Component of the plant cell wall polysaccharide acetylation pathway (PubMed:25681330, PubMed:30083810). Does not directly catalyze O-acetylation of xyloglucan but exhibits weak acetylesterase activity in vitro (PubMed:30083810). Severe growth defects and collapsed xylem (PubMed:25681330). Decreased xyloglucan acetylation and decreased O-acetylation of xylan (PubMed:25681330).