Calcium-regulated, actin-modulating protein that binds to the plus (or barbed) ends of actin monomers or filaments, preventing monomer exchange (end-blocking or capping). It can promote the assembly of monomers into filaments (nucleation) as well as sever filaments already formed. Binds to actin and to fibronectin. Isoform 1 and isoform 2 are ubiquitously expressed in early embryo. Isoform 1 is expressed in the fat body, and is abundant in hemolymph. Isoform 2 is expressed in parts of the gut. Belongs to the villin/gelsolin family. Lacks one of the cysteines to make the disulfide bridge in isoform 1. It is replaced by Val-233. Truncated N-terminus. Truncated N-terminus. Intron retention. Extended N-terminus. Truncated N-terminus.