May play an important role in formation and fusion of Golgi-derived vesicles during acrosome biogenesis. Interacts (via N-terminus) with TES (via LIM domain 2). Heterodimer with TES; the heterodimer interacts with ENAH to form a heterotrimer (PubMed:21278383). Interacts with ACTL9 (By similarity). Detected at the Golgi apparatus during acrosome biogenesis. Detected at the subacrosomal layer in round spermatids. Detected in sperm head and tail. Detected in testis. Detected at the acrosome of round spermatids (at protein level). Belongs to the actin family.