ATPase component of the chromatin remodeling INO80 complex which is involved in transcriptional regulation, DNA replication and DNA repair (By similarity). Binds DNA (By similarity). As part of the INO80 complex, remodels chromatin by shifting nucleosomes (By similarity). The INO80 complex controls ethylene-induced H2A.Z eviction dynamics (PubMed:31418686). Positive regulator of homologous recombination, but not an essential component of homologous recombination (PubMed:15525519). Not involved in the illegitimate repair pathway (PubMed:15525519). ATP + H2O = ADP + H(+) + phosphate Component of the INO80 chromatin-remodeling complex (By similarity). Associates with REF6/EIN6 (PubMed:31418686). A number of isoforms are produced. According to EST sequences. Not induced by methyl methanesulfonate (MMS) treatment. The DBINO region is involved in binding to DNA. Decreased homologous recombination frequency, but unchanged sensitivity to genotoxic agents and efficiency of T-DNA integration (PubMed:15525519). Differential expression of several genes (PubMed:31418686). Compromised ethylene-induced H2A.Z eviction dynamics (PubMed:31418686). The double mutant ref6-1 ino80-1 is insensitive to ethylene (ET) and exhibits reduced levels of EIN2 associated with a shift of the chromatin landscape to a repressive state at its locus (e.g. H3K27me3 and H2A.Z) (PubMed:31418686). Belongs to the SNF2/RAD54 helicase family.