Plays a pivotal role in the control of morphogenesis of shoot organs by negatively regulating the expression of boundary-specific genes such as CUC genes, probably through the induction of miRNA (e.g. miR164). Participates in ovule develpment (PubMed:25378179). Interacts with SPL. Expressed in cotyledons, particularly in the vascular region, in leaves, buds, flowers and immature siliques. Expressed during ovule development (PubMed:25378179).