Has a role in female meiotic chromosome segregation in females; proximal heterochromatin is involved in chromosome pairing during female meiosis. Is a dominant suppressor of both white and Stubble position-effect variegation (PEV), while it is a weak enhancer of brown variegation. Expressed in embryos. Flies exhibit brain cell metaphases where sister chromatids of all chromosomes are aligned parallel to each other instead of assuming the typical morphology, heterochromatin condensation or chromosome segregation are not affected. Belongs to the WAPL family.