The GINS complex plays an essential role in the initiation of DNA replication (PubMed:17556508). Required during embryogenesis (PubMed:22164284). Component of the GINS complex (By similarity). Interacts with EOL1 in the nucleus (PubMed:28428341). Expressed in callus 96 hours after initiation on callus-inducing medium. Embryo defective. Belongs to the GINS4/SLD5 family.