Plays a crucial role in homologous chromosome pairing and synapsis in meiosis. Does not seem required for cytokinesis (PubMed:19392701). Is essential for meiotic bouquet formation, homologous chromosome pairing and normal recombination, and synaptonemal complex (SC) assembly. Required for the proper association of PAIR2 with chromosomes (PubMed:21119003). During interphase-early leptotene, distributed as numerous punctuate foci throughout the chromatin. Associates with chromosomal axes at leptotene, zygotene and pachytene (PubMed:21119003). Associates with chromosomal axes at zygotene (PubMed:27436711). Expressed in pollen mother cells and the ovule tissues during meiosis. Sterility in both male and female gametes.