Redundantly with pde-1, plays a role in the AFD thermosensory neurons to regulate microvilli receptive ending morphology, possibly by regulating cGMP levels. a nucleoside 3',5'-cyclic phosphate + H2O = a nucleoside 5'-phosphate + H(+) Binds 2 divalent metal cations per subunit. Site 1 may preferentially bind zinc ions, while site 2 has a preference for magnesium and/or manganese ions. Belongs to the cyclic nucleotide phosphodiesterase family.