Acts redundantly with CYP98A9 as tricoumaroylspermidine meta-hydroxylase. Catalyzes also the meta-hydroxylation of the three triferuloylspermidine phenolic rings. Unable to use 5-O-(4-coumaroyl) D-quinate or 5-O-(4-coumaroyl) shikimate as substrates. Strongly expressed in inflorescence tips, young flower buds, seeds, stamen, tapetum and pollen. Drastic reduction in flower buds of the content in N1,N5,N10-tri-(hydroxyferuloyl)-spermidine and N1,N5-di(hydroxyferuloyl)-N10-sinapoyl-spermidine. Belongs to the cytochrome P450 family. Extended C-terminus.