Catalyzes the synthesis of dTDP-4-amino-4,6-dideoxy-D-galactose (dTDP-Fuc4N) from dTDP-4-keto-6-deoxy-D-glucose (dTDP-D-Glc4O) and L-glutamate. 2-oxoglutarate + dTDP-4-amino-4,6-dideoxy-alpha-D-galactose = dTDP-4-dehydro-6-deoxy-alpha-D-glucose + L-glutamate kcat is 0.38 sec(-1). Optimum pH is 7.5. Optimum temperature is 37 degrees Celsius. Bacterial outer membrane biogenesis; enterobacterial common antigen biosynthesis. Homotetramer. Mutants do not synthesize TDP-Fuc4NAc and enterobacterial common antigen (ECA). Belongs to the DegT/DnrJ/EryC1 family.