Part of the tectonic-like complex which is required for tissue-specific ciliogenesis and may regulate ciliary membrane composition. Part of the tectonic-like complex (also named B9 complex). Localizes at the transition zone, a region between the basal body and the ciliary axoneme. Ciliary phenotypes such as pericardial effusion, hydrocephalic brain, curved or kinked tail, gastrulation defects and other severe defects in left/right axis.