Involved in regulation of intracellular signaling pathways during development. Specifically thought to play a role in canonical and/or non-canonical Wnt signaling pathways through interaction with DSH (Dishevelled) family proteins. Binds to dvl2/dsh and impedes the degradation of beta-catenin (ctnnb1-A and possibly ctnnb1-B), thereby enhancing the transcriptional activation of target genes of the Wnt signaling pathway. Also promotes catenin delta/ctnnd1 stability which in turn promotes zbtb33/kaiso sequestration and thus is involved in the regulation of zbtb33/kaiso-mediated transcriptional repression. May also bind to and directly stimulate the transcriptional activity of tcf7l1-A. Required for eye development and neural patterning. Interacts with dbf4 and tcf7l1-A. Interacts with dvl2/dsh via the C-terminus. Expressed in the animal and dorsal marginal regions at late blastula and early gastrula stages. Expressed predominantly in the anterior neural plate at neurulation. Expressed mainly in the ectodermal placodes, including the eye anlagen and the otic vesicle, at later stages of development. Expressed bpth maternally and zygotically. The C-terminal PDZ-binding motif may mediate interaction with the PDZ domains of DSH (Dishevelled) family proteins. Belongs to the dapper family.