Component of a cytoskeletal structure that is required for the formation of endocytic vesicles at the plasma membrane level. Plays an important role in virulence. Expression is induced in absence of HOG1 under non-stressed conditions. Leads to defects in actin polarization, endocytosis, bud morphogenesis, as well as increased sensitivity to calcofluor white and Congo red. Shows also a strong defect of hyphal morphogenesis and attenuated virulence.