Involved in the regulation of leaf shape formation (PubMed:27473144). May function by coordinating the expression of genes associated with leaf and bulliform cell development (PubMed:27473144). Highly expressed in young leaves and panicles (PubMed:27473144). Expressed at low levels in roots (PubMed:27473144). Adaxially rolling and erect leaves which leads to plants with erect architecture and reduced lamina joint angle (PubMed:27473144). Abnormal bulliform cell number, size and arrangement in leaf blades (PubMed:27473144). Dark-green leaves with increased levels of chlorophylls (PubMed:27473144). Reduced number of tillers, altered grain morphology, and reduced number of grains per main panicle (PubMed:27473144). Reduced number and length of adventitious roots (PubMed:27473144).