Required for elongation of meiosis I spindle. Critical for ovulation, meiotic progression in oocytes and female courtship behavior, including their postmating changes. Regulates female meiosis by controlling calcineurin activity in the germline. Has a role in calcium signaling during egg activation; bcd mRNA polyadenylation and translation in the oocyte. Interacts with Pp2B-14D, CanA-14F and CanB2. Expressed in central nervous system of the third instar larvae, with a relatively intense signal in the brain and weak signals in the ventral ganglion. Relatively low, but ubiquitous expression level is observed in leg and wing imaginal disks, no signal is detected in the eye-antennal disks. Expressed in all neurons in the adult brain. Expressed both maternally and zygotically. Phosphorylation at Ser-215 and Ser-219 is essential for calcineurin activation and completion of female meiosis. Sgg is required for phosphorylation of Ser-215 in activated eggs. Ser-100, Thr-102 and Ser-219 are highly phosphorylated in both ovaries and activated eggs; however, phosphorylation at Ser-100 or Thr-102 is not required for sra function in completion of female meiosis. Flies exhibit spontaneous ovulation in virgins, female sterility with impaired meiotic progression (eggs are arrested at anaphase of meiosis I, they fail to fully polyadenylate and translate bicoid mRNA and the male pronucleus fails to mature), and compromised postmating responses with lower ovulation level, higher remating rate, and shorter period for restoration of receptivity. Belongs to the RCAN family.