Involved in the assembly of a N-linked pentasaccharide that decorates the S-layer glycoprotein and flagellins. Probably mediates or contributes to the translocation of the dolichol-phosphate-mannose across the membrane. Cell surface structure biogenesis; S-layer biogenesis. Mutants show accumulation of both tetrasaccharide- and mannose-linked dolichol phosphate. The final mannose residue of the N-linked glycan decorating the S-layer glycoprotein is not added. Belongs to the AglR/Agl15 family.