Involved in oxygen transport from gills to the various peripheral tissues. Heterotetramer of two alpha chains and two beta chains. Red blood cells. This fish has two hemoglobins: cathodic and anodic. The cathodic Hb and anodic Hb display small and large Bohr effects respectively. In addition, the cathodic Hb displays a reverse Bohr effect and appreciable phosphate effects. Belongs to the globin family.