Exhibits chaperone activity toward chloroplast outer envelope membrane, mitochondrion outer membrane, endoplasmic reticulum membrane and peroxisomal proteins, by recruiting specific proteins containing a single transmembrane associated with an AKR2A-binding sequence (ABS) and subsequently binding glycolipids (e.g. monogalactosyldiacylglycerol (MGDG) and phosphatidylglycerol (PG)) present in the membrane of the target organelle. Binds to chloroplast outer envelope membrane (OEM) protein targeting signals, as well as to chloroplasts. Binds specifically to two chloroplast glycolipids, monogalactosyldiacylglycerol (MGDG) and phosphatidylglycerol (PG) (By similarity). Interacts with APX3, APX5 and TOC34 (PubMed:20215589). The ankyrin repeats (ANK) mediate interactions with hexoses-containing lipids present in organellar membranes (e.g. chloroplast), such as monogalactosyldiacylglycerol (MGDG) and phosphatidylglycerol (PG).