Could play a role in maintaining cochlear stereocilia bundles that are involved in sound detection. Expressed in sensory hair cells in the cochlea and vestibular organ. Mice appear normal at postnatal week 2, but develop progressive hearing loss associated with early defects in orientation and organization of cochlear stereocilia bundles. They have normal vestibular function. Belongs to the GRXCR1 family. Although it shares high sequence similarity with GRXCR1, it does not contain a canonical glutaredoxin domain.