Transcription factor involved in the expression of a broad class of genes including snRNAs. Required for sporulation and DNA-damage repair. Prevents the spreading of SIR silencing at telomeres and protects histone H4, but not H3, from deacetylation. Interacts with the TFIID subunit TAF7 and with acetylated histones H3 and H4. Phosphorylated by the casein kinase CK2 complex. Heterochromatin spreading downstream of the silent mating-type locus HMR. Present with 8100 molecules/cell in log phase SD medium.