Scaffolding component of the CCR4-NOT complex which is one of the major cellular mRNA deadenylases and is linked to various cellular processes including bulk mRNA degradation, miRNA-mediated repression, translational repression during translational initiation and general transcription regulation (PubMed:23843623, PubMed:28204614). Positively regulates the accumulation of the CCR4-NOT complex component ccr-1 (PubMed:23843623). Within the complex promotes germ cell development and fertility in hermaphrodites (PubMed:23843623). Additional complex functions may be a consequence of its influence on mRNA expression (By similarity). Its scaffolding function implies its interaction with the catalytic complex module and diverse RNA-binding proteins mediating the complex recruitment to selected mRNA 3'UTRs (By similarity). Mediates the recruitment of the CCR4-NOT complex to miRNA targets and to the RISC complex (Probable). Acts as a transcriptional repressor (By similarity). Represses the ligand-dependent transcriptional activation by nuclear receptors (By similarity). In embryos, plays a role in female pronucleus and mitotic spindle positioning during the first cleavage divisions after fertilization (PubMed:16971515). This may partly be through negatively regulating the accumulation of zyg-9 at the centrosome (PubMed:16971515). Negatively regulates the formation of long astral microtubules in developing embryos (PubMed:16971515). Required for the stabilization and degradation of maternal mRNAs such as nos-2 in somatic blastomeres (PubMed:18692039). Component of the CCR4-NOT complex at least composed of ccf-1, ccr-4 and let-711, which is required for germ cell development in hermaphrodites (PubMed:23843623). Within the complex interacts with ccf-1 and ccr-4; the interactions are direct (PubMed:23843623). Highly expressed in the germline of hermaphrodites. Highly expressed in embryos. Contains Leu-Xaa-Xaa-Leu-Leu (LXXLL) motifs, a motif known to be important for the association with nuclear receptors. RNAi-mediated knockdown results in sterility and the production of inviable embryos which are osmotically sensitive, exhibit nuclear rotation defects and irregular positioning of the mitotic spindle which is located towards the posterior or lateral cortex during anaphase (PubMed:23843623, PubMed:16971515). RNAi-mediated knockdown may also result in a reduced brood size and larval lethality (PubMed:23843623). RNAi-mediated knockdown disrupts the arrangement and differentiation of oocytes in the proximal region and as a result small oocyte-like cells arrange in several rows in the germline (PubMed:23843623). RNAi-mediated knockdown reduces the levels of ccf-1, but does not alter ccr-4 levels (PubMed:23843623). RNAi-mediated knockdown results in reduced global poly(A) tail deadenylation (PubMed:23843623). RNAi-mediated knockdown results in reduces degradation of nos-2 mRNA, impairs the recruitment of lsm-1 and decreases the number of patr-1-positive granules in somatic blastomeres (PubMed:18692039). Belongs to the CNOT1 family.