RNA-binding protein that binds to misfolded non-coding RNAs, pre-5S rRNA, and several small cytoplasmic RNA molecules known as Y RNAs (By similarity). May play roles in cilia formation and/or maintenance (PubMed:21289087). Identified in a IGF2BP1-dependent mRNP granule complex containing untranslated mRNAs (By similarity). Found in a complex with PUF60 and Y5 RNA (By similarity). Interacts with RAB11FIP5 (By similarity). Localized in cytoplasmic mRNP granules containing untranslated mRNAs. Highest in brain, followed by lung, muscle, kidney and heart. Lower levels are found in testis, liver and spleen. The horseshoe-shaped TROVE domain is built with 7 helical HEAT-like repeats, and is closed by the VWFA-like domain giving rise to a ring-shaped monomer. Single-stranded RNA is bound in the positively charged central cavity (By similarity). The MIDAS-like motif in the VWFA-like domain binds divalent metal cations. Cilia absent or reduced, virtually no cilia of the normal 5 uM mean length. Belongs to the Ro 60 kDa family.