DNA polymerase with primer-independent templated DNA polymerization activity, primer-dependent DNA polymerization activity with strand displacement, translesion synthesis activity across non-bulky base damage, 3'-5' exodeoxyribonuclease activity, and de novo primer synthesis activity. The enzyme is processive and faithful. Translation synthesis across abasic sites is coupled to de novo primer synthesis. Overexpression of wild-type protein increases survival of cells upon mitomycin C or UV treatment. a 2'-deoxyribonucleoside 5'-triphosphate + DNA(n) = diphosphate + DNA(n+1) Translesion synthesis across abasic sites is more efficient with Mn(2+) compared to Mg(2+), replication of undamaged template is more efficient with Mg(2+). Encoded in a mobile genetic element called pipolin.