Involved in dynein arm assembly, is important for expression and transporting outer dynein arm (ODA) proteins from the cytoplasm to the cilia (PubMed:23122589, PubMed:23527195, PubMed:33403504). Acts as a crucial component in the formation and motility of spermatozoal flagella (PubMed:33403504). Interacts (via CS domain) with ZMYND10 (via C-terminus). Expressed predominantly in testis and in nasal epithelial cells. Localized in the cytoplasm of early spermatogonia and spermatids of different spermatogenic stages and in the flagella of mature epididymal spermatozoa. The disease is caused by variants affecting the gene represented in this entry. Belongs to the tilB family.