Regulates intracellular cholesterol distribution from recycling endosomes to the trans-Golgi network through interactions with RAB11 and OSBP. Functions in membrane tethering and promotes OSBP-mediated cholesterol transfer between RAB11-bound recycling endosomes and OSBP-bound Golgi-like membranes. Interacts with RAB11A (VIA-GTP form) (PubMed:29514919). Interacts with RAB11B (PubMed:29514919). Interacts (via the third HEAT repeat) with OSBP (via C-terminus) (PubMed:29514919). Found in a complex composed of RELCH, OSBP1 and RAB11A (PubMed:29514919). Colocalization with RAB11A in recycling endosomes (PubMed:29514919). Translocated to the trans-Golgi network area in an OSBP-dependent manner (By similarity). The sequence differs from that shown because it is derived from pre-RNA.