Regulatory subunit of the condensin-2 complex, a complex which establishes mitotic chromosome architecture and is involved in physical rigidity of the chromatid axis (By similarity). May promote the resolution of double-strand DNA catenanes (intertwines) between sister chromatids (By similarity). Required for plant vigor, fertility, chromatin condensation and sister chromatid cohesion both during mitosis and meiosis (PubMed:23929493, PubMed:27385818). Necessary to maintain normal structural integrity of the meiotic chromosomes during the two nuclear divisions of gametogenesis, especially to prevent interchromosome connections at metaphase I (PubMed:25065716). Seems also involved in crossover formation during meiotic prophase I (PubMed:25065716). Prevents centromeric and pericentromeric heterochromatin repeats association (PubMed:23929493). Component of the condensin-2 complex. Observed throughout euchromatic nucleoplasm (PubMed:23929493, PubMed:27385818). Associates with the chromosomes throughout meiosis (PubMed:25065716). Present in buds. Mostly expressed at bolting, flowering and during seed formation. Regulated in a cell cycle-dependent manner with an increase during G2 phase, highest levels in the middle of G2 and a drop during mitosis (PubMed:23929493). Induced by MMD1 (PubMed:27385818). Dwarf plants with small rosette leaves (PubMed:25065716). Reduced pollen fertility and seed set (PubMed:23929493, PubMed:27385818). Lower chromatin density and frequent (peri)centromere association in interphase nuclei as well as impaired sister chromatid cohesion (PubMed:23929493). Altered interphase chromatin architecture in differentiated nuclei (PubMed:23929493). Defect chromosome condensation in male meiocytes and in centromeres orientation (PubMed:27385818). During meiosis, extensive interchromosome connections at metaphase I as well as a slight reduction in crossover formation (PubMed:25065716).