Transcriptional activator that binds specifically to the DNA sequence 5'-[AG]CCGAC-3' (PubMed:11798174). Binding to the C-repeat/DRE element mediates high salinity- and dehydration-inducible transcription (PubMed:11798174). Promotes the expression of heat stress-inducible genes by contributing to the formation of a heat stress-specific transcriptional complex with NF-Y subunits (e.g. DPB3-1, NF-YA2 and NF-YB3) at the promoter of target genes, thus promoting heat tolerance (PubMed:25490919). Interacts with MED25 (PubMed:18552202, PubMed:21536906). Binds to DPB3-1 in the nucleus during heat-stress (PubMed:25490919). A number of isoforms are produced. According to EST sequences. Expressed preferentially in roots and stems, and at a lower level in leaves. By high-salt and drought stresses. Ubiquitinated by DRIP1 and DRIP2. Ubiquitination probably leads to its subsequent degradation, thus negatively regulating response to drought. Belongs to the AP2/ERF transcription factor family. ERF subfamily.