Involved in the TCA cycle. Catalyzes the stereospecific interconversion of fumarate to L-malate. (S)-malate = fumarate + H2O Carbohydrate metabolism; tricarboxylic acid cycle; (S)-malate from fumarate: step 1/1. Homotetramer. There are 2 substrate-binding sites: the catalytic A site, and the non-catalytic B site that may play a role in the transfer of substrate or product between the active site and the solvent. Alternatively, the B site may bind allosteric effectors. Belongs to the class-II fumarase/aspartase family. Fumarase subfamily. Truncated N-terminus.