Involved in cell division and chromosome segregation (PubMed:24065632). Plays a central role in the initiation of sporulation through the activation of multiple components of the cell division machinery (PubMed:24065632). Acts as a transcriptional regulator that is involved in the direct control of genes required for key steps in sporulation, including the cessation of aerial growth, the initiation of cell division, and chromosome segregation. Functions almost equally as an activator and as a repressor (PubMed:24065632). Required for DNA binding by WhiB. May function cooperatively with WhiB (PubMed:27094333). DNA binding depends on WhiB. Autoregulated. Expressed throughout vegetative growth and sporulation. Could be post-translationally regulated. Mutants form unusually long aerial hyphae that fail to initiate septation or chromosome segregation. Belongs to the WhiA family.