Acts as a positive regulator of hedgehog signaling and regulates ciliary function (PubMed:29290584). Found in a complex with TEDC1, TEDC2, TUBE1 and TUBD1. Associated with centrioles (PubMed:10753753). Both cytoplasmic and nuclear (PubMed:10753753). In the elongating spermatid it is associated with the manchette, a specialized microtubule system present during reshaping of the sperm head (PubMed:10753753). Highly expressed in testis. Belongs to the tubulin family.