ATP-dependent DNA helicase implicated in DNA replication, DNA repair and the maintenance of genomic stability. Acts as an anti-recombinase to counteract toxic recombination and limit crossover during meiosis. Regulates meiotic recombination and crossover homeostasis by physically dissociating strand invasion events and thereby promotes noncrossover repair by meiotic synthesis dependent strand annealing (SDSA) as well as disassembly of D loop recombination intermediates (PubMed:25516598, PubMed:25595823). Also plays a role in preserving the stability of 45S rDNA repeats (PubMed:27760121). ATP + H2O = ADP + H(+) + phosphate The PIP-box (PCNA interacting peptide) motif mediates the interaction with PCNA and localization to replication foci. Increased homologous recombination and replication defects (PubMed:25516598, PubMed:25595823). Growth inhibition due to defective cell proliferation. Hypersensitivity to DNA cross-link agents (PubMed:25595823). Loss of 45S rDNA repeats (PubMed:27760121). Belongs to the helicase family. RAD3/XPD subfamily.