Involved in the negative regulation of dendrite outgrowth. Homotetramer, and heterotetramer with other DPYS-like proteins. Interacts with DPYSL2, DPYSL3 and DPYSL4. Interacts with SEPTIN4 isoform 4. Interacts with MAP2 and TUBB3 (By similarity). Translocates into the mitochondria upon interaction with SEPTIN4 isoform 4. Detected in brain. Detected in whole embryos after 11 days of development. Detected in embryonic head. Highly expressed in newborns and up to 7 days after birth. Expression is decreased after 14 days. Belongs to the metallo-dependent hydrolases superfamily. Hydantoinase/dihydropyrimidinase family. Lacks most of the conserved residues that are essential for binding the metal cofactor and hence for dihydropyrimidinase activity. Its enzyme activity is therefore unsure.