RNA-binding protein that functions in both mitochondrion and nucleus. In mitochondrion, it is associated with polysomes and may play a role in translation. Required during embryogenesis. In nucleus, might be involved in the regulation of its own gene expression. Interacts with NAP1;1 and TCP8. Able to bind mitochondrial RNA in vitro. Can localize to both mitochondrion and nucleus. Expressed in root tips, lateral root primordia and leaf primordia. Highly detected in the mature pollen grains. Embryo-lethal at a early stage of development. Belongs to the PPR family. P subfamily.