Transcriptional activator (PubMed:9687312, PubMed:23707421). It is likely involved in the regulation of keratinocytes terminal differentiation in squamous epithelia and hair follicles (By similarity). Required for the maintenance of spermatogenesis (PubMed:22266914). It is involved in the positive regulation of oocyte maturation, probably acting through the control of BMP15 levels and regulation of AKT signaling cascade (By similarity). May also play a role in the early development of embryos (PubMed:9687312). Interacts with HSF2BP (via C-terminus). Relocates to the midpiece of the flagellum during late spermiogenesis in spermatids. Epidermis and germ cells of testis and ovary. Appears to be expressed by primary and secondary spermatocytes and spermatids. Detected in oocytes within the primary follicles. It is also present in secondary oocytes. Phosphorylation on Ser-505 and Ser-509 leads to cytoplasmic localization. Viable, although fertility is severely reduced in both males and females. Males have significantly reduced testis size, with progressive degeneration of seminiferous epithelium and loss of germ cells from 8 weeks of age. Sperm motility is also reduced.