Plant-specific single-stranded DNA-binding protein required for efficient heterologous recombination-dependent DNA repair in nuclear and mitochondrial compartments. Forms large nucleo-protein complexes with WHY2 in mitochondria. Binds ssDNA with high affinity, but with little sequence specificity (PubMed:22762281). Involved in double-stranded DNA break repair (PubMed:22202891). Involved in the hydrolytic splicing pathway in mitochondrion. Facilitates the excision of two cis-spliced group II introns, NAD1 intron 2 and NAD2 intron 1 (PubMed:26048959). Interacts with WHY2. A number of isoforms are produced. According to EST sequences. Expressed in root vascular tissue, tips of primary and secondary roots, young leaves, hydathodes, stomatal guard cells, cauline leaves, flower buds, stipules, carpels, pistils and anther filaments. Reduced fertility, increased sensitivity to mitomycin C, and decreased levels of intrachromosomal recombination. Belongs to the RAD52 family.