Implicated in enhancement of V-sense gene expression. Acts a an inhibitor of C-sense gene transcription (By similarity). Homooligomer. Interacts (via LXCXE domain) with host retinoblastoma-related protein 1 (RBR1), and may thereby deregulate the host cell cycle. Part of the C- and V-complexes which are RepA-Rep-DNA complexes involved in the c-sense and v-sense transcription (By similarity). There are 3 rolling circle replication (RCR) motifs. RCR-2 may be involved in metal coordination. RCR-3 is required for phosphodiester bond cleavage for initiation of RCR. The LXCXE motif specifically binds to host RBR1. Produced from the unspliced transcript. Belongs to the geminiviridae Rep protein family.