Vacuolar iron transporter involved in the transfer of iron ions from the cytosol to the vacuole for intracellular iron storage (PubMed:22731699). Vacuolar iron storage is required for seed embryo and seedling development (Probable). May be involved in the regulation of iron translocation between flag leaves and seeds (Probable). Can transport zinc ions from the cytosol to the vacuole (PubMed:22731699). Homodimer. The dimeric interaction is mediated by both the transmembrane domains (TMDs) and the cytoplasmic metal binding domain (MBD). Localizes to the tonoplast. Expressed in leaf sheaths and at lower level in leaf blades. Induced by treatment with iron in roots and shoots (PubMed:22731699). Down-regulated under iron deficiency in roots and shoots (PubMed:22731699). Down-regulated by treatment with zinc in roots and shoots (PubMed:22731699). The cytoplasmic metal binding domain (MBD) is located between transmembrane 2 (TM2) and transmembrane 3 (TM3). No visible phenotype under normal growth conditions, but mutant embryos exhibit higher levels of iron and zinc, and leaf blades of mutant plants show decreased levels of iron and zinc. Can mediate sequestration of iron ions into vacuoles when expressed in the yeast ccc1 mutant (PubMed:22731699). Can mediate zinc ions sequestration into vacuoles when expressed in the yeast zrc1 mutant (PubMed:22731699). Belongs to the CCC1 family.