Required for the post-translational delivery of tail-anchored (TA) proteins to the endoplasmic reticulum. Acts as a membrane receptor for soluble GET3, which recognizes and selectively binds the transmembrane domain of TA proteins in the cytosol (By similarity). Involved in the control of root hair growth through the regulation of syntaxin SYP123 expression (PubMed:28096354). Forms homodimers (PubMed:28096354). Interacts with GET3A (PubMed:28096354). Strong reduction of root hair length. Belongs to the WRB/GET1 family.