Plays a role in the process of sperm capacitation and acrosome reactions (PubMed:9322250, PubMed:9820206). Probable receptor for the putative fertilization-promoting peptide (FPP) at the sperm membrane that may modulate the activity of the adenylyl cyclase cAMP pathway (PubMed:9322250, PubMed:9820206). Found in a complex at least composed of MROH2B isoform 2, PRKACA isoform 2 and TCP11 (PubMed:27105888). Interacts with MROH2B isoform 2 (PubMed:27105888). Interacts with PRKACA isoform 2 (PubMed:27105888). Interacts with ODF1 (via leucine zipper motif) (By similarity). Localizes on the acrosomal cap region of acrosome-intact, but not acrosome-reacted sperm (PubMed:9322250). Colocalizes with MROH2B and PRKACA on the acrosome and tail regions in round spermatids and spermatozoa regardless of the capacitation status of the sperm (PubMed:27105888). Testis-specific (PubMed:1893875). Expressed in mature epididymal spermatozoa (at protein level) (PubMed:9322250). First expressed in the pachytene spermatocyte stage (PubMed:1893875). Constitutively phosphorylated on serine, threonine and tyrosine residues within the head and tail regions of noncapacitated spermatozoa (PubMed:27105888). Phosphorylation on tyrosine residues increases upon sperm capacitation within the acrosomal region in a protein kinase A (PKA)-dependent signaling pathway (PubMed:27105888). Belongs to the TCP11 family.