Component of the linker nucleocytoskeleton and cytoskeleton (LINC) complex that regulates telomere movement and meiotic recombination during meiosis (PubMed:17245108, PubMed:18585352, PubMed:32967926). Connects the spindle pole body with the nuclear envelope through its interaction with MPS2 and mediates meiotic bouquet formation and rapid chromosome movements in meiotic prophase (PubMed:16923827, PubMed:17495028, PubMed:22017544). Functions as an integral membrane anchor for telomeres and is a nuclear receptor for the SIR4 pathway of telomere tethering and gene inactivation (PubMed:18039933, PubMed:19390087). Essential for nuclear division and fusion and required for the first step of spindle pole body duplication in G1 (PubMed:12486115, PubMed:12493774). Functions in sister chromatid cohesion establishment (PubMed:15355977, PubMed:16682351). Recruits double-strand breaks (DSBs) to the nuclear periphery for chromosome healing (PubMed:19390086, PubMed:19217407, PubMed:20016273). Component of the linker nucleocytoskeleton and cytoskeleton (LINC) complex composed of at least MPS2, MPS3 and CSM4 (PubMed:32967926). Interacts with HTZ1 (PubMed:21518795). Interacts with REC8 (PubMed:30417519). Interacts with CSM4; ECO1, EST1, JEM1, CDC31, NDJ1, MPS2 and SIR4 (PubMed:12486115, PubMed:12493774, PubMed:15355977, PubMed:16923827, PubMed:17245108, PubMed:17495028, PubMed:18039933, PubMed:18585352). Localizes to the spindle pole body half bridge throughout the cell cycle (PubMed:15282802). Relocalizes from the spindle pole body to the nuclear envelope in meiosis, via the interaction with HTZ1 (PubMed:22017544, PubMed:21518795). The cohesin component REC8 promotes localization on nuclear envelope in mitotic cells (PubMed:30417519). The SUN domain is involved in the binding to MPS2 and is important for modulating chromosome motion events that act in meiotic chromosome juxtaposition and by extension, promoting proper morphogenesis of the synaptonemal complex. Phosphorylation at Ser-189 and Ser-190 by CDC7 and CDC28 during meiosis regulates of the localization on nuclear envelope for meiotic chromosome motion and nuclear envelope remodeling.