Involved in iron loading of the seeds. Acts probably as a transporter of iron- and metal-nicotianamine chelates. Low levels of expression in leaves and shoots, but not detected in roots. Restricted to the vasculature, in the xylem parenchyma surrounding xylem tubes. Expressed in pollen grains, in the vasculature of petals and sepals, in the carpel veins, in the style underneath the stigmatic papillae, in the vascular tissue of the funiculus and in the chalazal endosperm. Highly expressed during leaf senescence. Slight induction by manganese, copper or zinc deficiency. Inhibited upon iron deficiency and induced by iron overload. Plants do not show visible phenotype, but a decreased iron and nicotianamine content in seeds resulting in a transient defect in germination. Belongs to the YSL (TC 2.A.67.2) family.