Prevents homeodomain proteins (e.g. STM) association to plasmodesmata and, consequently, cell-to-cell transport. Binds to RNA. Alters STM RNA binding capacity (PubMed:17965274). Regulates cytoskeleton (e.g. actin) organization that determinates cell shape (PubMed:8893552, PubMed:19074626). Regulates stomata patterning and drought tolerance (PubMed:19074626). Involved in restricting tobamovirus (e.g. oilseed rape mosaic virus) infectivity, probably by interfering with cell-to-cell virus movement (PubMed:19074626). Interacts with STM. Microtubule-associated and stabilizer (PubMed:17965274, PubMed:19074626). Localized in cytosolic punctae when associated with STM (PubMed:17965274). Expressed in seedlings, roots, flowers and developing ovules. In young seedlings, expressed in the vascular tissues of cotyledons, leaf primordia, and shoot apical meristem (SAM). In roots, confined to a limited number of epidermal cells in proximity to the root apical meristem. In flowers, present in carpels and ovules. In developing ovules, observed in the funiculus and integuments. Belongs to the microtubule binding protein 2C family.