Required for sperm flagellum axoneme organization and function (PubMed:30686508). Involved in axonemal central pair complex assembly and/or stability (PubMed:30686508). Male mice are infertile, while female fertility is not affected (PubMed:30686508). Spermatozoa exhibit multiple morphologic abnormalities including short, thick, and/or coiled flagella, whereas sperm heads conserve an overall typical hooked shape (PubMed:30686508). Due to intron retention.