Component of a deubiquitination module (DUB module) that specifically deubiquinates monoubiquinated histone H2B (H2Bub) (PubMed:29588169, PubMed:30192741). Does not seem to be a component of the TREX-2 complex (PubMed:29588169). Seems to act independently of the SAGA multiprotein complex (PubMed:30192741). The DUB module is responsible for the major H2Bub deubiquitinase activity in Arabidopsis (PubMed:30192741). Component of a deubiquitination module (DUB module) formed by ENY2, SGF11, and UBP22 in Arabidopsis (PubMed:29588169, PubMed:30192741). Interacts directly with SGF11, but not with UBP22 (PubMed:29588169, PubMed:30192741). Interacts with MOS4 (PubMed:29588169). Does not localize in nucleolus and displays a rather patchy distribution forming a speckled pattern in the nucleoplasm (PubMed:29588169, PubMed:30192741). Does not localize in the nuclear pore complex at the nuclear periphery (PubMed:29588169, PubMed:30192741). Expressed in roots, cotyledons, leaves and upper part of sepals. Belongs to the ENY2 family.