Participates in the later stages of centriole assembly through the interaction with Ana3 leading to the centriole to centrosome conversion in somatic cells. Interacts with Ana3; this complex is recruited to daughter centrioles before their conversion to centrosomes. Localizes to the more distal part of the lumen of the centriole. Recruited into zone I during interphase after the formation of the cartwheel at the end of mitosis (PubMed:32543652). The Rcd4 recruitment to the daughter centriole is dependent on Ana3 (PubMed:32543652). Homozygous flies for rcd4 are lethal at the late pupal and adult stages. The flies that emerged are slow and uncoordinated. Female mutant flies are sterile. Male mutants are fertile. Belongs to the PPP1R35 family.