Small GTPase required for proper localization of RNA polymerase II and III (RNAPII and RNAPIII). May act at an RNAP assembly step prior to nuclear import (PubMed:23267056). Required for establishment of sister chromatid cohesion (PubMed:18439903). Heterodimers with NPA3/GPN1 or GPN3 (PubMed:21844196, PubMed:23324351). Binds to RNA polymerase II (RNAPII) (By similarity). Present with 9230 molecules/cell in log phase SD medium. Belongs to the GPN-loop GTPase family.