Component of the endomembrane trafficking machinery that is involved in protein recycling to the plasma membrane. Forms a complex with BIG5/MIN7/BEN1 and actin to modulate the function of the trans-Golgi network /early endosome at the intersection of the exocytic and endocytic pathways. May be linking post-Golgi traffic with the actin cytoskeleton. Interacts with BIG5. The C-terminal domain (496-565) is responsible for the localization to the trans-Golgi network/early endosome. Hypersensitivity to latrunculin B1 (LatB) and cytochalasin, reduced primary root length and root hair growth defects.