Polycomb group (PcG) protein. Involved in flowering processes by repressing unknown target genes and preventing reproductive development. Participates in polycomb group (PcG) protein complex-mediated (probably in complex with EMF1) silencing of the flower homeotic genes AGAMOUS (AG), PISTILLATA (PI), and APETALA3 (AP3), as well as of some regulatory genes such as ABSCISIC ACID INSENSITIVE3 (ABI3), LONG VEGETATIVE PHASE1 (LOV1), and FLOWERING LOCUS C (FLC) during vegetative development, by mediating trimethylation of histone 3 lysine 27 on the AG chromatin (H3K27me3). PcG proteins act by forming multiprotein complexes, which are required to maintain the transcriptionally repressive state of homeotic genes throughout development. PcG proteins are not required to initiate repression, but to maintain it during later stages of development. They probably act via the methylation of histones, rendering chromatin heritably changed in its expressibility. In plants, PcG complexes are probably composed of a member of the EZ family (CLF or MEA), FIE, and a member of the VEFS family (FIS2, VRN2 or EMF2) (By similarity). Binds to ALP1 (PubMed:26642436). Widely expressed throughout the life cycle with higher levels in proliferating tissues. Expressed in both vegetative and the reproductive shoot meristems. Expressed in the developing embryos and endosperm, then decreases when embryos mature and soon after cellularization in the endosperm. After germination, it is expressed in the shoot apical meristems (SAMs), leaf primordia, and young leaves. In the reproductive shoots, it is expressed in both the influorescence and floral meristems. Later, it is expressed in floral organ primordia. In coflorescences, it is expressed in SAMs and lateral organs. In roots, it is expressed in root tips. Belongs to the VEFS (VRN2-EMF2-FIS2-SU(Z)12) family. Was originally thought to correspond to two different genes At5g51230 and At5g51240. Was originally thought to correspond to two different genes At5g51230 and At5g51240.