Involved in epithelial cell integrity by acting on the dynamics of the actin cytoskeleton (PubMed:30361697). Positively regulates the actin polymerization, by inhibiting the interaction of actin-capping proteins with actin (By similarity). Directly interacts with actin-capping proteins CAPZA1, CAPZA2 and CAPZB; this interaction decreases the binding of capping proteins to actin. Expressed in the small intestine (at protein level). Knockout mice display adenoma development in the small intestine in an age-dependent manner. They also develop pulmonary lesions resembling early small cell lung cancer and solid-pseudopapillary neoplasm of the pancreas. Tumors from mutant animals show increased expression of Wnt/beta-catenin target genes without the alteration of other signaling pathways. Intestinal tumors exhibit disorganized and decreased levels of F-actin.