Putative transcription factor that acts as a key negative regulator of cell accumulation in shoot and floral meristems. Negatively regulates the size of the WUSCHEL (WUS)-expressing organizing center in inflorescence meristems. May act by down-regulating expression of WUS. Acts as an antirepressor that counteracts EMF1 action through modulation of trimethylated 'Lys-4' on histone H3 (H3K4me3) marks on target gene loci (including genes involved in salt stress response and flower development) (PubMed:23632855). Collaboratively with RBL and CYP40/SQN, influences floral meristem (FM) determinacy in an AGAMOUS and SUPERMAN-dependent manner, thus contributing to the floral developmental homeostasis (PubMed:18441215). Interacts with HHO5 (PubMed:26903506). Associates with ATX1 for trimethylating 'Lys-4' on histone H3 (H3K4me3) at flower MADS box gene loci (PubMed:23632855). A number of isoforms are produced. According to EST sequences. Expressed at low levels in seedlings, roots, shoots, leaves, stems, inflorescences, pollen, flowers and siliques, with highest levels dividing tissues including inflorescence. Expressed in embryonic shoot apical meristems, in inflorescence and floral meristems, and in developing stamens, carpels and ovules. Also expressed in vegetative meristems and leaf primordia. Delayed flowering and higher floral organs number with extra petals and sepals (PubMed:23632855). Increased H3K27me3 marks but decreased H3K4me3 marks on target gene loci (PubMed:23632855). Plants missing both EMF1 and ULT1 have rescued normal H3K27me3 marks and H3K4me3 marks (PubMed:23632855). Plants lacking both CRC and ULT1 exhibit strong floral meristem (FM) indeterminacy with reiterations of extra floral whorls in the center of the flower associated with reduced AGAMOUS and SUPERMAN levels (PubMed:18441215). It is uncertain whether Met-1 or Met-8 is the initiator. Truncated N-terminus.