5,6-dihydrouracil + H2O = 3-(carbamoylamino)propanoate + H(+) Binds 2 Zn(2+) ions per subunit. Homotetramer. In L1-L2 larvae, expressed in body hypodermal cells, hemidesmosomes and in a neuronal cell between the pharynx and ring neuropil. In adults, expression is seen in body hypodermal cells and pharynx. Expressed in dorsal regions of embryos at late gastrula stage, transiently expressed in the developmental process of 3-fold embryo to L1-L2 larval stage. Carboxylation allows a single lysine to coordinate two zinc ions. Belongs to the metallo-dependent hydrolases superfamily. Hydantoinase/dihydropyrimidinase family.