Able to repress WUS when constitutively overexpressed, but have no effect on CLV3. Expressed in the shoot apex, stamens, anthers and young siliques. Detected in provascular and vascular tissue. Expressed throughout development. During germination, initially restricted to the hypocotyl, but expression gradually shifts to the root and on day 6, to the vasculature of the cotyledons and then of the leaves. Increased expression in the shoot apex during the transition to flowering. Induced in the inflorescence vasculature and young flower buds as flowering commenced. Expressed in developing embryos from the early heart stage until torpedo stage. Belongs to the fantastic four family.