Negative regulator of flowering controlling the H4K5 acetylation levels in the FLC and FT chromatin. Positively regulates FLC expression. Component of the transcription factor IID (TFIID) complex that is essential for mediating regulation of RNA polymerase transcription. Component of the SWR1 complex which mediates the ATP-dependent exchange of histone H2A for the H2A variant HZT1 leading to transcriptional regulation of selected genes by chromatin remodeling. Component of a NuA4 histone acetyltransferase complex which is involved in transcriptional activation of selected genes principally by acetylation of nucleosomal histones H4 and H2A. Component of the TFIID complex. TFIID is composed of TATA binding protein (TBP) and a number of TBP-associated factors (TAFs) whose MWs range from 14-217 kDa. Component of the SWR1 chromatin-remodeling complex. Interacts with TAF4B, TAF5, TAF8, TAF12, TAF12B, TAF13 and TAF15B. Interacts with the transcription activator complex FRI-C. Interacts with SWC6, FRI and (via C-terminus) with FLX, SUF4, FRIL1 and FES1 (PubMed:17340043, PubMed:21282526). Interacts with EAF1A and EAF1B (via HSA domain) (Ref.7). Expressed in roots, leaves, inflorescence and flowering tissues. No visible phenotype, due to the redundancy with TAF14B (PubMed:21282526). Decreased FLC expression leading to earlier transition from vegetative to reproductive phase (Ref.7). Taf14 and taf14b double mutants show a pleiotropic phenotype that includes small size and abnormal leaf morphology (PubMed:21282526). Taf14 and taf14b double mutants flower significantly earlier than single mutants under both long days and short days conditions (Ref.7). Belongs to the YAF9 family.