Protein tyrosine phosphatase that exhibits also lipid phosphatase activity. Can use phosphatidylinositol substrates such as PtdIns(3,4,5)P(3) as substrate. Pollen-specific phosphatase required for pollen development. H2O + O-phospho-L-tyrosyl-[protein] = L-tyrosyl-[protein] + phosphate a 1,2-diacyl-sn-glycero-3-phospho-(1D-myo-inositol-3,4,5-trisphosphate) + H2O = a 1,2-diacyl-sn-glycero-3-phospho-(1D-myo-inositol-4,5-bisphosphate) + phosphate Inhibited by vanadate. Expressed exclusively in pollen grains during the late stage of development (at protein level). Pollen cell death after mitosis, during the late stage of development, characterized by collapsed pollen grains, shrunken, with highly deformed exines and reduced size. Belongs to the PTEN phosphatase protein family.