Probable tRNA nucleus export receptor which regulates tRNA processing and facilitates tRNA translocation across the nuclear pore complex. Is required for correct leaf initiation at different developmental stages and may play a role in floral patterning. Shuttles between the nucleus and the cytoplasm. Expressed in roots, stems, leaves, flowers and embryos. Reduced size in non-mature plants, delayed panicle heading, reduced seed number and low seed fertility. Belongs to the exportin family.