Catalyzes the hydrolysis of cutin, a polyester that forms the structure of plant cuticle (PubMed:20393707, PubMed:22183084, PubMed:25910960, PubMed:33387709). Shows esterase activity towards p-nitrophenol-linked aliphatic esters (pNP-aliphatic esters) (PubMed:20393707, PubMed:22183084, PubMed:25910960, PubMed:33387709). Capable of degrading the plastic poly(ethylene terephthalate) (PET), the most abundant polyester plastic in the world (By similarity). Can also depolymerize the synthetic polyesters poly(epsilon-caprolactone) (PCL), poly(butylene succinate-co-adipate) (PBSA), poly(butylene succinate) (PBS), and poly(lactic acid) (PLA) (PubMed:20393707, PubMed:22183084). an acetyl ester + H2O = acetate + an aliphatic alcohol + H(+) (ethylene terephthalate)(n) + H2O = (ethylene terephthalate)(n-1) + 4-[(2-hydroxyethoxy)carbonyl]benzoate + H(+) a butanoate ester + H2O = an aliphatic alcohol + butanoate + H(+) cutin + H2O = cutin monomers. a hexanoate ester + H2O = an aliphatic alcohol + H(+) + hexanoate an octanoate ester + H2O = an aliphatic alcohol + H(+) + octanoate Can also bind other divalent metal ions with lower efficiency (PubMed:22183084). Calcium ion binding contributes to the thermostability of the protein (PubMed:22183084). Activated by calcium ions (PubMed:22183084). Activated by magnesium ions (PubMed:22183084). Activated by manganese ions (PubMed:22183084). Inhibited by the serine hydrolase inhibitor phenylmethanesulfonyl fluoride (PMSF) (PubMed:22183084). Inhibited by the chelator ethylenediaminetetraacetic acid (EDTA) (PubMed:22183084). Inhibited by iron ions (PubMed:22183084). Inhibited by aluminum ions (PubMed:22183084). Inhibited by rubidium ions (PubMed:22183084). Inhibited by lithium ions (PubMed:22183084). kcat is 2.06 sec(-1) with pNP-acetate as substrate (at 37 degrees Celsius and pH 7) (PubMed:22183084). kcat is 4.48 sec(-1) with pNP-butyrate as substrate (at 37 degrees Celsius and pH 7) (PubMed:22183084). kcat is 3.28 sec(-1) with pNP-hexanoate as substrate (at 37 degrees Celsius and pH 7) (PubMed:22183084). kcat is 2.45 sec(-1) with pNP-octanoate as substrate (at 37 degrees Celsius and pH 7) (PubMed:22183084). Optimum pH is 6. Optimum temperature is 50 degrees Celsius. Monomer. Expression is not induced by the synthetic polyester poly(butylene succinate-co-adipate) (PBSA). Shows promising applications in ethyl alcohol distillery processes as it may be utilized for treatment of molasses wastewater (PubMed:33387709). Has potential for application in biological recycling of plastic waste products (PubMed:20393707, PubMed:22183084). Belongs to the AB hydrolase superfamily.