Citrate transporter responsible for loading citrate into xylem tissues, which helps facilitate iron transport to shoots (PubMed:12172022, PubMed:15310833, PubMed:17351051, PubMed:18826429). Mediates the citrate release in the apoplastic spaces during plant development allowing iron nutrition between symplastically disconnected tissues (PubMed:21742986). Expressed in roots in the pericycle and cells internal to the pericycle and surrounding the vascular tissue (PubMed:12172022, PubMed:15310833). Also expressed in seed and flower (PubMed:21742986). Expressed during embryogenesis and during the early stages of germination. Two-fold induction by iron deficiency. Not induced by aluminum (PubMed:12172022, PubMed:18826429). Induced by abscisic acid (ABA) (PubMed:24111973). Chlorotic (PubMed:12172022, PubMed:8754685, PubMed:18826429, PubMed:21742986). Constitutive expression of strategy I iron deficiency response and accumulation of iron, manganese and zinc in shoots. No reduction in aluminum tolerance (PubMed:12172022, PubMed:18826429). Accumulation of Mn, Cu, Zn and Mg in leaves and accumulation of Fe in roots (PubMed:8754685). Altered pollen development (PubMed:21742986). Belongs to the multi antimicrobial extrusion (MATE) (TC 2.A.66.1) family.