Putative transcription factor. Homo- and heterodimer with other ZFHD proteins (By similarity). Interacts with MIF1, MIF2 and MIF3; these interactions prevent nuclear localization and DNA-binding to inhibit transcription regulation activity. Binds to ZHD1, ZHD2, ZHD4, ZHD10 and ZHD11. Interacts with HIPP30 (PubMed:18974936). Interactions with MIF proteins prevent nuclear subcellular location and leads to a scattered repartition throughout the cytoplasm. Mostly expressed in flowers and inflorescence. The homeodomain differs form the typical one by having namely 4 instead of 3 extra amino acids inserted in the loop between helix 1 and helix 2.