Exhibits lactonase activity. Acts in cells with perturbed membrane integrity and is possibly related to the membrane homeostasis. Contributes to bacitracin resistance (By similarity). Binds 2 Ca(2+) ions per subunit. Induced by cell wall-affecting antibiotics such as oxacillin, vancomycin, fosfomycin, and bacitracin. Belongs to the SMP-30/CGR1 family.