Required during ciliogenesis for tubulin glutamylation in cilium. Probably acts by participating in the transport of TTLL6, a tubulin polyglutamylase, between the basal body and the cilium. Found in a complex with TTLL6. Localizes mainly to the cilium basal body and in primary cilia. At 10-13 dpc, embryos show a range of phenotypes, such as malformed hindbrain, exencephaly, brain hemorrhage, dilated pericardial sac and lethality. For an unclear reason, some homozygous mutants develop normally, suggesting presence of extragenic phenotypic modifiers. Belongs to the CEP41 family.