High affinity carnitine transporter involved in the active cellular uptake of carnitine. Also transports organic cations. Regulates lateral root development. Expressed in vascular tissues and at sites of lateral root formation. Mostly present in floral buds, flowers and immature siliques. In flowers, expressed in anthers, sepals, petals and peduncles. Accumulates during germination in embryos, reaching a maximum during the radicle emergence. In seedlings, restricted to the vasculature, with highest levels in the collar and the root. Later present in both elongation and root hair zones in roots. Also present in root cap columella. Detected before and during lateral root initiation and emergence and thereafter remained in vascular tissues of each lateral root. Accumulates in embryo upon imbibition. Reduced sensitivity to carnitine and high degree of root branching. Belongs to the major facilitator (TC 2.A.1) superfamily. Organic cation transporter (TC 2.A.1.19) family.