Required for incorporation of 9(10),16-dihydroxy-hexadecanoic acid into cutin. Expressed in root caps and lateral root emerging sites, in trichomes, in epidermis in stems, sepals and anther filaments, and in pollen grains and torpedo stage seeds. Expressed in all 3 inner integumenta layers and 2 outer integument layers in young seeds, but specifically restricted to the inner integumenta in mature seeds. Smaller rosettes, collapsed or tangled trichomes, postgenital fusions between the rosette leaves and flower buds, fusions between sepals, altered cuticle permeability, loss of cuticular folding in petals, abnormal flower and silique development, altered seeds surface, defective mucilage extrusion and semisterility. Increased susceptibility to salinity, osmotic and water deprivation stresses. Belongs to the plant acyltransferase family.