Catalyzes the dehydration of cis-3-hydroxy-L-proline (c3LHyp) to Delta(1)-pyrroline-2-carboxylate (Pyr2C). Is likely involved in a degradation pathway that converts c3LHyp to L-proline, which allows L.aggregata to grow on c3LHyp as a sole carbon source. Also catalyzes the epimerization of c3LHyp to trans-3-hydroxy-D-proline (t3DHyp), a competing reaction occurring from the same enolate anion intermediate. L-proline, t3LHyp, t4LHyp, c4DHyp and their methylated derivatives are not substrates. cis-3-hydroxy-L-proline = 1-pyrroline-2-carboxylate + H2O Binds 1 Mg(2+) ion per subunit. kcat is 14 sec(-1) for c3LHyp epimerization, and 9 sec(-1) for c3LHyp dehydration. Is up-regulated when the bacterium is grown on c3LHyp or t3LHyp as sole carbon source. Belongs to the mandelate racemase/muconate lactonizing enzyme family.