Acetyltransferase required for the establishment of sister chromatid cohesion (PubMed:20671110). Involved in preservation of genome integrity and meiosis (PubMed:23750584, PubMed:20671110). Required for DNA repair and for the regulation of chromosome segregation during mitotic cell division (PubMed:23750584, PubMed:20671110). Knock-down mutants are extremely dwarf (PubMed:20671110). Regulator of sister chromatid cohesion in meiosis which negatively regulates cohesin association with chromatin, acting as an antagonist of WAPL1 and WAPL2 (PubMed:26813623). Expressed in roots, stems, leaves, young seedlings and flower buds. Detected in the embryo, but not in the endosperm. Autoacetylated. Embryo lethality, but normal endosperm development (PubMed:20671110). No effect on pollen mitosis but minor alterations in female gametophyte development (PubMed:20671110). Defective microsporangenesis and anthesis (PubMed:20671110). Major defects in vegetative growth and development, as well as complete sterility; these phenotypes are partially restored in plants also missing WAPL1 and WAPL2 (PubMed:26813623). Belongs to the acetyltransferase family. ECO subfamily.