Essential protein required during embryogenesis (PubMed:15266054). Component of ribonuclease P, a protein complex that generates mature tRNA molecules by cleaving their 5'-ends (By similarity). Also a component of RNase MRP (By similarity). Component of nuclear RNase P and RNase MRP ribonucleoproteins (By similarity). Interacts with GAF1/RPP30 (PubMed:22509260). Defective embryo arrested at the globular stage. Belongs to the eukaryotic/archaeal RNase P protein component 2 family. The predicted gene At1g04640 has been split into 2 genes: At1g04635 and At1g04640.