Component of the outer dynein arm-docking complex (ODA-DC) that mediates outer dynein arms (ODA) binding onto the doublet microtubule. Plays an essential role for the assembly of ODA-DC and in the docking of ODA in ciliary axoneme. Component of the outer dynein arm-docking complex along with ODAD1, ODAD2 and ODAD3. Morpholino knockdown of the protein causes severe curvature of the body at 72 hpf. Morphants also exhibit hydrocephalus, abnormal otoliths and disturbed left-right asymmetry patterning. Some animals (17%) tend to develop pronephric cysts. At the 7-somite stage, cilia number and length are reduced in the Kupffer's vesicle. At 60 hpf, cilia motility is markedly impaired in the pronephric duct.