Involved in the dorsovental asymmetry of flowers. Promotes dorsal identity. Specifically expressed in the dorsal region of developing flowers. Early expressed at stage two (plastochrons 7 and 8), when the floral meristem comprised a loaf-shaped bulge of cells. Expression was in the dorsal region of the floral meristem. At stage four, when sepal primordia had emerged around the meristem dome, expression could be seen in the dorsal sepal primordium. At stage five, expressed in emerging dorsal primordia of whorls two and three. In older flower buds (stages seven and eight), expression was found mainly in the dorsal petals and staminode. Up-regulated by both CYC and DICH. Radially symmetric flowers. Loss of functional stamens in dorsal positions. Organs reduction with five organs per whorl rather than six.