Essential for normal spermatogenesis. Testis specific. Expressed at the spermatid stage. Undetectable until 20 days post partum (dpp), and becomes detectable only after 22-30 dpp (PubMed:22110678). Expressed after postnatal day 15 and expression gradually increases with age (PubMed:29700843). Mice have smaller testis and show impaired spermatogenesis.