Transcriptional regulator involved in the regulation of cell differentiation in meristems (By similarity). Binds DNA without sequence preference (PubMed:22226340). Localizes to the cytoplasm in non-differentiating cells and to the nucleus in differentiating cells. Expressed in rosette leaves, stipules, stems, flowers, siliques and mature seeds. Expressed in the vascular bundles of xylem in shoot parenchyma cells. Expressed in the remnant cytoplasm of differentiated fiber cells and in protoxylem element of parenchymal cells. Contains a bacterial GIY-YIG-like domain.