D-arabinono-1,4-lactone oxidase that catalyzes the final step of biosynthesis of D-erythroascorbic acid, an important antioxidant and one of the virulence factors enhancing the pathogenicity (PubMed:11349062, PubMed:7957197). Is also able to oxidize L-galactono-1,4-lactone, L-xylono-1,4-lactone and L-gulono-1,4-lactone (PubMed:7957197). D-arabinono-1,4-lactone + O2 = dehydro-D-arabinono-1,4-lactone + H(+) + H2O2 L-galactono-1,4-lactone + O2 = H(+) + H2O2 + L-ascorbate L-gulono-1,4-lactone + O2 = H(+) + H2O2 + L-ascorbate L-xylono-1,4-lactone + O2 = dehydro-L-arabinono-1,4-lactone + H(+) + H2O2 Cofactor biosynthesis; D-erythroascorbate biosynthesis; dehydro-D-arabinono-1,4-lactone from D-arabinose: step 2/2. Abolishes D-arabinono-1,4-lactone oxidase activity and impairs the production of D-erythroascorbic acid (PubMed:11349062). Leads to increased sensitivity towards oxidative stress, defective hyphal growth and attenuated virulence (PubMed:11349062). Belongs to the oxygen-dependent FAD-linked oxidoreductase family.