Transcription factor of the NAC family associated with the grain protein content (GPC). Accelerates senescence and increases nutrient remobilization from leaves to developing grains. The tetraploid cultivated wheat (T.durum) and the hexaploid wheat (T.aestivum) carry a one bp insertion generating a non-functional allele of this gene. They have a delayed senescence and a lower grain protein, zinc and iron content (PubMed:17124321). The NAC domain includes a DNA-binding domain and a dimerization domain. High grain protein content (GPC) is a desirable trait in breadmaking and pasta wheat varieties because of its positive effects on quality and nutritional value while Low GPC is a desirable trait for barley malt and beer production.