Rhodanese domain-containing protein required for anchoring ferredoxin--NADP reductase to the thylakoid membranes and sustaining efficient linear electron flow (LEF). Component of high molecular weight thylakoid LFNRs-containing protein complexes containing LIR1, LFNR1, LFNR2, TIC62 and TROL proteins. Mainly localized to the non-appressed regions of the thylakoid membrane. Expressed in leaves and stems, and at lower levels in flowers and siliques (at protein level). Slight reduction in rosette size, yellowish inflorescences and siliques and small chloroplasts with irregular morphology and no starch grains.