Regulates translation as part of a complex with DENR. Specifically required for translational re-initiation in mRNAs containing upstream open reading frames (uORFs). Not required for standard translational initiation. Regulates expression of a subset of gene products including mbc, InR and EcR. Interacts with DENR. RNAi-mediated knockdown results in pupal lethality. The abdominal epidermis is soft and transparent whereas the head and thorax appear normal. Belongs to the MCTS1 family.