Receptor-like protein kinase that controls pollen tube behavior by directing rupture at proper timing to release the sperm cell. ATP + L-seryl-[protein] = ADP + H(+) + O-phospho-L-seryl-[protein] ATP + L-threonyl-[protein] = ADP + H(+) + O-phospho-L-threonyl-[protein] Expressed in pollen, but not in pistils or seedlings. No effect on male fertility and pollen germination, but siliques slightly shorter. Anx1 and anx2 double mutants show defects in male gametophytes due to premature pollen tube rupture. Male paralog of FERONIA, a female factor expressed in synergid cells that controls pollen tube behavior. Named Anxur after the husband of the Etruscan goddess of fertility Feronia. Belongs to the protein kinase superfamily. Ser/Thr protein kinase family.