Involved in regulation of actin cytoskeleton organization and cytokinesis. Interacts with calmodulin in the absence of Ca(2+). In living cells localizes to the cell cortex and nucleus. In formaldehyde fixed cells localizes to the nucleolus. Expressed during vegetative growth. Expression decreases rapidly during early development and is not detected after 12 hours of development. Cells fail to complete the final step of cytokinesis and form multinucleated cells when grown on a solid surface. Null mutants show excessive accumulation of F-actin around the periphery of the ventral surface. When grown in suspension, mutants lacking dwwA complete cytokinesis normally. Mutants lacking the N-terminal C2 domain fail to localize to the cell cortex.