Glycosyltransferase required for the regulation of cellulose biosynthesis in the cell wall (PubMed:18939965, PubMed:27708650). Required for the biosynthesis of hexoses (glucose, mannose and galactose) in both cellulosic and non-cellulosic (pectins and hemicelluloses) components of cell walls (PubMed:27708650). Required for the formation of arabinogalactan proteins which contribute to the strengthening of cell walls (PubMed:18939965). Possesses low glycosyltransferase activity (PubMed:18939965). Localizes in punctuate patterns. Expressed in roots, culms, leaves and panicles (PubMed:18939965). Expressed in vascular bundles of leaf sheaths and stems where sclerenchyma cells are developing (PubMed:18939965). Expressed in mechanical tissues of young organs, such as young leaf sheaths, stems and tiller buds (PubMed:18939965). Retarded growth and reduced mechanical strength (brittleness) due to reduced cell wall thickness in sclerenchyma and bundle sheath fiber cells. Belongs to the glycosyltransferase 14 family.