Functions in vesicle-trafficking events required for site-specific cell wall modifications during pollen germination and for anchoring of the cell plate to the mother wall at the correct cortical position. Interacts with CLC2 and CHC2. Localized in the forming cell plate and the cortical division zone (CDZ) during cytokinesis. Expressed at the pollen tube exit site in germinating pollen. Male sterility due to the production of shriveled pollen unable to germinate. Belongs to the TPLATE family.