Involved in target of rapamycin (TOR)-regulated translation control, especially under energy-deficient conditions. Binds to EIF4A1 (PubMed:29084871). The association with ribosomes is modulated by cellular energy status and TOR activity (PubMed:29084871). Mostly expressed, at low levels, in rosette leaves and flower buds, and, to a lower extent, in roots, stems, cauline leaves and flowers. Induced by dark and starvation but repressed by glucose feeding subsequent to starvation in a TOR-dependent manner. Increased susceptibility to dark and starvation, and to treatment with the TOR inhibitor (PubMed:29084871). Decreased translation activity associated with altered ribosome patterns, especially in the dark and starvation conditions, in which mRNAs distribution is altered and rRNA abnormally degraded (PubMed:29084871). Slightly early flowering time under long-day conditions (PubMed:29084871). Belongs to the PDCD4 family.