Nucleolar protein which binds to RNA polymerase I and rDNA and is required for efficient RNA polymerase I-mediated rDNA transcription (PubMed:24722283). Maintains the epigenetically active status of rDNA chromatin which facilitates rDNA transcription and sustains germline development, ensuring fertility (PubMed:24722283). Plays a role in the modulation of nucleolus size (PubMed:27003693). May play a role in the regulation of lifespan (PubMed:11743719). May form dimers (PubMed:20405020). Interacts with RNA polymerase I (PubMed:24722283). Prior to fertilization, during the diakinesis stage of meiotic prophase, transcription ceases and mature oocytes harbor no nucleolus. During this time, localizes to nucleoli of transcriptionally active cells in the rest of the gonad. From the 2-cell stage of embryonic development, localizes to the nucleoplasm. Localizes to nucleoli upon their development in 6- to 8-cell stage embryos. Localizes to the nuclear periphery in developing embryos. In postembryonic cells, predominantly localizes to the nucleolus, but also localizes to small punctate structures throughout the nucleoplasm. Co-localizes with rDNA and fib-1 in nucleoli. Expressed in the nerve ring and hypodermal tissues (PubMed:24722283). Expressed in the intestine (PubMed:20405020, PubMed:24722283). Expressed in the germline (PubMed:20405020). Expressed throughout embryonic development and in adults (PubMed:22768349). During embryogenesis, expressed from the 2-cell stage and persists throughout development in transcriptionally active cells (PubMed:22768349). Expression is higher in dauer larvae than in adults (PubMed:11743719). Belongs to the NOLC1 family.