Catalytic component of the CCR4-NOT complex which is one of the major cellular mRNA deadenylases and is linked to various cellular processes including bulk mRNA degradation, miRNA-mediated repression, translational repression during translational initiation and general transcription regulation (PubMed:23843623, PubMed:28204614). Within the complex, plays a role in miRNA-mediated deadenylation in embryos (PubMed:28204614). Within the complex promotes germ cell development and fertility in hermaphrodites (PubMed:23843623). Additional complex functions may be a consequence of its influence on mRNA expression. Exonucleolytic cleavage of poly(A) to 5'-AMP. Component of the CCR4-NOT complex at least composed of ccf-1, ccr-4 and let-711, which is required for germ cell development in hermaphrodites (PubMed:23843623). Within the complex interacts with let-711 (PubMed:23843623). Highly expressed in the germline (PubMed:23843623). In particular, highly expressed in germ cells that enter meiosis and progress through the pachytene stage (PubMed:23843623). Expressed at all developmental stages in males and hermaphrodites (PubMed:23843623). Expressed at low levels in embryos and L1 stage larvae (PubMed:23843623). RNAi-mediated knockdown either results in sterility or a reduced brood size (PubMed:23843623). RNAi-mediated knockdown disrupts the arrangement, differentiation and maturation of oocytes in the proximal region and as a result small oocyte-like cells arrange in several rows in the germline (PubMed:23843623). RNAi-mediated knockdown reduces miRNA-mediated deadenylation (PubMed:28204614). RNAi-mediated knockdown reduces ccr-4 and ntl-1 protein levels (PubMed:28204614). In another study, RNAi-mediated knockdown does not alter the levels of let-711 or ccr-4 (PubMed:23843623). RNAi-mediated knockdown results in reduced global poly(A) tail deadenylation (PubMed:23843623). Belongs to the CAF1 family.