Acts as one of several non-catalytic accessory components of the cytoplasmic dynein 2 complex (dynein-2 complex), a motor protein complex that drives the movement of cargos along microtubules within cilia and flagella in concert with the intraflagellar transport (IFT) system (By similarity). Required for proper retrograde ciliary transport (PubMed:26044572). Morpholino knockdown of the protein produces a dose dependent typical ciliopathy phenotype, with ventrally curved body axis, hydrocephalus, abnormal otoliths and small eyes. The few embryos surviving to 4 days post fertilization display severe generalized edema, pronephric cysts, and defects of the craniofacial cartilage. Belongs to the dynein light chain Tctex-type family.