Component of the monoterpenoid pyrethrins biosynthesis; pyrethrins are widely used plant-derived pesticide (PubMed:30468448). Acyltransferase that catalyzes the esterification of terpene acids and lipid alcohol substrates into pyrethrins; mediates the transfer of a chrysanthemoyl moiety from the coenzyme A (CoA) thio-ester chrysanthemoyl CoA to pyrethrolone, and, to a lower extent, to jasmololone and cinerolone thus producing pyrethrins (e.g. pyrethrin type I) (PubMed:22385412). Can also use pyrethroyl CoA as substrate (PubMed:22385412). Has also esterase activity, being able to cleave the ester bond of pyrethrin I, p-nitrophenyl butanoate and p-nitrophenyl octanoate to produce pyrethrolone and p-nitrophenol, respectively (PubMed:22385412). (1R,3R)-chrysanthemoyl-CoA + (Z,S)-pyrethrolone = CoA + pyrethrin I (1R,3R)-pyrethroyl-CoA + (Z,S)-pyrethrolone = CoA + pyrethrin II (1R,3R)-chrysanthemoyl-CoA + (Z,S)-jasmololone = CoA + jasmolin I (1R,3R)-chrysanthemoyl-CoA + (Z,S)-cinerolone = cinerin I + CoA (1R,3R)-pyrethroyl-CoA + (Z,S)-jasmololone = CoA + jasmolin II (1R,3R)-pyrethroyl-CoA + (Z,S)-cinerolone = cinerin II + CoA Optimum pH is 7.5. Optimum temperature is 25 degrees Celsius. Isoprenoid biosynthesis. Restricted to the pericarp during achene maturation (PubMed:23104830). Expressed in the leaves of mature plants and seedlings, as well as in buds and flowers (PubMed:22385412, PubMed:29122986). Present in disk florets (PubMed:29122986). Mostly expressed in ovaries of flowers at stages S1 to S7 (bud to overblown), before the first disk florets opening, and prior embryos formation. By wounding. Belongs to the 'GDSL' lipolytic enzyme family.