Hydroxycinnamoyl transferase involved in the conjugation of feruloyl CoA to spermidine (PubMed:19077165, PubMed:19762055, PubMed:33519864). Catalyzes the three conjugating steps required for the biosynthesis of N(1),N(4),N(8)-triferuloyl-spermidine (PubMed:19077165, PubMed:33519864). Spermidine is the only acceptor substrate while feruloyl CoA > caffeoyl CoA > coumaroyl CoA > cinnamoyl CoA >> sinapoyl CoA are efficient acyl donors. No activity with hydroxyferuloyl CoA (PubMed:19077165). Required for the biosynthesis of these conjugated spermidine derivatives, specifically in anther tapetum (PubMed:22912643). Expressed in flower buds and inflorescences. Restricted to anther tapetum. Expressed during bud growth and diappears when the flower opens. Not induced by pathogens or by wounding. Glossy and sticky pollen, but very little changes in the exine patterns. Drastic reduction in flower buds of the content in N1,N4,N8-tri-(hydroxyferuloyl)-spermidine and N1,N4-di(hydroxyferuloyl)-N8-sinapoyl-spermidine. Belongs to the plant acyltransferase family.