Involved in repair of UV radiation-induced DNA damage. Catalyzes the photoreactivation of pyrimidine [6-4] pyrimidone photoproduct (6-4 products). Binds specifically to DNA containing 6-4 products and repairs these lesions in a visible light-dependent manner. Not required for repair of cyclobutane pyrimidine dimer (CPD). (6-4) photoproduct (in DNA) = 2 pyrimidine residues (in DNA). Binds 1 FAD per subunit. Expressed in siliques, flowers and leaves. Not detected in roots. Expressed at all developmental stages. Not induced by white or UV-B light. Belongs to the DNA photolyase class-1 family. Truncated N-terminus. Truncated N-terminus.