Metalloendoprotease which functions in fertility and memory formation (PubMed:24395329, PubMed:27629706). Required in the dorsal paired medial neurons and alpha/beta mushroom body neurons for the proper formation of long-term and middle-term memories (PubMed:27629706). Required in males to maximise egg-laying in female mates and is also required in females for their fertility (PubMed:24395329). Preferential cleavage of polypeptides between hydrophobic residues, particularly with Phe or Tyr at P1'. Binds 1 zinc ion per subunit. Expressed in the testicular tube, near and in the seminal vesicles. In adults and third-instar larvae, expressed in the midgut and in the mushroom bodies of the brain and neurons in the pars intercerebralis. Also expressed in neurons of the ventral ganglion and imaginal disks (wing and leg) of third-instar larvae. In stage 17 embryos, expressed in the peripheral nervous system, pharynx and midgut. RNAi-mediated knockdown females lay fewer eggs and display a reduced hatch rate when mated to wild-type males, and wild-type females lay fewer eggs when mated to RNAi-mediated knockdown males (PubMed:24395329). RNAi-mediated knockdown in the dorsal paired medial neurons impairs middle-term (MTM) and long-term memory (LTM), but has no effect on normal aversion learning and anesthesia-resistant memory (ARM) (PubMed:27629706). RNAi-mediated knockdown in alpha/beta mushroom body neurons impairs MTM and LTM (PubMed:27629706). RNAi-mediated knockdown in all mushroom body neurons has no effect on learning and ARM (PubMed:27629706). Belongs to the peptidase M13 family.