Endosomal E3 ubiquitin-protein ligase that regulates the recycling endosome pathway (PubMed:23353890). May play an indirect role in regulation of gene expression during embryonic mesoderm formation (PubMed:8462875). S-ubiquitinyl-[E2 ubiquitin-conjugating enzyme]-L-cysteine + [acceptor protein]-L-lysine = [E2 ubiquitin-conjugating enzyme]-L-cysteine + N(6)-ubiquitinyl-[acceptor protein]-L-lysine. Protein modification; protein ubiquitination. Embryonic visceral mesoderm and primordia of somatic musculature. Flies are viable and fertile. Truncated N-terminus.