May act as a negative regulator of ASF1 in chromatin assembly. Interacts with ASF1A and ASF1B. Found in a cytosolic complex with ASF1A, ASF1B, IPO4 and histones H3.1 and H4. Mainly detected as a cytoplasmic protein. Widely expressed in adult mice, the highest levels can be measured in erythropoietic cells. Expressed ubiquitously during early embryogenesis. Embryonic lethal; embryos die before 7.5 days of gestation.