Component of the ceramide synthase complex that catalyzes the transfer of the acyl chain from acyl-CoA to a sphingoid base, with high selectivity toward hexacosanoyl-CoA (C26:0-CoA) (PubMed:15692566, PubMed:11694577). N-acylates sphinganine and phytosphingosine bases to form dihydroceramides and phytoceramides, respectively (PubMed:15692566, PubMed:11694577). Redundant with LAG1. Facilitates ER-to-Golgi transport of GPI-anchored proteins. a sphingoid base + a very long-chain fatty acyl-CoA = an N-(very-long-chain fatty acyl)-sphingoid base + CoA + H(+) hexacosanoyl-CoA + sphinganine = CoA + H(+) + N-hexacosanoylsphinganine eicosanoyl-CoA + sphinganine = CoA + H(+) + N-eicosanoylsphinganine a fatty acyl-CoA + sphinganine = an N-acylsphinganine + CoA + H(+) (4R)-hydroxysphinganine + a fatty acyl-CoA = an N-acyl-(4R)-4-hydroxysphinganine + CoA + H(+) Lipid metabolism; sphingolipid metabolism. Composed of the ceramide synthase complex composed of at least LAC1, LAG1 and LIP1. Expression is controlled by PDR1 and PDR3 which bind its propmoter PDRE element, and by ROX1 and CBF1. Present with 2840 molecules/cell in log phase SD medium. Belongs to the sphingosine N-acyltransferase family.