Plays a key role in chromosome recombination during meiosis. Required for the initiation of meiotic recombination and the recruitment of PAIR2 onto meiotic chromosomes. Essential for meiotic DNA double-strand break (DSB) formation. Interacts with PAIR1 and ZEP1. Appears as punctuate foci on chromosomes at leptotene. Colocalizes with ZEP1 to the central region of the synaptonemal complex. Highly expressed in young panicles. Male and female sterility due to defects in meiosis. Belongs to the AAA ATPase family. PCH2 subfamily.