Possesses intrinsic ATP-dependent nucleosome-remodeling activity. Constitutes the catalytic subunit of several complexes capable of forming ordered nucleosome arrays on chromatin (By similarity). Involved in the formation of nucleosome distribution patterns (PubMed:24606212). Involved in nuclear proliferation during megagametogenesis and cell expansion in the sporophyte (PubMed:16286646). Required for the maintenance of the plant vegetative phase. In association with RLT1 or RLT2 may prevent the early activation of the vegetative-to-reproductive transition by regulating key genes that contribute to flower timing, such as FT, SEP1, SEP3, AGL8/FUL, SOC1 and FLC (PubMed:22694359). Necessary to acquire heat stress (HS) memory (PubMed:27680998). Interacts with RLT1 and RLT2 (PubMed:22694359). Interacts (via C-terminus) with RLT1 (via the DDT domain), RLT2 (via the DDT domain), PTM (via the DDT domain) and DDR4 (via the DDT domain) (PubMed:23691993). Binds to FGT1 (PubMed:27680998). Highly expressed in growing tissues such as inflorescence and flower meristems, young leaves and floral organs. Expressed in roots, rosette and cauline leaves, stems, flowers, inflorescences and siliques. In developing ovule, expressed in all cells of the young nucellus, including the functional megaspore. After the initiation of megagametogenesis, expressed in most cells of the ovule, including the integuments, the developing megagametophyte, and the funiculus. In mature ovules, strongly expressed in the cellularized megagametophyte. After double fertilization, expressed in the developing embryo and the free nuclear endosperm until seed maturity. Expressed in developing male gametophytes and mature pollen grains. Reduced heat stress (HS) memory. Premature decline of expression of HSA32, HSP18.2, HSP21, HSP22 and HSP101 after HS in the double mutant plants chr11-1 and chr17-1 (PubMed:27680998). No visible phenotype under normal growth conditions, but the double mutant plants chr11-1 and chr17-1 are very small and display early flowering and sterility (PubMed:22694359). Plants silencing CHR11 display reduced plant height and small cotyledonary embryos with limited cell expansion. Plants silencing CHR11 specifically at the onset of female gametogenesis (megagametogenesis) have defective female gametophytes arrested before the completion of the mitotic haploid nuclear divisions. Belongs to the SNF2/RAD54 helicase family. ISWI subfamily.