Regulates negatively the cuticle development by interacting with the HD-ZIP IV transcription factor HDG1. Interacts with BHLH122/CFLAP1 and BHLH80/CFLAP2 (PubMed:26745719). Binds to HDG1 (PubMed:21954461). Mostly observed in roots, flowers and siliques (PubMed:21954461). Expressed in cells differentiated from epidermal cells such as trichomes, stigmatic papillar cells and guard cells, as well as in tissues undergoing abscission and dehiscence (PubMed:21954461). Expressed in seedlings and all organs of adult plants (PubMed:21954461). In roots, present in endodermis and central cylinder (PubMed:21954461). In flowers, detected in stigmatic papillar cells (PubMed:21954461). When petals and sepals are withering, accumulates in the abscission zone at the bottom of the silique (PubMed:21954461). Later observed along the valve margin-replum boundary, where dehiscence and pod shatter occur (PubMed:21954461). Increased cuticle development leading to ectopic cuticle biosynthesis in trichomes. Truncated N-terminus.