Seminal fluid metalloprotease which is transferred to females during mating and is required for processing of two other seminal fluid proteins Acp26Aa and Acp36DE in mated females. Binds 1 zinc ion per subunit. Secreted into seminal fluid. Produced in the male accessory glands and secreted into seminal fluid. In mated females, confined to the reproductive tract and also detected in eggs laid by mated females (at protein level). Undergoes cleavage in the male during mating with a cleaved product detected in the ejaculatory duct and/or bulb of males by 8-10 minutes after the start of mating (PubMed:17116868). Further cleavage occurs in the mated female (PubMed:15979005). May undergo cleavage in a two-step process where it is first cleaved by Sems, making it susceptible to activational cleavage which may be carried out by another protease or by autocleavage (PubMed:24514904). Loss of cleavage of accessory gland proteins Acp26Aa and Acp36DE in females mated with mutant males (PubMed:24514904). RNAi-mediated knockdown in males results in incomplete and delayed processing of Acp26Aa and Acp36DE in females mated with these males. Extended N-terminus.