Involved in ribosome biogenesis. Required for normal pre-rRNA processing in internal transcribed spacer 1 (ITS1). May be involved in the movements of the replication forks. Interacts with the 35S, 23S and 20S pre-rRNAs and with the U3 snoRNA. Transiently up-regulated during the initial 15 minutes of dough-fermentation. Present with 3670 molecules/cell in log phase SD medium. Belongs to the SLX9 family.