Acts as component of the GARP complex that is involved in retrograde transport from early and late endosomes to the trans-Golgi network (TGN). The GARP complex facilitates tethering as well as SNARE complex assembly at the Golgi (By similarity). Probably involved in pollen tube elongation and other polar growth. Component of the Golgi-associated retrograde protein (GARP) complex, composed by VPS52, VPS53 and VPS54. Interacts directly with VPS53. Localized in the GARP complex in the Golgi and post-Golgi compartments. Present in pollen. Mostly expressed in roots and flower buds, and, at lower levels, in vegetative tissues and mature flowers. Lethal when homozygous. In hemizygous plants, male-specific transmission defect. Belongs to the VPS54 family.