Required during somitogenesis for the formation of somite boundaries. Represses the expression of genes involved in somite segmentation by acting with the corepressor tle4 to down-regulate the transcriptional activity of tbx6. May act by regulating the activity of tle4. Represses transcription of delta2, thy1 and ripply2.2/bowline itself. Interacts with tle4 and tbx6, and mediates interaction between these proteins. Expressed in the presomitic mesoderm (PSM) in the anterior halves of somitomeres S-I, S-II and S-III. Part of a negative feedback loop during somitogenesis. Expression is activated by tbx6 together with thy1 and tcf3. The ripply homology domain is required for transcriptional repression. The WRPW motif is required for binding to tle/groucho proteins and transcriptional repression. Belongs to the ripply family.