Plant non-specific lipid-transfer protein that binds phospholipids in vitro (PubMed:33093144). Required for correct pollen exine patterning by controlling the continuity and homogeneity of the primexine distribution (PubMed:33093144). Localizes to the microspore plasma membrane. Expressed in young panicles (PubMed:30675653). Specifically expressed in pollen mother cells and young microspores (PubMed:33093144). Abnormal exine pattern of pollen grains and complete male sterility. Belongs to the plant LTP family.