Participates in the transport of viral genome to neighboring plant cells directly through plasmodesmata, without any budding. TGBp2 and TGBp3 are necessary for intracellular delivery of TGBp1-containing vRNPs to plasmodesmata. Can gate plasmodesmata and increase their size exclusion limit. To a lesser extent than TGB3, induces host actin cytoskeleton network thickening, which probably plays a major role in virus cell-to-cell movement (By similarity). Binds ssRNA in a sequence non-specific manner (PubMed:12093178). Interacts with movement protein TGB3 (PubMed:12093178, PubMed:15608333). TGB1-TGB3-TGB2 complex formation is enhanced by ATP hydrolysis (By similarity). Probably localizes to plasmodesmata-associated membrane compartments called peripheral membrane bodies (PMBs). Associates with host actin filaments, possibly for the intracellular transport to the plasmodesmata (By similarity). The viral replication sites are located at the host chloroplast membrane (PubMed:23269927). Belongs to the virgaviridae/benyvirus TGB2 movement protein family.