Sequence-specific DNA binding factor that recognizes the 5'-CCAAATTTTAAGTTT-3' sequence. Recruits the FRI-C complex to the FLC promoter. Required for FRI-mediated FLC activation, but has no effect on the expression of MAF1, MAF2, MAF3, MAF5, UFC and CO. Dispensable for the reactivation of FLC in early embryogenesis, but required to maintain high levels of FLC expression in later embryonic and vegetative development. Homodimer. Component of the transcription activator complex FRI-C composed of FRI, FRL1, SUF4, FLX and FES1. Interacts with LD, ASHH2, FRL1, (via C-terminus) with FRI (via C-terminus), and with SWC6, a component of the SWR1 chromatin-remodeling complex. Binds to MED18 to regulate flowering time; recruits MED18 to FLC promoter (PubMed:24451981). Expressed in root, shoot apex, leaves, stem and flowers. Expressed in expanding leaves, in the vasculature of fully expanded leaves, in the inflorescence, throughout young floral primordia, in the carpels of older flowers and in fertilized ovules. Barely detectable in 3-days-old seedling, and then increases. Not regulated by photoperiod or vernalization. Early flowering. Truncated N-terminus.