Component of the MIS complex, a complex that mediates N6-methyladenosine (m6A) methylation on some mRNAs during meiosis and is required for sporulation. Component of the MIS complex, composed of SLZ1, MUM2 and IME4. Localizes in the nucleolus during the period of m6A mRNA accumulation. Mutant is sporulation defective and fails to perform premeiotic DNA synthesis. Present with 3400 molecules/cell in log phase SD medium. Belongs to the fl(2)d family.