Involved in transport of phospholipids and in regulation of pollen plasma membrane lipid asymmetry. ATP + H2O + phospholipidSide 1 = ADP + phosphate + phospholipidSide 2. Ala6 and ala7 double mutants are hypersensitive to temperature stress and are impaired in pollen fitness with an altered lipid composition and short and slow pollen tubes. Belongs to the cation transport ATPase (P-type) (TC 3.A.3) family. Type IV subfamily.