Transcription activator that plays a role in the regulation of cell expansion in leaf and cotyledons tissues. Component of a network formed by miR396, the GRFs and their interacting factors (GIFs) acting in the regulation of meristem function, at least partially through the control of cell proliferation. Predominantly expressed in shoot tips and flowers. microRNA 396 (miR396a or miR396b) negatively regulates growth-regulating factors (GRF1-4 and GRF7-9). The QLQ domain and WRC domain may be involved in protein-protein interaction and DNA-binding, respectively. Belongs to the GRF family.