Scaffold protein that may play a role in cell adhesion, cell spreading and in the reorganization of the actin cytoskeleton. May inhibit cell growth (By similarity). Regulates cranial neural crest migration. Acts together with prickle1 to control axial elongation. Detected along actin stress fibers. Expressed in the animal hemisphere at the 4-cell stage. By stage 18, expressed in cells adjacent to the anterior neural plate. In late neurula, expressed in the cranial neural crest. At tail bud stages, expressed strongly in the head, ventral to the developing eye, branchial arches and lateral line placodes. Also localized in the otic vesicle, dorsal fin and notochord with weaker expression at intersomitic junctions of tail bud embryos. Expressed maternally. Expression levels decline during the cleavage stages with only weak expression by gastrula stages. Levels then increase at neurula stages and are high in tail bud embryos. The N-terminal and the C-terminal halves of the protein can associate with each other, thereby hindering interactions with other proteins. Belongs to the prickle / espinas / testin family.