Molecular adapter that acts as a bridge between a variety of multiprotein complexes, and which is involved in embryonic development, immunity, myogenesis and brain development (PubMed:28193841). Plays a key role in nuclear protein degradation by promoting import of proteasomes into the nucleus: acts by bridging fully assembled 20S proteasomes with nuclear import receptor ipo9 (By similarity). Involved in both neural precursor maintenance and terminal neural differentiation: bridges gmnn and actl6a/baf53a in neural progenitor cells, antagonizing the activity of gmnn, thereby suppressing sox2 expression (PubMed:28193841). Also required for proper activation of neurod1 and neuronal differentiation (PubMed:28193841). Involved in myogenesis: required for skeletal muscle formation and skeletal development, possibly by regulating expression of muscle differentiation factors (By similarity). Homodimer (By similarity). Interacts with actl6a/baf53a (PubMed:28193841). Interacts with gmnn (PubMed:28193841). Belongs to the akirin family.