Probable floral-specific cell division component, required for proper organ formation in flowers. Regulates the floral meristem cell division and the inflorescence meristem organization. Plays a role in development of both male and female reproductive tissues. Ubiquitous but expressed mostly in flowers with highest levels in developing ovules and microspores. Expressed at the first stage of pollen development in uninucleate microspores and bicellular pollen but not in tricellular and mature pollen. The cysteine-rich domain CRC binds zinc in vitro. Despite a wild-type flower phenotype, the null mutant presents defects in pollen, carpel and ovule development, including a failure in megasporogenesis. Small siliques. Belongs to the lin-54 family. Truncated N-terminus.