Component of the sarcomere that tethers together nebulin (skeletal muscle) and nebulette (cardiac muscle) to alpha-actinin, at the Z lines. Interacts with TTN/titin, NEB, NEBL, ACTN2 and CARP. Bound to sarcomere both at the Z-line periphery and in the central I-band region. Belongs to the myotilin/palladin family.