Required for accurate chromosome segregation in meiosis. Required for pairing to occur between homologous chromosomes. Acts in early recombination steps and ensures pairing fidelity and proper repair of meiotic DNA double-strand-breaks (PubMed:14704428, PubMed:19918061). Regulates recombination and pairing of homologous chromosomes during meiotic prophase by controlling transport of RAD50 from cytoplasm to the nucleus. May affect pairing of the gene-rich fraction of the genome rather than preventing pairing between repetitive DNA elements (PubMed:19918061). Localizes to the cytoplasm during leptotene, zygotene and pachytene. Male and female sterility. During meiosis, synapsis is uncoupled from recombination and pairing in the mutant gametes.