Preferentially hydrolyzes para-nitrophenyl-beta-D-galactoside. Can hydrolyze para-nitrophenyl-beta-D-fucoside with 5 time less efficiency. Hydrolysis of terminal non-reducing beta-D-galactose residues in beta-D-galactosides. Measured at pH 4.5 and 37 degrees Celsius for all experiments. Ubiquitous, with higher expression levels in roots and siliques. By sugar starvation and exposure to darkness for 24 hours. Belongs to the glycosyl hydrolase 35 family.