E2-like enzyme required for the cytoplasm to vacuole transport (Cvt), autophagy and nucleophagy (By similarity). Acts as an E2-like enzyme that catalyzes the conjugation of ATG12 to ATG5 (By similarity). ATG12 conjugation to ATG5 is required for proper localization of ATG8 to the preautophagosomal structure (PAS) (By similarity). Likely serves as an ATG5-recognition molecule (By similarity). Autophagy is required for proper vegetative growth, asexual/sexual reproduction, and full virulence (PubMed:28894236). Autophagy is particularly involved in the biosynthesis of deoxynivalenol (DON), an important virulence determinant (PubMed:28894236). Forms homooligomers (By similarity). Interacts with ATG7 and ATG12 (By similarity). Significantly decreases the radial growth of colonies under nutrient-rich conditions (PubMed:28894236). Belongs to the ATG10 family.