Putative microtubule-associated force-producing protein, able to bind and hydrolyze GTP (By similarity). Collaboratively with DRP1A, participates in clathrin-coated vesicle formation during endocytosis (PubMed:20231465). With DRP1A and PIP5K3, required for the precise coordination of polar ARAC3/ROP6 and ARAC4/ROP2 placement and subsequent root hair positioning during planar polarity formation in root hair-forming cells (PubMed:27251533). GTP + H2O = GDP + H(+) + phosphate Interacts with DRP1A at the plasma membrane and in forming clathrin-coated vesicles (CCV). Accumulates in a sterol-enriched, polar membrane domain during root hair initiation. Ubiquitous. Preferentially expressed in siliques. Basal shift of ARAC3/ROP6 and ARAC4/ROP2 positioning in root hair-forming cells leading to basal shift of root hair positions. Belongs to the TRAFAC class dynamin-like GTPase superfamily. Dynamin/Fzo/YdjA family.