Probable component of ribonuclease P, a ribonucleoprotein complex that generates mature tRNA molecules by cleaving their 5'-ends (By similarity). May also be a component of the MRP ribonuclease complex, which cleaves pre-rRNA sequences (By similarity). Required for female gametophyte development and male competence (PubMed:22509260). Probable component of nuclear RNase P and RNase MRP ribonucleoproteins (By similarity). Interacts with POP5 (PubMed:22509260). Mostly expressed in inflorescence and roots, to a lower extent in leaves, and, at low levels, in siliques, seedlings and stems. Expressed in seedlings, young leaves, root tips and lateral primordia (PubMed:22509260). In reproductive organs, present in the inflorescence, especially in pollen grains, nucellar cells and embryo sacs (PubMed:22509260). Impaired progression of the gametophytic division during female gametogenesis leading to arrested embryo sacs at stages ranging from FG1 to FG7 (PubMed:22509260). Normal pollen development, but weaker pollen tube fitness associated with a reduced transmission through the male gametes (PubMed:22509260). Belongs to the eukaryotic/archaeal RNase P protein component 3 family.