Transcriptional regulator involved in the regulation of cell differentiation in meristems. Probably regulates the expression of various KNAT genes involved in the maintenance of the cells in an undifferentiated, merismastic state. Plays a role in the regulation of gibberellin 20 oxidase and the gibberellin-regulated protein GASA4. Localizes in the nucleus during the cellular differentiation state and may act via a single strand cutting domain (PubMed:17991462). Transcriptional regulator required for the induction of dormancy during late seed development. Interacts genetically with FUS3 and may be component of the same regulatory pathway during embryogenesis. Binds both linear and supercoiled DNA without sequence preference (PubMed:22226340). Localizes to the cytoplasm in non-differentiating cells and to the nucleus in differentiating cells. Expressed in vascular tissues of stems, hypocotyls, leaves and flowers. Expressed in the vascular bundles of xylem in shoot parenchyma cells. Expressed in the remnant cytoplasm of differentiated fiber cells and in protoxylem element of parenchymal cells. During embryogenesis, expressed in the embryo at the late torpedo/cotyledon stage, at the onset of maturation. Not expressed during early stages of seed development or in mature seeds. Contains a bacterial GIY-YIG-like domain which can functionally replace the bacterial GIY-YIG domain in the UVRC protein. Reduced levels of lignin in leaves and stems (PubMed:17991462). Reduced seed germination rate (PubMed:22226340).