Acts as both a repressor and activator of transcription. Binds to the consensus sequence 5'-GACCACCCAC-3'. In the embryo, expressed at high levels in the kidney and testis. In the adult, expressed at high levels in the kidney and uterus and at lower levels in the brain, lung, skeletal muscle and pancreas. In the embryo, expression is detected between days 6.5 and 14.5, particularly during neurulation. At embryonic days 11.5 to 12.5, expression is high in the interdigital regions destined to undergo apoptosis. Belongs to the GLI C2H2-type zinc-finger protein family.