Involved in oxygen transport from the gills to various peripheral tissues. Heterotetramer of two alpha and two beta chains. Red blood cells. Acetylated on Ser-1. This fish has two types of hemoglobin: one cathodic Hb and four anodic Hbs. The cathodic Hb displays a small normal Bohr effect and a reverse Bohr effect in the presence and absence of phosphate respectively. In addition, the cathodic Hb displays a large phosphate effect. Belongs to the globin family.