Regulates tapetum degeneration and pollen maturation during anther development. Interacts with SKP1A. Expressed in flower buds, developing anthers, pollen grains, siliques, rosette leaves and roots. Detected at lower levels in open flowers, stems and cauline leaves. Expressed in young seedling in the hydathodes, shoot apical meristem, root tips and lateral root primordia. Expressed to a high level in young floral buds, but decreases in the later floral developmental stages. Expression restricted to microspores and pollen grains at the later developmental stages of anther.