Ferric chelate reductase involved in iron mobilization from the cytosol into the chloroplast. May participate in the transport of electrons to a Fe(3+) ion via FAD and heme intermediates. Might be involved iron homeostasis in trichomes. 2 a Fe(II)-siderophore + H(+) + NAD(+) = 2 a Fe(III)-siderophore + NADH Expressed in shoots, flowers and siliques. Detected in cotyledons, leaves and trichomes, but not in roots. Expressed during cell or organ differentiation. Up-regulated in shoots by iron deficiency and down-regulated in shoots by copper deficiency. No visible phenotype when grown under normal conditions. Reduced photosynthetic electron transport efficiency, chloroplast ferric chelate reductase activity and iron content. Seedling lethal when grown in alkaline soil. Belongs to the ferric reductase (FRE) family.