Essential for pollen development. Involved in protein N-glycosylation in the endoplasmic reticulum (ER), especially in the female gametophyte. Mediates pollen tube (PT) reception in synergids through protein glycosylation. CTP + di-trans,poly-cis-dolichol = a dolichyl phosphate + CDP + H(+) Pollen tube (PT) overgrowth inside the female gametophyte (FG) without PT rupture. Degenerated pollen grains before maturation, during the early tricellular stage. Belongs to the polyprenol kinase family.