In hippocampal neurons, regulates dendritic and spine growth and synaptic transmission. Interacts with CTCF, MYO1C and with the transcriptional machinery, including RNA polymerase II and TBP. Cell-permeable protein. 22 hours after injection in the hippocampal area CA1, internalized by most cells at the injection site (By similarity). Localizes at the chromosome periphery during mitosis (By similarity). Belongs to the learning-associated protein family.