Regulates synaptic differentiation through the organization of actin cytoskeleton possibly by activating Rho-like GTPases. Is likely a factor in the cascade of Rac1 or Cdc42 in the neurons (PubMed:8999801). May play a role in maintaining proper septate junction functions. Required for eye development and most likely affects corneal lens-formation (PubMed:30290152). Localizes to the submembranous region of synaptic terminals. In the adult eye is expressed in the two primary pigment cells in the subapical region of the eye. Also present in photoreceptors. At stage 14, expression occurs in each segment of the central nervous system. At stage 17, expression becomes restricted to the synaptic regions of the brain and ventral nerve cord, where synapses undergo maturation (PubMed:8999801). At 45 hours after puparium formation (apf) expression is enriched in the shaft of bristle cells in the sub-apical region. Present in all photoreceptors (PubMed:30290152).