Involved in organization of the endomembrane system and is required for endoplasmic reticulum morphology and organelle distribution. May act by inhibiting the formation of PYK10 complex by binding to GLL23 and exporting it from the ER. Required for proper subcellular localization of myrosinase TGG2. Has no lipase or esterase activity. Interacts with the PYK10 complex and TGG2, but not with TGG1 or PEN2. Also found in ER bodies. Retained in the ER when mutated. Expressed throughout the seedling, rosette leaves, roots, inflorescence and imbibed seed, but not in pollen. Expressed during all developmental stages, with the highest accumulation in germinated seeds. Up-regulated by methyl jasmonate and upon pathogen infection. Defects in endoplasmic reticulum (ER) protein export and organizational defects in the ER, including aggregation of ER around the nucleus. Increased sensitivity to salt. Belongs to the 'GDSL' lipolytic enzyme family. Lacks the conserved active site 'GDSL' motif and has no lipase activity. Truncated N-terminus. The stop codon at position 16 is created by sequencing errors.