Plays a fundamental role in building the proper chromosome structure at the beginning of meiosis in male meiocytes. Required for the transition from leptotene to zygotene in meiocytes. Required for homologous chromosome pairing, and initiation and progression of meiotic recombination. Regulates meiocyte cytoskeleton organization. Diffuse localization in the nucleus during the initiation of meiosis. Binds to chromatin in early meiotic prophase I. Sterile phenotype due to male meiosis arrested at leptotene.