Essential for normal crossover (CO) formation during meiosis (PubMed:30589140, PubMed:31266799). Essential component for the formation of class I meiotic COs (PubMed:31266799). Interacts with SHOC1, another meiotic component, to regulate CO formation, possibly by stabilizing the recombination intermediates during meiosis (PubMed:30589140, PubMed:31266799). PTD and SHOC1 may form transient heterotrimeric or heterotetrametric complexes with HEI10 and/or ZIP4 to promote class I COs formation (PubMed:31266799). Does not seem to be involved in early meiotic recombination steps involving double-strand break (DSB) formation, processing, and single-strand invasion (PubMed:31266799). Does not seem to be involved in homologous pairing or synaptonemal complex (SC) assembly (PubMed:31266799). Interacts with SHOC1 (via C-terminus) (PubMed:30589140, PubMed:31266799). Interacts with HEI10 (PubMed:31266799). Predominantly localized in the nucleus (PubMed:30589140). Localized in punctuate foci onto meiocyte chromosomes from leptotene to early pachytene with a maximal number of foci at zygotene (PubMed:31266799). Highly expressed in anthers and pistil during meiosis (PubMed:30589140). Expressed in pollen mother cells (PMCs) during meiosis (PubMed:30589140). Expressed at low levels in roots, shoots, leaves, flowers, and glumes (PubMed:30589140). Complete sterility of male and female gametophytes due to altered meiosis (PubMed:30589140, PubMed:31266799). Striking reduction in the number of meiotic crossovers (PubMed:30589140, PubMed:31266799). Belongs to the ERCC1/RAD10/SWI10 family.