Binds to NGFI-B response elements. Plays an important role in late stages of epidermal metamorphosis. Forms a heterodimer with USP. Ubiquitously expressed in preblastoderm embryos, specifically in central nervous system and intestinal tract. Highly expressed in third instar larval imaginal disks and brain complexes, but not in ovaries. Low levels in 0-8 hours embryos and adults. Higher in late embryogenesis and during larval and pupal stages. Short isoform is enriched in pupae and adults, long isoform in larvae. Belongs to the nuclear hormone receptor family. NR4 subfamily.