Protein kinase that may be involved in microsporogenesis. ATP + L-seryl-[protein] = ADP + H(+) + O-phospho-L-seryl-[protein] ATP + L-threonyl-[protein] = ADP + H(+) + O-phospho-L-threonyl-[protein] Activated by calcium/calmodulin binding after calcium-induced autophosphorylation. Autophosphorylation is associated with a time-dependent loss of kinase activity sensitive to reaction pH and ATP concentration. In vitro inactivation leads to the formation of network-like structures. Expressed during anther development. Autophosphorylation stimulated by calcium and inhibited by calcium/calmodulin. Occurs probably by an intermolecular mechanism. Belongs to the protein kinase superfamily. CAMK Ser/Thr protein kinase family. CaMK subfamily.