Component of the PAF1 complex (PAF1C) which is involved in histone modifications such as methylation on histone H3 'Lys-4' (H3K4me3) (PubMed:20363855). Involved in regulation of flowering time. Required for the expression of the flowering repressor and MADS box gene FLC (PubMed:12750345, PubMed:18725930). Required for histone H3 trimethylation on 'Lys-4' (H3K4me3) and histone dimethylation on 'Lys-36' (H3K36me2) at the FLC locus. Prevents trimethylation on 'Lys-27' (H3K27me3) at the same locus (PubMed:18725930). Not required for meiotic recombination or progression (PubMed:16716192). Component of the SKI complex which is thought to be involved in exosome-mediated RNA decay and associates with transcriptionally active genes in a manner dependent on PAF1 complex (PAF1C) (PubMed:22511887). Required for proper progression of cell differentiation process (PubMed:23134555). Component of the nuclear PAF1 complex (PAF1C), which consists of VIP2/ELF7/PAF1, VIP3/SKI8/WDR61, VIP4/LEO1, VIP5/RTF1, VIP6/ELF8/CTR9 and CDC73 (PubMed:20363855). Component of the cytoplasmic SKI complex, which consists of SKI2, SKI3 and VIP3/SKI8 (PubMed:22511887). Interacts with VIP4 and VIP6 (PubMed:15472079). Early flowering, reduced plant size and defects in floral morphology in whorls 1-3, but fully fertile flowers (PubMed:12750345, PubMed:16716192). Growth defects due to extra shoot apical meristem (SAM) formation (PubMed:23134555).