Promotes nuclear translocation of the transcription factor OSH1, which directly suppresses the auxin biosynthetic gene YUCCA4 during the late development of anthers (PubMed:29915329). Reduction of auxin levels at late stage of anther development, after meiosis of microspore mother cells, is necessary for normal anther dehiscence and seed setting (PubMed:29915329). Required for jasmonate (JA) biosynthetic genes expression and JA production in anthers (PubMed:29915329). Interacts with OSH1. May be associated with the endoplasmic reticulum membrane in the close vicinity of the plasma membrane. Expressed in roots, stems, lemma, palea, pistils and ovules (PubMed:29915329). Expressed at low levels in leaves (PubMed:29915329). Highly expressed in developing anthers before pollen mitotic divisions from stage 8 to stage 12, with a peak of expression at stage 9 (at protein level) (PubMed:29915329). Expressed in developing seeds from 3 to 10 days after flowering (at protein level) (PubMed:29915329). Male sterility due to defect in anther dehiscence and mature pollen grain release at anthesis. Belongs to the MCTP family.