Transcription activator essential for the anti-viral defense called virus basal resistance response pathway (PubMed:11041886, PubMed:15629774, PubMed:18785827, PubMed:24418554). Not involved in HRT-mediated hypersensitive response (HR) and resistance to TCV (PubMed:18785827). Binds DNA non specifically (PubMed:15629774). Activated by proteolytic cleavage through regulated intramembrane proteolysis (RIP) (By similarity). (Microbial infection) Compromised function in defense response pathway when interacting with the invading viral capsid protein (CP) of turnip crinkle virus (TCV) due to abnormal subcellular localization. (Microbial infection) Interacts via its C-terminal region with the N-terminal region of the turnip crinkle virus (TCV) capsid protein (CP); this interaction prevents its nuclear localization and inhibits its function in basal resistance response pathway. (Microbial infection) Nuclear localization is blocked by its interaction with the coat protein (CP) of turnip crinkle virus (TCV), thus leading to its accumulation in inclusion-like structures peripheral to the nuclei. Induced by bacterial pathogens type III effector proteins (TTEs). (Microbial infection) Accumulates 2 days post infection with turnip crinkle virus (TCV) (PubMed:24418554). The NAC domain includes a DNA binding domain and a dimerization domain. Phosphorylated at Thr-142. Phosphorylation at Thr-142 is required for nuclear import. Increased replication of turnip crinkle virus (TCV) and cucumber mosaic virus (CMV) (PubMed:18785827, PubMed:24418554). Normal HRT-mediated hypersensitive response (HR) and resistance to TCV (PubMed:18785827). Delayed flower development in antisense asTIP plants (PubMed:24418554).