Acyltransferase which mediates the conversion of lysophosphatidylinositol (1-acyl-sn-glycero-3-phosphatidylinositol or LPI) into phosphatidylinositol (1,2-diacyl-sn-glycero-3-phosphoinositol or PI) (LPIAT activity). Prefers sn-2-LPI rather than sn-1-LPI as the acyl acceptor. Lysophospholipid acyltransferases (LPLATs) catalyze the reacylation step of the phospholipid remodeling pathway also known as the Lands cycle. Involved in the selective incorporation of arachidonoyl-CoA ((5Z,8Z,11Z,14Z)-eicosatetraenoyl-CoA) and (5Z,8Z,11Z,14Z,17Z)-eicosapentaenoyl-CoA (EPA-CoA) into PI (PubMed:18094042). Besides its role in biomembranes, PI is a precursor of PI 3-phosphate (PIP3) and its fatty acid composition has an important role in PI3P signaling (PubMed:22862955). (5Z,8Z,11Z,14Z,17Z)-eicosapentaenoyl-CoA + 1-octadecanoyl-sn-glycero-3-phospho-(1D-myo-inositol) = 1-octadecanoyl-2-(5Z,8Z,11Z,14Z,17Z-eicosapentaenoyl)-sn-glycero-3-phospho-(1D-myo-inositol) + CoA (5Z,8Z,11Z,14Z,17Z)-eicosapentaenoyl-CoA + a 1-acyl-sn-glycero-3-phospho-(1D-myo-inositol) = a 1-acyl-2-(5Z,8Z,11Z,14Z,17Z-eicosapentaenoyl)-sn-glycero-3-phospho-(1D-myo-inositol) + CoA (5Z,8Z,11Z,14Z)-eicosatetraenoyl-CoA + a 1-acyl-sn-glycero-3-phospho-(1D-myo-inositol) = a 1-acyl-2-(5Z,8Z,11Z,14Z-eicosatetraenoyl)-sn-glycero-3-phospho-(1D-myo-inositol) + CoA Lipid metabolism; phospholipid metabolism. Expressed ubiquitously throughout development from early embryo to larval and adult stages. In adults, strongly expressed in pharyngeal muscle, body wall muscle, vulval cells, distal tip cells, intestinal cells and spermatheca. Mutants show larval arrest at an early developmental stage and egg-laying defects; 14% of mutants accumulate unlaid eggs that hatched internally. Belongs to the membrane-bound acyltransferase family.