Polo kinase inhibitor required to maintain G2 arrest in the meiotic cell cycle in females (PubMed:18052611). Holds heterochromatically paired homologs together from the end of pachytene until metaphase I (PubMed:14573476). Haploinsufficient locus for homologous achiasmate segregation and may be required for the maintenance of heterochromatic pairings (PubMed:14573476). Interacts with polo (PubMed:18052611). Interacts with cort (PubMed:24019759). Expressed from the end of pachytene until the completion of meiosis I. Probably ubiquitinated: degraded during the oocyte-to-embryo transition by the anaphase promoting complex/cyclosome (APC/C) containing cort protein. Mutants exhibit a strong dominant effect on achiasmate segregation, inducing both X and fourth chromosome non-disjunction in females (PubMed:14573476). Precocious nuclear envelope breakdown in oocytes, due to defects in meiotic arrest in G2 (PubMed:18052611).