Cross-links actin filaments (F-actin) in a calcium independent manner. Induces the formation of actin bundles. Stabilizes and prevents F-actin depolymerization mediated by latrunculin B (LatB). Interacts with F-actin. Expressed in mature pollen. Delayed pollen germination and inhibition of pollen tube growth due to the disorganization and redistribution of actin filaments. Cross-links actin with a constant of dissociation of 0.71 uM.