Required for DNA double-strand breaks (DSBs) formation during meiotic recombination (PubMed:31003867, PubMed:31000436). Regulates the spatial and temporal patterns of pre-DSB recombinosome assembly and recombination activity by acting as a scaffold that anchors REC114 and other factors to specific genomic locations, thereby regulating DSB formation (PubMed:31003867, PubMed:31000436). Plays a key role in recombination in the pseudoautosomal regions of sex chromosomes (PubMed:31000436). Interacts with REC114; the interaction is direct (PubMed:31003867, PubMed:31000436). Interacts with IHO1 (PubMed:31000436). Localizes on chromatin between preleptotene and early pachytene (PubMed:31000436). Associates with the chromosome axes, but disappears from axes upon synaptonemal complex formation (PubMed:31000436). Present in meiotic cells (at protein level). Fertility defects (PubMed:31003867, PubMed:31000436). Although female mice are fertile, they display fertility defects faster than wild type in advanced age (PubMed:31000436). Male mice are infertile (PubMed:31003867, PubMed:31000436). Defects are caused by delayed recombination initiation (PubMed:31000436). DNA double-strand breaks (DSBs) distribution during meiosis is altered because of reduced selectivity for sites that normally attract DSBs (PubMed:31000436). Spermatocytes have altered DSB locations and fail to target DSBs to the pseudoautosomal regions of sex chromosomes (PubMed:31003867, PubMed:31000436).