Accessory component of the DNA polymerase epsilon complex (By similarity). Participates in DNA repair and in chromosomal DNA replication (By similarity). Has a role in the entrance and progression through S phase (PubMed:24224125). Has a role in endoreplication (PubMed:24224125). Essential for viability and tissue development (PubMed:24224125). Component of the epsilon DNA polymerase complex consisting of four subunits: the catalytic subunit PolE1/DNApol-epsilon255 and the accessory subunits PolE2/DNApol-epsilon58, Chrac-14/DNApolE3 and PolE4. Expressed in the embryo and larva (at protein level). Pupal lethal (PubMed:24224125). Results in smaller eye, leg and wing disk, brain lobe and salivary glands (PubMed:24224125). Results in defective entry and/or progression though S phase during the cell cycle in eye imaginal disk cells (PubMed:24224125). Results in defective endoreplication in salivary glands (PubMed:24224125). Belongs to the DNA polymerase epsilon subunit B family. Truncated N-terminus. Extended N-terminus. Truncated N-terminus.