Involved in the pathway that organizes the shaping and sizing of the prospore membrane (PSM) during sporulation. May be required for the formation of ADY3 and DON1-containing protein coats at the leading edge of the PSMs during meiosis II. Interacts directly with ADY3. Probable component of a spindle pole body (SPB) complex composed of ADY3, SSP1, DON1, MPC54, SPO21/MPC70, NUD1 and CNM67. Localizes to precursors of the PSM and to the leading edge, which cover the ring-shape opening of the PSMs during meiosis II. Some fraction colocalizes with DON1. Meiosis-specific. Expressed from 3 to 9 hours after induction of sporulation. Not expressed during mitosis. Phosphorylated.