Transcription activator involved in the spatiotemporal regulation of anthocyanin biosynthesis specifically in the inner pericarp of red-fleshed kiwifruits (PubMed:30912865). Functions in association with BHLH42 to activate the promoters of LDOX (ANS) and F3GT1 that encode the dedicated enzymes for anthocyanin biosynthesis (PubMed:30912865). Interacts with BHLH42. Expressed in the inner pericarp of maturing fruits.