Transcription factor required for terminalia development. Negative regulator of the JAK/STAT pathway: represses JAK/STAT-dependent expression of ventral veins lacking (vvl) in the posterior spiracles. Expressed from stage 5 in two rather faint stripes at positions of 64% (anterior domain; AD) and 17% (posterior domain; PD) egg length. During early gastrulation, at stage 6, these two stripes become more evident and detectable at the region posterior to the cephalic furrow and in the hindgut primordium. The AD disappears as gastrulation proceeds, while the PD remains. At stage 15, the AD appears again in the foregut, and PD expression in the hindgut and anal pad. In imaginal disks, it is ubiquitously expressed in both males and females in genital and eye-antennal disks. Not expressed in the brain. In genital disks, it is expressed along the margin of the anterior bulbus in males, while in females it is expressed in the posterior compartment along the anterior-posterior border, with medial expansion in the most posterior region. Expressed both maternally and zygotically. Highly expressed in embryos and pupae, and at lower level in larvae. In adults, it is expressed at higher level in females. Flies show low mating success and reduced copulation duration. Men and female genitalia often remain inside the body, and genitalia and analia are missing in some homozygous flies. These flies are named after the famous dolls who also lack these 'features'.