Transcriptional coactivator; part of the gene cluster that mediates the biosynthesis of trypacidin, a mycotoxin with antiprotozoal activity and that plays a role in the infection process (PubMed:26278536, PubMed:26242966). With tpcE, coregulates the production of trypacidin (PubMed:26278536, PubMed:26242966). Specifically expressed in conidia (PubMed:22319557). Expression is positively regulated by the transcription factors brlA and laeA (PubMed:26242966). Resulted in loss of trypacidin production (PubMed:26278536, PubMed:26242966).