Vacuolar transmembrane transporter that participates in the first stage of the beta-lactam biosynthesis (the formation of the ACV tripeptide), probably taking part in the supply of amino acids from the vacuolar lumen to the vacuole-anchored ACV synthetase. Leads to a drastic reduction in the production of penicillin and its biosynthetic intermediates delta-(L-alpha-aminoadipyl-L-cysteinyl-D-valine) and isopenicillin N. Belongs to the CSC1 (TC 1.A.17) family.