Probable phosphodiesterase (PDE) that catalyzes the hydrolysis of cyclic diguanylate (c-di-GMP). Increases motility and decreases biofilm formation in vivo. cyclic di-3',5'-guanylate + 2 H2O = 2 GMP + 2 H(+) Transcripts are more abundant in biofilm cells than in planktonic cells.