Ca(2+)-independent actin-binding protein. Binds actin microfilaments (MFs). Involved in actin filament bundling, severing and capping. Caps the barbed end of actin filaments and protects them from disassembly. Promotes VLN3-mediated MF severing. Expressed in roots, young leaves, and inflorescences, mostly in the vasculature of roots, leaves, and filaments of the anthers. Also detected in guard cells. Belongs to the villin/gelsolin family.