Exhibits lactonase activity for dihydrocoumarin or 2-coumaranone, although its natural substrate is not known. Acts in cells with perturbed membrane integrity and is possibly related to the membrane homeostasis. Contributes to bacitracin resistance. Binds 2 Ca(2+) ions per subunit. Activity is decreased by EGTA or EDTA. Induced by cell wall-affecting antibiotics such as oxacillin, beta-lactams, vancomycin, fosfomycin, bacitracin and detergents such as Triton X-100, SDS, Nonidet P40, and CHAPS. Not essential for cells to maintain the levels of resistance to most of the cell wall-affecting antibiotics although they can effectively induce its expression. Belongs to the SMP-30/CGR1 family.