Involved in sister chromatid cohesion (PubMed:10882066). Forms a complex with SCC2, which is required for the association of the cohesin complex with chromosomes (PubMed:10882066, PubMed:26038942). Binds to the nucleosome-free promoter regions of ribosomal protein genes and tRNA genes (PubMed:25173104). Interacts with SCC2 to form the cohesin loading complex. Associates with chromatin. Present with 907 molecules/cell in log phase SD medium. Belongs to the SCC4/mau-2 family.