Transcription factor that binds to the motif 5'-GTAACNT-3' in the promoter of target genes (e.g. DD11 and DD18) and promotes their expression within synergid cells (e.g. in the filiform apparatus) in ovules (PubMed:16214903, PubMed:17693534, PubMed:18410484, PubMed:17937500). Required for the formation of the filiform apparatus during synergid cell differentiation in the female gametophyte (PubMed:16214903). Involved in pollen tube guidance to the micropyle (PubMed:16214903, PubMed:17937500, PubMed:23093426). Localized to the nuclei of the synergid cells. Expressed at high levels in the synergid cells of the female gametophyte, and at lower levels in the endosperm of young seeds and the trichomes of young leaves and sepals. Expressed in cellularized but not uncellularized female gametophytes. Impaired in pollen tube attraction associated with a reduced cytoplasmic calcium burst during pollen tube tips growing (PubMed:17937500, PubMed:23093426). Altered expression of several ovule-specific genes (PubMed:17937500). Extended C-terminus.