Transcription factor that determines dorsal-ventral body axis. Monomer. At the neurula stage, high expression in dorsal embryo region including neural folds and somites. Also high expression in adult brain (CNS) and low expression in oocytes. Expressed both maternally and zygotically. Threonine (predominantly) and serine residues are phosphorylated during oocyte maturation, when CDK1 is active.