Vacuolar metal transporter involved in intracellular metal homeostasis. Can transport iron (Fe), manganese (Mn) and cadmium (Cd). Regulates metal accumulation under Fe starvation. Acts redundantly with NRAMP3 to mobilize vacuolar Fe and provide sufficient Fe during seed germination. In association with NRAMP3, required for optimal growth and photosynthesis under Mn deficiency. Exports Mn from vacuoles in leaf mesophyll cells, making Mn available for functional photosystem II in chloroplasts. Expressed in vascular tissues. By iron starvation. No visible phenotype under normal growth condition, but slightly enhanced resistance of root growth in presence of Cd and increased accumulation of Mn and Zn under Fe starvation. Belongs to the NRAMP (TC 2.A.55) family.