Hydroperoxy fatty acid reductase essential for the removal of lipid hydroperoxides under normal and stress conditions, leading to the protection of membrane integrity. a hydroperoxy polyunsaturated fatty acid + H(+) + NADPH = a hydroxy polyunsaturated fatty acid + H2O + NADP(+) Mercaptosuccinate, pCMB, and nethylmaleimide act as inhibitors of the catalytic activity. Optimum pH is 8.2. Optimum temperature is 37 degrees Celsius. Monomer. High light, methylviologen, and salt stress conditions increase the expression level. Belongs to the glutathione peroxidase family.