Member of two distinct condensin I complexes, the condensin I complex and the condensin I-like dosage compensation complex (PubMed:19781752, PubMed:19119011). The condensin I complex is required for conversion of interphase chromatin into mitotic-like condensed chromosomes and for chromosome segregation in meiosis and mitosis (PubMed:19781752, PubMed:19119011). As a member of the condensin I complex, further controls crossover number and distribution in meiosis by restricting double strand break formation, probably by influencing higher-order chromosome structure (PubMed:19781752). Regulatory subunit of the condensin I-like dosage compensation complex that associates specifically with hermaphrodite X chromosomes to reduce their gene transcription during interphase, possibly through chromatin reorganization (PubMed:19119011). Component of the condensin I complex, which contains the mix-1/SMC2 and smc-4/SMC4 heterodimer, and three non-SMC subunits that probably regulate the complex: dpy-26, capg-1 and dpy-28 (PubMed:19781752, PubMed:19119011). Within the complex, interacts with dpy-26, mix-1, smc-4 and dpy-28 (PubMed:19781752, PubMed:19119011, PubMed:28301465). Component of the dosage compensation complex, which consists of the condensin I-like components mix-1/SMC2 and dpy-27/SMC4, and the three non-SMC subunits dpy-26, capg-1 and dpy-28 (PubMed:19119011). Within the complex, interacts with mix-1, dpy-27, dpy-26 and dpy-28 (PubMed:19119011, PubMed:28301465). Interacts with smcl-1 (PubMed:28301465). During meiosis and mitosis, localizes to condensed chromosomes in both sexes (PubMed:19119011). Localizes to the spindle midzone in between separating chromosomes during anaphase and to the midbody during cytokinesis (PubMed:23684975). During interphase, localizes specifically to X chromosomes in hermaphrodites after the onset of dosage compensation (PubMed:19119011). Expressed in embryos and in somatic and germline tissues in adult animals (at protein level). RNAi-mediated knockdown disrupts X-chromosome localization of dpy-27, mix-1, dpy-26 and sdc-3. Results in chromosome segregation defects in mitosis leading to aneuploidy. In a sex-1 mutant background, leads to high embryonic lethality. Belongs to the CND3 (condensin subunit 3) family.