Lipid-transfer proteins that possesses lipid-binding activity in vitro. Involved in the development of lipidic orbicules/Ubisch bodies and pollen exine during anther development (PubMed:20610705). May be regulated by the transcription factor UDT1 in developing anthers and play a role in tapetum development (PubMed:16141453). Positively regulated by the transcription factor TDR in developing anthers and may play a role in tapetum programmed cell death (PCD) (PubMed:17138695). During anther development, expressed in tapetal cells of immature anthers, but not in mature pollen (PubMed:1463854, PubMed:20610705). Weakly expressed in microspores from stage 9 to stage 11 of anther development (PubMed:20610705). Belongs to the plant LTP family. Sequencing errors.