Putative lipid transfer protein required for systemic acquired resistance (SAR) long distance signaling. May interact with a lipid-derived molecule to promote long distance signaling associated with SAR. Together with AZI1, required for glycerol-3-phosphate- (G3P) and azelaic acid- (AA) induced systemic acquired resistance (SAR). Component of plant systemic immunity involved in priming defenses in a AA-dependent manner, by modulating production and/or translocation of a mobile signal(s) during SAR. Is able to bind with high affinity monoacylated phospholipids, mainly lysophosphatidylcholines (PubMed:18552128). Binds 1 zinc ion per subunit. Self-interacts and binds to AZI1 (PubMed:23602565). Does not interact with PDLP1 (PubMed:27078071). Induced by glycerol-3-phosphate (G3P). No visible phenotype under normal growth condition, but compromised pathogen-induced glycerol-3-phosphate-(G3P) and azelaic acid- (AA) dependent systemic acquired resistance (SAR). Belongs to the A9/FIL1 family.