Transcriptional regulator required for Agrobacterium-mediated stable genetic transformation by T-DNA integration in host genome, but not for T-DNA transient expression. Binds to VIP1 (By similarity). Interacts with Agrobacterium tumefaciens VirE2. Forms a complex made of Agrobacterium VirE2, VIP1, VIP2 and single-stranded DNA (ssDNA). Upon infection with a T-DNA transfer-competent Agrobacterium strain. Impaired stable genetic transformation by Agrobacterium T-DNA. Belongs to the CNOT2/3/5 family.