Binds actin and actin filament bundles in a Ca(2+)/calmodulin-insensitive manner, but is unable to sever, cap, and nucleate actin filament formation in vitro. Does not protect individual filaments from severing by VLN3 (AC O81645). A number of isoforms are produced. According to EST sequences. Expressed in all tissues examined. Mainly detected in the vascular tissue and the pericycle of roots and in the vasculature of leaves. Not expressed in the root cap. Belongs to the villin/gelsolin family.