Scaffolding protein that plays a role in ciliogenesis, cilium positioning and dendrite anchoring in sensory amphid neurons including AWB, AWA, AWC, ADL and ASI and the phasmid neurons PHA and PHB. Its role in cilium positioning may be through regulation of the localization of cell adhesion proteins such as the apical junction protein ajm-1, and the ciliary scaffolding protein Rootletin/che-10. Enriched at the proximal ends of centrioles. Broadly expressed during embryogenesis with high expression at the presumptive nose during the 1.5-fold stage of embryogenesis. Expressed in AWB sensory neurons in larvae and in PHA/PHB tail neurons at the L1 stage of larval development. The C-terminal domain is required for localization to the cilium basal body and role in ciliogenesis. Embryonic lethal with few malformed larvae that develop to the L1 stage of larval development. Belongs to the CCDC88 family.