Catalyzes the cross-linking of proteins and the conjugation of polyamines to proteins. L-glutaminyl-[protein] + L-lysyl-[protein] = [protein]-L-lysyl-N(6)-5-L-glutamyl-[protein] + NH4(+) Binds 1 Ca(2+) ion per subunit. Mainly expressed in hemocytes, hepatopancreas, and gastric tissues. On the other hand nothing was detected in the heart, intestine and muscle. Belongs to the transglutaminase superfamily. Transglutaminase family.