Seymour Benzer
At Purdue University, Benzer developed the "T4 rII system", a new genetic technique. This used recombination in T4 bacteriophage rII mutants to map the inside structure of genes. Benzer realized that by generating many r mutants and recording the recombination frequency between different r strains, he could make a detailed map of the gene, much as Alfred Sturtevant had done for chromosomes.<ref name=Weiner>Weiner, Jonathan 1999. Time, love, memory: a great biologist and his quest for the origins of behavior. New York: Knopf. </ref>