Involved in the biosynthesis of vindoline, a precursor of vinblastine and vincristine. 4-O-deacetylvindoline + acetyl-CoA = CoA + vindoline Optimum pH is 7.5-9.0. Alkaloid biosynthesis; vindoline biosynthesis. Monomer. Predominantly expressed in young leaves of mature plants. Low expression in stems and flowers and not detected in roots. Confined to the laticifer and idioblast cells of leaves, stems, and flower buds (PubMed:11154328). Inhibited by tabersonine, coenzyme A, K(+), Mg(2+) and Mn(2+). Induced by light with a maximum 48 hours after initiation of the light treatment. Belongs to the plant acyltransferase family. Was originally purified as a heterodimer of the N- and C-terminal end of the DAT gene product, but the cleavage of DAT protein appears to be a purification artifact.