Regulates floral organ identity and cell proliferation in the inner floral whorls. Probably specifies the identities of lodicule and stamen through positive regulation of MADS16 expression. May contribute to morphogenesis by suppressing OSH1 expression in the lateral organs. Expressed in leaf primordia, inflorescence meristem, rachis branch meristems, floral meristem and floral organ primordia. Male sterility. Increased number of inflorescence branches, modified floral organ identities, flowers with varied numbers of inner floral organs and amorphous tissues without floral organ identity in whorls 3 and 4. Malformation of leaf blades until the fourth-leaf stage.