Hydrolyzes glycerol-phospholipids at the terminal phosphodiesteric bond to generate phosphatidic acids (PA). Plays an important role in various cellular processes, including phytohormone action, vesicular trafficking, secretion, cytoskeletal arrangement, meiosis, tumor promotion, pathogenesis, membrane deterioration and senescence. Can use phosphatidylserine or N-acylphosphatidylethanolamine as substrates. a 1,2-diacyl-sn-glycero-3-phosphocholine + H2O = a 1,2-diacyl-sn-glycero-3-phosphate + choline + H(+) Ca(2+). Requires micromolar level (PIP2-dependent). Inhibited by neomycin. Expressed in stems, and to a lower amount in leaves, flowers and siliques. Activated by wounding, methyl jasmonate, heavy metal, osmotic and salt stresses. Up-regulated by phosphate limitation (PubMed:16384909). C2 domain is a calcium-binding fold, and the binding promotes the protein association with membranes. In PLD beta, all the calcium-coordinating acidic amino acids are conserved. Belongs to the phospholipase D family. C2-PLD subfamily.