May play a role in sucrose partitioning during seed development and in stress response. Hydrolysis of terminal non-reducing beta-D-fructofuranoside residues in beta-D-fructofuranosides. Associated to the cell wall. Expressed in roots, leaves and flowers. Weakly expressed in seeds. Expressed from 1 to 3 days after flowering mainly in the maternal tissues of the developing caryopsis, corresponding to the early grain filling stage. By sugar in excised leaves. By rice blast fungus (M.grisea) 4 hours after infection. Belongs to the glycosyl hydrolase 32 family.