Involved in the biogenesis of the 60S ribosomal subunit. May play a part in the quality control of pre-60S particles (By similarity). Component of the pre-66S ribosomal particle. Interacts with nop7 and rrp1. Interacts with rsa4 (via WD repeats). Belongs to the eukaryotic ribosomal protein eS8 family. Ribosome biogenesis protein NSA2 subfamily.