DNA polymerase II, which participates in chromosomal DNA replication. Required for the timing and determination of cell fate during plant embryogenesis and root pole development, by promoting cell cycle and cell type patterning. Necessary for proper shoot (SAM) and root apical meristem (RAM) functions. Involved in maintaining epigenetic states, controlling hypersensitive response (HR), and mediating abscisic acid (ABA) signaling. Required for flowering repression through a mechanism involving epigenetic gene silencing. May participate in processes involved in chromatin-mediated cellular memory. a 2'-deoxyribonucleoside 5'-triphosphate + DNA(n) = diphosphate + DNA(n+1) Binds 1 [4Fe-4S] cluster. Heterotetramer (By similarity). Subunit of the DNA polymerase II (PubMed:16212602, PubMed:19947980). Interacts (via C-terminus) with DPB2 (PubMed:16212602). Interacts with LHP1/TFL2 (PubMed:19947980). Mostly expressed at low levels in inflorescence (floral meristem and flowers until anthesis), and, to a lower extent, in roots, seeds and leaves. Present in actively dividing cells such as root and shoot meristematic regions, young leaves and stems, inflorescences and siliques. Follows a cell-cycle-dependent expression with a maximal induction in the S phase and another induction at the G2/M transition. The DNA polymerase activity domain resides in the N-terminal half of the protein, while the C-terminus is necessary for maintenance of the complex. The CysA-type zinc finger is required for PCNA-binding. The CysB motif binds 1 4Fe-4S cluster and is required for the formation of polymerase complexes. Lethal, with sporophytic embryo-defective with an arrest at the globular stage during embryo development. Abnormal cell division characterized by several rounds of mitosis with aberrant planes of division. Belongs to the DNA polymerase type-B family.