Polycomb group (PcG) protein. PcG proteins act by forming multiprotein complexes, which are required to maintain the transcriptionally repressive state of homeotic genes throughout development. PcG proteins are not required to initiate repression, but to maintain it during later stages of development. They probably act via the methylation of histones, rendering chromatin heritably changed in its expressibility. Required to prevent the proliferation of the central cell by repressing unknown target genes before fertilization. Probably also involved in floral repression mechanism established during early plant development. Regulates the anteroposterior organization of the endosperm. Interacts with the promoter and represses the transcription of genes such as PHE1, that are paternally active and maternally silenced. Interacts directly with MEA. These two proteins are probably indirectly associated with FIS2. In plants, PcG complexes are probably composed of a member of the EZ family (CLF or MEA), FIE, and a member of the VEFS family (FIS2, VRN2 or EMF2). Component of the plant homeodomain / polycomb repressive complex 2 (PHD-PRC2) large complex during prolonged cold, composed of core PRC2 components (VRN2, EZA1, FIE and MSI1), and three related PHD finger proteins (VIL1, VIL2 and VIN3) that mediates histone H3 trimethylation on 'Lys-27' (H3K27me3). Binds to ALP1 (PubMed:26642436). Excluded from the nucleolus. Expressed in cauline leaves, root and stems. In the male reproductive organ, it is expressed in the developing anther; and is abundant in microspore mother cells, in microsporocytes and in the tapetum, but is absent from vascular bundles, the connective tissue and the filament. It is also absent from pollen grains at subsequent developmental stages. In the developing female reproductive organs, it is highly expressed in all cells of the young ovules primordium before archesporial differentiation. Then, it is highly expressed in the ovule sporophytic tissue and the megaspore mother cell before meiosis, but is absent from placenta or the developing carpel. Then, it decreases. Expressed maternally and zygotically. Expressed in both egg and central cell before fertilization, and in the embryo and endosperm after fertilization. Belongs to the WD repeat ESC family.