Functions as a cytoplasmic linker protein. Involved in TGN-endosome dynamics (By similarity). Homodimer. Localized to Golgi stacks as well as on tubulovesicular elements juxtaposed to Golgi cisternae. Microtubule association is inhibited by the ANK repeats and the Golgi localization region (GoLD). The N-terminal half is dispensable for proper Golgi targeting, whereas the GoLD region is required.