Transcription factor required for lateral development of the lemma and palea (PubMed:22203474, Ref.9, PubMed:23526395, PubMed:25224972, PubMed:26486996, PubMed:29057928, PubMed:30725309). Involved in the regulation of grain hull development (PubMed:23526395). Possesses transactivation activity in yeast, and may determine grain shape and size by modifying gene expression in the grain hull (PubMed:23526395). Regulates the expression of cell proliferation and expansion related genes (PubMed:26486996). Acts as transcriptional repressor and regulates cell expansion during the lateral development of spikelet (PubMed:29057928). May act upstream of hormone-related genes and starch and sucrose metabolism-related genes, which may be responsible for lemma and palea development, and grain filling, respectively (PubMed:30725309). Does not seem to function at stages of floral-organ initiation and patterning (Ref.9). Homodimer. Expressed in lemma, palea, rachis branches, flag leaves and young embryos (PubMed:22203474). Highly expressed in lemmas and paleas of young spikelets (Ref.9, PubMed:23526395, PubMed:26486996). Expressed in stamens, pistil and leaf sheaths (Ref.9). Triangular hull with tortuous lemma and palea (PubMed:22203474, PubMed:29057928). Defects in development of lemma and palea, which have a beak-like form (Ref.9, PubMed:25224972, PubMed:30725309). Beak-shaped grains of decreased width, thickness and weight with a loosely interlocked lemma and palea that are unable to close tightly (PubMed:23526395, PubMed:26486996, PubMed:29057928). Poor grain filling (PubMed:22203474, Ref.9, PubMed:23526395, PubMed:25224972, PubMed:26486996, PubMed:29057928, PubMed:30725309). Belongs to the plant homeotic and developmental regulators ALOG protein family.