Involved in chromosome segregation during meiosis. Promotes efficient recombinational repair and functions in the protection of the genome from spontaneous and induced DNA damage like mutations and gross chromosomal rearrangements (GCRs). Component of the SHU complex composed of at least CSM2, PSY3, SHU1 and SHU2. Present with 414 molecules/cell in log phase SD medium. Belongs to the CSM2 family.