of the Nuba, 20% of the Shilluk, 16.7% of the Nuer, and 15% of the Dinka. Hassan et al. attributed the atypically high frequencies of the haplogroup in the Masalit to either a recent population bottleneck, which likely altered the community’s original haplogroup diversity, or to geographical proximity to E1b1b’s place of origin in North Africa. The researchers suggest that the clade “might have been brought to Sudan […] after the progressive desertification of the Sahara around 6,000–8,000 years ago”. Henn et al. (2008) similarly observed Afro-Asiatic influence in the Nilotic Datog of northern Tanzania, 43% of whom carried the M293 sub-clade of E1b1b.