May act either alone or in interaction with glutaredoxin as a redox-regulated transcriptional regulator, or as a factor regulating Fe-S cluster biogenesis (Probable). The GRXS17-BOLA2 heterodimer binds a labile, oxygen sensitive iron-sulfur cluster (PubMed:24714563). Homodimer (PubMed:24203231). Interacts in vitro with GRXS14, GRXS15, GRXS16 and GRXS17, but not with GRXC5 (PubMed:24203231). Interacts in vivo only with GRXS17 (PubMed:24203231, PubMed:24714563). The putative nucleic acid binding region (34-77) coincides with the interaction surface with glutaredoxin. Can be either glutathionylated or forming covalent homodimers, depending on the oxidation state. Belongs to the bolA/yrbA family.