Plays a central role in chromosome cohesion during meiosis I by preventing premature dissociation of cohesin complex from centromeres after prophase, when most of cohesin complex dissociates from chromosomes arms. Required for maintenance of centromeric cohesion before prophase II and correct segregation of chromatids during meiosis II. Has apparently no function in mitosis. Localizes to the centromere during meiosis I. REC8 is required for centromeric localization. Highly expressed in tissues containing meiocytes. Expressed at much lower level in leaves and pollen-containing flowers. Belongs to the shugoshin family.