Probable beta-1,3-glucanase that may be involved in the degradation of callose walls around the microspore tetrad during pollen development (Probable). May be required for pollen exine formation (PubMed:21849515). Hydrolysis of (1->3)-beta-D-glucosidic linkages in (1->3)-beta-D-glucans. Anthers. Contains two additional disulfide bonds, but it is unclear if they are between the pairs Cys-409-Cys-416 and Cys-425-Cys-471 or between the pairs Cys-409-Cys-471 and Cys-416-Cys-425. Embryonic lethality due to division arrested at one-cell zygotic stage. Belongs to the glycosyl hydrolase 17 family.