Controls cell shape changes in the neuroepithelium during neural tube closure. Induces apical constriction in epithelial cells by promoting the apical accumulation of F-actin and myosin II, and probably by bundling stress fibers (By similarity). Induces apicobasal cell elongation by redistributing gamma-tubulin and directing the assembly of robust apicobasal microtubule arrays (By similarity). Interacts with F-actin (By similarity). Interacts with ROCK1 (PubMed:22493320). Colocalizes with F-actin in stress fibers and adherens junctions. The ASD1 domain mediates F-actin binding. The ASD2 domain mediates the interaction with ROCK1 and is required for apical constriction induction. Belongs to the shroom family. It is uncertain whether Met-1 or Met-2 is the initiator. Met-2 is more conserved than Met-1 among the orthologs. Truncated N-terminus. Truncated N-terminus.