Component of SUN-protein-containing multivariate complexes also called LINC complexes which link the nucleoskeleton and cytoskeleton by providing versatile outer nuclear membrane attachment sites for cytoskeletal filaments (By similarity). Required for nucleus structure organization (e.g. size and shape) (PubMed:24308514, PubMed:23396599). Core component of the LINC complex which is composed of inner nuclear membrane SUN domain-containing proteins coupled to outer nuclear membrane WIP proteins, the nucleoskeletal CRWN/LINC proteins, and, possibly, KAKU4. Recruited to the nucleus envelope (NE) by SUN proteins and is immobilised therein (By similarity). Punctate or bundle-like structures are detected, especially in trichomes where bundle-shape localization pattern along the long axis of the nucleus is observed. During prometaphase to anaphase, localized diffusely in the cytoplasm. Later transferred from the cytoplasm to the chromatin surface, preferentially assembling to the distal surface of the chromatin. Relocalized to the nuclear periphery during late telophase (PubMed:23396599). Expressed at low levels in roots, leaves, flowers and flower stalks. Altered nuclear morphology. Plants lacking both CRWN1 and CRWN3 exhibit markedly smaller rosettes. Plants lacking CRWN1, CRWN3 and CRWN4 are extremely stunted and set few seed. Belongs to the CRWN family.