Component of the Mediator complex, a coactivator involved in the regulated transcription of nearly all RNA polymerase II-dependent genes (By similarity). Mediator functions as a bridge to convey information from gene-specific regulatory proteins to the basal RNA polymerase II transcription machinery (By similarity). Mediator is recruited to promoters by direct interactions with regulatory proteins and serves as a scaffold for the assembly of a functional pre-initiation complex with RNA polymerase II and the general transcription factors (By similarity). Involved in development of gonad and seam cells (PubMed:19500563, PubMed:15020414). Plays a role in temporal regulation of seam cell division, probably acting as a part of the Mediator complex (PubMed:19500563). Component of the Mediator complex (By similarity). May interact with sel-7 and sel-8 (PubMed:15020414). RNAi-mediated knockdown has no effect on seam cell division or differentiation, but in a sel-7 mutant background decreases number of seam cells (PubMed:19500563). Partially suppresses the hermaphrodite gonadal development defect in a lin-12 mutant background (PubMed:15020414). Belongs to the Mediator complex subunit 29 family.