Promotes flowering by suppressing FLC-mediated inhibition of flowering through the regulation of FCA pre-mRNA alternative processing (PubMed:18298670). Regulates mRNA maturation including alternative polyadenylation, splicing or transcription initiation of stress-associated genes (PubMed:21364912). Homodimer. Forms a complex with cleavage and polyadenylation specificity factor (CPSF) subunits CPSF30, CLPS3 and PCFS1. Expressed in embryos, seedlings, roots, leaves and flowers. During embryogenesis, first observed at the heart stage. Expressed through the whole developing embryo from the torpedo stage. Disappears in the root tip of the mature embryo and in adult plants. In seedlings, mostly present in the shoot apical meristem and young leaves. In flowers, detected in sepals, stamens and pistils, but not in petals. Delayed flowering mediated by FLC under both long-day and short-day conditions, but normal response to vernalization treatment. Abnormal alternative polyadenylation-dependent FCA expression profile of the known transcripts (PubMed:18298670). Global differentially processed (DPG) and differentially expressed (DEG) stress-associated genes due to alternative polyadenylation, splicing or transcription initiation (PubMed:21364912).