Inhibits voltage-gated ion channels. Expressed by the venom duct. Only expressed in adults. The presence of a 'disulfide through disulfide knot' structurally defines this protein as a knottin. The cysteine framework is VI/VII (C-C-CC-C-C). Belongs to the conotoxin O2 superfamily. Contains only 5 Cys residues instead of the 6 residues that are usually found in the O2 superfamily.