Required during cellulose deposition (PubMed:20388664). May act as a bridging protein that binds pectin and other cell wall polysaccharides. Probably involved in maintaining esterification of pectins (Probable). May be involved in the specific O-acetylation of cell wall polymers (By similarity). Expressed in leaf vasculature, growing part of the root, expanding inflorescence stems and trichomes. Contains 2 motifs that are conserved in esterases, but it is unlikely that this protein belongs to the catalytically active pectin esterases (PubMed:20657172). Tbr mutants are lacking leaf and stem trichome birefringence characteristic of plant cells that contain highly ordered cellulose in their secondary walls (PubMed:20388664). Belongs to the PC-esterase family. TBL subfamily.