Component of the peroxisomal protein import machinery. Together with PEX6, mediates the ATP-dependent relocation and recycling of the peroxisomal targeting signal-1 (PTS1) import receptor PEX5 from the peroxisomal membrane to the cytosol, where it is then available for another round of protein import into the organelle. Interacts with PEX6; forms a high-molecular-mass heterooligomer in the cytosol. The PEX1-PEX6 heterooligomers associate with the peroxisomal importomer via interaction of PEX6 with the peroxisomal membrane anchor PEX15. Shuttles between the cytoplasm and the peroxisomal membrane. By oleate. AAA-cassette D1 is required for interaction with PEX6. ATP-binding in AAA-cassette D2 is required for interaction with PEX6. ATP-binding and hydrolysis in AAA-cassette D2 is required for proper function in PEX5 dislocation. Present with 2100 molecules/cell in log phase SD medium. Belongs to the AAA ATPase family.