Involved in vesicular trafficking at the Golgi apparatus level. May play a role in delivery of transport vesicles containing GPI-linked proteins from the trans-Golgi network through its interaction with MACF1. Involved in endosome-to-Golgi trafficking. Homodimer. Interacts with GTP-bound ARL1 and ARL3. Interacts with MACF1. Directly interacts with TBC1D23. Interacts with FAM91A1; this interaction may be mediated by TBC1D23. Expressed in the head of epididymal sperm but not in testicular sperm (at protein level). Extended rod-like protein with coiled-coil domains.