Splicing factor involved in pre-mRNA splicing and component of the spliceosome (PubMed:18702672, PubMed:19168457). Essential for reproduction. In female gametophyte, is necessary for the egg cell and central cell fate determination and hence reproductive success. Involved in a mechanism that prevents accessory cells from adopting gametic cell fate. Is necessary to restrict LIS expression to interfere with egg-cell specification (PubMed:18702672). Probable component of U5 small nuclear ribonucleoprotein (snRNP) that is required for pre-mRNA splicing. Plays an essential role in female gametogenesis and embryo development (PubMed:19261069). Required for the control of polarized cell growth and cell proliferation during floral organ morphogenesis (PubMed:19168457). Interacts with BRR2A and PRP8A. Colocalizes with CLO (PubMed:18702672). Colocalizes with SC35 (PubMed:19168457). Expressed in flower buds, open flowers and siliques. Expressed at low levels in rosettes leaves, cauline leaves and stems. Expressed in developing embryos until heart stage. Gametophytic lethality due to female gametophyte defect, when homozygous. This protein was named 'VAJRA', because the shape of flower buds when ready to open resemble VAJRA, an instrument used in esoteric Buddhism. Belongs to the TRAFAC class translation factor GTPase superfamily. Classic translation factor GTPase family.