Plays a role in innate immunity against the oomycete pathogen A.arabidopsidis (Hpa). Interacts with SUN1 and SUN2. Expressed in epidermal cells, mesophyll cells, trichomes and root cells. The KASH domain, which contains a transmembrane domain, mediates the nuclear envelope targeting and is involved in the binding to the SUN proteins. No visible phenotype. More susceptible to the oomycete pathogen H.arabidopsidis (Hpa) infection.