Probable tRNA nucleus export receptor which regulates tRNA processing and facilitates tRNA translocation across the nuclear pore complex. Is required for proper activity of the shoot apical meristem (SAM) and correct leaf initiation at different developmental stages, and may play a role in floral patterning. Shuttles between the nucleus and the cytoplasm. Expressed in young leaves, growing leaf blades, young floral organs and root tips. Delayed leaf formation during vegetative development, abnormal SAM layered structure, altered inflorescences, disrupted phyllotaxy, reduced number of seeds and reduced number of lateral roots. Belongs to the exportin family.