Involved in resistance to abiotic stress. Heterodimers (PubMed:29272523). Interacts with WIH1/CYSTM13 (PubMed:29272523). Mostly expressed in roots and flowers and, to a lower extent, in stems, siliques and leaves. Induced by heat in shoots (PubMed:29272523). Repressed in roots in response to drought, oxidation stress and salt (PubMed:29272523). Belongs to the CYSTM1 family.