Aldehyde decarbonylase involved in the conversion of aldehydes to alkanes. Core component of a very-long-chain alkane synthesis complex. Involved in epicuticular wax biosynthesis and pollen fertility. a long-chain fatty aldehyde + H(+) + 2 NADPH + O2 = a long-chain alkane + formate + H2O + 2 NADP(+) Homodimer. Interacts with CER3, CYTB5-B, CYTB5-C, CYTB5-D and CYTB5-E. Expressed in seedlings, stems, leaves, flowers, fruits and siliques. Not detected in roots, pollen and seeds. Expressed in trichomes, cotyledons, shoot apical meristem and leaf primordia. Preferentially associated with young leaves rather than mature leaves. Expressed in the epidermis of the stem and caulines leaves, in the carpels and the sepals. Down regulated by cold and dark stresses. Up-regulated by low humidity, osmotic stress and abscisic acid treatment. No effet of methyl jasmonate, GA3, salicylic acid, cytokinin or auxin treatments. Glossy stem and fruit and reduced fertility due to the inability of the pollen grains to rehydrate on the stigma surface. Disappearance of the wax crystals. Decreased C29, C31 and C33 alkane contents. Increased susceptibility to soil water deficit. Belongs to the sterol desaturase family. Truncated C-terminus.