Possesses triterpene oxidizing activity. Catalyzes the C28 hydroxylation of alpha-amyrin, beta-amyrin, and lupeol, producing uvaol, erythrodiol, and betulin, respectively. Catalyzes the C28 carboxylation of alpha- and beta-amyrin. Possesses 22alpha-hydroxylation activity against alpha- and beta-amaryn. Belongs to the cytochrome P450 family. Was originally thought to correspond to two different genes At5g36130 and At5g36140. Was originally thought to correspond to two different genes At5g36130 and At5g36140. Was originally thought to correspond to two different genes At5g36130 and At5g36140. Was originally thought to correspond to two different genes At5g36130 and At5g36140.