Component of the replication protein A complex (RPA) required for DNA recombination, repair and replication. The activity of RPA is mediated by single-stranded DNA binding and protein interactions. Plays an essential role at later stages of meiotic recombination events required for the formation of class I crossovers. Is essential for normal progression through meiosis in pollen mother cells. Is involved in repair of double-strand DNA breaks (DSBs) induced by genotoxic stresses, but does not seem to be required for the repair of meiotic DSBs. Heterotrimer of RPA1, RPA2 and RPA3 (canonical replication protein A complex) (By similarity). Interacts with RPA2A. Associated with meiotic chromosomes from leptotene through to early pachytene. A number of isoforms are produced. According to EST sequences. Expressed in roots, leaves, stalks and flower buds. By the genotoxic agents methyl methanesulfonate (MMS) and bleomycin. There are differing accounts of the mutant phenotypes. According to an early study with a limited number of progeny from different sources (PubMed:15978034), it is embryonic lethal when homozygous. However, a more recent study describes homozygous lines that show normal vegetative growth, but reduced fertility and meiotic defects (PubMed:19153602). Belongs to the replication factor A protein 1 family.