Acts as a positive regulator of hedgehog signaling and regulates ciliary function. Found in a complex with TEDC1, TEDC2, TUBE1 and TUBD1. Associated with centrioles. Both cytoplasmic and nuclear. In the elongating spermatid it is associated with the manchette, a specialized microtubule system present during reshaping of the sperm head. Belongs to the tubulin family.