Required to maintain the prospore membrane to the nucleus during sporulation in order to capture the daughter nuclei and form the spores. Mediates the fusion of exocytic vesicles with the plasma membrane during sporulation through its interactions with the t-SNARE SSO1 and v-SNARE SNC2. Interacts with the t-SNARE SSO1 and the v-SNARE SNC2. Membrane-associated to the plasma during vegetative growth and prospore membrane associated during sporulation. The inhibitory region sequesters the protein in the nucleus. The positive regulatory region is essential and contains an amphipathic helix with hydrophobic and positive charged faces involved in the localization of the protein to the membranes through its binding to phospholipids. Belongs to the SNAP-25 family.