Acts as a non-catalytic accessory component of a dynein complex (By similarity). Part of a complex with bicd-1 and egal-1, which is recruited to the nuclear envelope by unc-83, where in turn, it recruits dynein to the nuclear surface and regulates nuclear migrations in hypodermal precursor cells (PubMed:20005871). Probably within a dynein motor complex, plays a role in the cell fate specification of the germline and oogenesis (PubMed:19752194, PubMed:27864381). In particular, it inhibits germ cell proliferation (PubMed:19752194). Regulates the function and localization of the RNA-binding protein fbf-2 in the germline (PubMed:27864381). Plays a role in mitotic and meiotic processes (PubMed:19752194, PubMed:26483555). Involved in the pairing of homologous chromosomes (PubMed:26483555). Independently of its dynein-mediated functions, plays a role in germ cell apoptosis (PubMed:24030151). Interacts with mett-10; the interaction is direct, and is required for the nuclear localization of mett-10 (PubMed:19752194). Component of a dynein-regulating complex composed of at least bicd-1, dlc-1 and egal-1 (PubMed:20005871). Interacts with egal-1 and unc-83 (PubMed:20005871). Interacts with fbf-2 (PubMed:27864381). Probably recruited to the nuclear envelope by unc-83 (PubMed:20005871). Localizes to perinuclear patches in the transition zone (PubMed:27864381). Localizes to P-granules in the mitotic region and transition zone (PubMed:27864381). Broadly expressed in tissues including the intestine, body wall muscles, germs cells, oocytes, the rectal valve and cells in the head. Expressed at all developmental stages from the one-cell embryo. RNAi-mediated knockdown results in viable animals, with progeny that have nuclear migration defects in the hyp7 hypodermal precursor cells at the L1 stage of larval development (PubMed:20005871). RNAi-mediated knockdown results in increased germ cell apoptosis, but does not affect somatic cell death (PubMed:24030151). RNAi-mediated knockdown results in reduced mett-10 accumulation in nuclei (PubMed:19752194). RNAi-mediated knockdown prevents fbf-2 localization to P-granules (PubMed:27864381). Furthermore, RNAi-mediated knockdown in a fbf-2 loss of function background results in sterility and masculinization of the germline where only sperm, and not oocytes, are produced (PubMed:27864381). RNAi-mediated knockdown in a dhc-1 loss of function (or195) mutant background results in failed homologous chromosome pairing during meiosis (PubMed:26483555). RNAi-mediated knockdown in a glp-1 gain of function sensitized mutant background results in disrupted meiotic and mitotic processes (PubMed:19752194). This in turn results in polyploid nuclei, that are larger in size and contain more DNA as compared to wild-type (PubMed:19752194). In addition, many oocytes from these animals also contain unpaired chromosomes (PubMed:19752194). Belongs to the dynein light chain family.