Required to maintain the structure of the centrosomal microtubule organizing center (MTOC) during mitosis. May have a preferential role in regulating neurogenesis. Required for germ cell mitosis and oocyte differentiation. During interphase in syncytial embryos distribution is cytoplasmic. On entering mitosis, moves to polar regions of the spindle immediately surrounding the centrosome. At telophase, migrates to microtubules on the spindle side of both daughter nuclei. The nuclear-cytoplasmic distribution could be regulated by the availability of calmodulin. Expressed both maternally and zygotically in embryos. Intron retention.