Plays a pivotal role in the control of morphogenesis of shoot organs by negatively regulating the expression of boundary-specific genes such as CUC genes, probably through the induction of miRNA (e.g. miR164). Binds to the 3'-ACC-5' repeats in the light-responsive promoter (LRP) of psbD, and activates its transcription. Participates in ovule develpment (PubMed:25378179). Interacts with AHL27 and AHL29 (PubMed:24218605). Interacts with SPL (PubMed:25527103, PubMed:25378179). Interacts with KIN10; KIN11 and FLZ3 (Ref.11). Expressed in cotyledons, particularly in the vascular region, in leaves, buds, flowers and immature siliques, and, to a lower extent, in roots. Expressed during ovule development (PubMed:25378179).