RNA-directed RNA polymerase that replicates the viral (+) and (-) genome. a ribonucleoside 5'-triphosphate + RNA(n) = diphosphate + RNA(n+1) Forms a ribonucleoprotein complex with the viral RNA. Since the viral RNA is not encapsidated, ribonucleoprotein complex formation appears to be the strategy to survive in the host as persistent virus. The virus probably exists as a ribonucleoprotein viral particle and is transmitted via maternal inheritance through ascospores with a high rate of transmission through conidia.