Probable acetyltransferase required for the establishment of sister chromatid cohesion and couple the processes of cohesion and DNA replication to ensure that only sister chromatids become paired together. In contrast to the structural cohesins, the deposition and establishment factors are required only during S phase. The relevance of acetyltransferase function remains unclear (By similarity). Interacts with pds5. In the C-terminal section; belongs to the acetyltransferase family. ECO subfamily. In the N-terminal section; belongs to the DNA polymerase type-Y family.