Transcription factor that specifically binds AT-rich DNA sequences related to the nuclear matrix attachment regions (MARs) (PubMed:25336567). Encodes a nuclear matrix protein that acts in the maintenance of genomic integrity by silencing TEs and repeat-containing genes through epigenetic machinery. Acts as a chromatin remodeling factor that modifies the architecture of FLC and FWA chromatin by modulating both H3 acetylation and methylation leading to the regulation of FLC and FWA expression (PubMed:23394836, PubMed:23836195). Negatively regulates floral repressors including MAF4 and MAF5 (PubMed:23733063). Plays a transcription activation role in anther development. Regulates the expression of arabinogalactan proteins (AGPs) involved in the formation of the nexine layer of the pollen wall (PubMed:25336567, PubMed:24804694). Binds AGP6, AGP11, AGP23 and AGP40 promoters (PubMed:25336567). Interacts with FVE/MSI4 and MSI5 which are components of HDAC corepressor complexes. Preferentially expressed in the inflorescence meristem and young floral buds, as well as in seedling-stage vegetative meristems (PubMed:23394836). Widely expressed in flowers, roots and stems, with relatively low expression in leaves (PubMed:24804694). Expressed in tapetum during the tetrad stage. The PPC domain mediates interactions between AHL proteins. Abnormal pollen wall with the absence of the nexine and intine layers causing microspore abortion and male sterility. AHL16/TEK knockdown results in late flowering phenotype.