Transcription factor involved in the determination of epidermal cell identity (Probable). Required for correct morphological development and maturation of trichomes (PubMed:7926739). Regulates the frequency of trichome initiation and determines trichome spacing (PubMed:11844112). Acts as a negative factor for root hair development (PubMed:8620852, PubMed:8811855, PubMed:26486447). Required for ectopic repression of root hair development in a subset of epidermal cells (PubMed:8811855). May suppress hair formation in root epidermis by promoting differentiation into hairless epidermal cells (PubMed:8620852). Directly suppresses the bHLH transcription factor genes, RHD6, RSL1, RSL2, LRL1, and LRL2, which have diverse functions in root hair development (PubMed:26486447). Required for normal development of seed coat mucilage (PubMed:7926739, PubMed:21883555). Involved in the control of seed oil accumulation (PubMed:16514561, PubMed:21883555). Acts as a negative regulator of anthocyanin biosynthesis (PubMed:26017690). May directly repress the expression of some component genes from the MYB-bHLH-WD40 (MBW) transcriptional activator complex (PubMed:26017690). The MBW complex activates the transcription of late biosynthesis genes in the flavonoid pathway, leading to the production of anthocyanins (Probable). Interacts with GIR1 and GIR2. A number of isoforms are produced. According to EST sequences. Expressed in individual developing trichome cells of the emerging leaf primordia (PubMed:7926739). Expressed in differentiating hairless cells of root epidermis (PubMed:8620852). Down-regulated by GEM. Defect in trichome development (PubMed:7926739, PubMed:28526410). Defect in accumulation of seed coat mucilage (PubMed:7926739, PubMed:21883555). Excessive root hair formation (PubMed:8811855, PubMed:8620852, PubMed:28526410). High seed oil content phenotype (PubMed:16514561, PubMed:21883555). The gain-of-function gl2-1D mutants (T-DNA tagging) exhibit reduced endogenous levels of anthocyanins. Belongs to the HD-ZIP homeobox family. Class IV subfamily. Truncated N-terminus. Truncated N-terminus.