Involved in Fe(3+) uptake from the rhizosphere and phloem transport of iron. Plays an important role in iron homeostasis during the early stages of growth. Transports Fe(3+)-phytosiderophore, but not Fe(3+)- or Fe(2+)-nicotianamine. May not transport other chelated metals. Expressed in root phloem and at low levels in the shoot companion cells. Before anthesis, expressed in the vascular bundles of spikelets, stamens, pistils, and husk. After fertilization, expressed in the husk, spikelets, and pistil. From 3 to 10 days after anthesis, expressed in the dorsal vascular bundles of developing ovaries. At 15 days after anthesis, expressed in the embryo and dorsal vascular bundles. At 30 days after anthesis, expressed in the scutellum and endosperm. In roots by iron deficiency. Belongs to the YSL (TC 2.A.67.2) family.