In hippocampal neurons, regulates dendritic and spine growth and synaptic transmission. Interacts with CTCF, MYO1C and with the transcriptional machinery, including RNA polymerase II and TBP. Cell-permeable protein. 22 hours after injection in the hippocampal area CA1, internalized by most cells at the injection site (By similarity). Localizes at the chromosome periphery during mitosis (PubMed:20813266). Belongs to the learning-associated protein family.