Receptor for the leucine-rich nuclear export signal (NES). Binds cooperatively to the NES on its target protein and to the small GTPase Ran in its active GTP-bound form (By similarity). Required for the maternal-to-embryonic transition and during gametophyte development (PubMed:18791220). Present in mature pollen grains, unpollinated pistils, and 2-week-old seedlings. Gametophyte defective when both XPO1A and XPO1B are disrupted. Abnormal pollen germination and tube growth, impaired female gametophyte development and embryo lethal. Belongs to the exportin family. Truncated N-terminus. Extended N-terminus.