Involved in regulation of intracellular signaling pathways during development. Specifically thought to play a role in canonical and/or non-canonical Wnt signaling pathways through interaction with DSH (Dishevelled) family proteins. Positive regulator of the Wnt signaling pathway which acts downstream of wnt1 indicative for non-canonical Wnt signaling. Also negatively regulates the Nodal signaling pathway, possibly by promoting the lysosomal degradation of Nodal receptors. Required for convergent extension movements in gastrulation. Interacts with dvl2. Predominantly cytoplasmic. Also found in late endosomes, the nucleus and at the cell membrane. Expressed maternally. Restricted to the dorsal blastoderm margin in early gastrula and the future dorsal side of the embryo at the sphere stage. At the shield stage, expressed throughout the shield and in lateral areas. Expressed in the anterior brain and axial mesoderm at the tailbud stage, and subsequently maintained in the lateral mesoderm. During somitogenesis, up-regulated in the anterior of the older somites, and expressed in the posterior presomitic mesoderm and tailbud. The C-terminal PDZ-binding motif may mediate interaction with the PDZ domains of DSH (Dishevelled) family proteins. Belongs to the dapper family.