Induces disassembly of actin filaments in conjunction with ADF/cofilin family proteins. Essential for organismal and cell viability. Required for the development of normal wing cell planar polarity. At the cell periphery of wing cells. Expressed in pupal wing cells. Pupals exhibit grossly abnormal epidermal hairs on wings, an abnormal accumulation of F-actin and microtubules, and disruption of the frizzled-based planar cell polarity system. Belongs to the WD repeat AIP1 family.