Plays a role in the cell cycle. Involved in a late stage of septum formation leading to the separation of the daughter cells. Involved in the correct orientation of forespore membrane extension during sporulation. Binds phosphatidylinositol 4-phosphate. Component of the septin complex composed of two copies of each spn1, spn2, spn3 and spn4. Component of the sporulation-specific septin complex composed of at least spn2, spn5, spn6 and spn7. Localizes to the medial ring at the cell cortex of dividing cells. The sporulation-specific septin complex associates to the forespore membrane and forms partial or complete ring-like structures that curl around each haploid nucleus. Belongs to the TRAFAC class TrmE-Era-EngA-EngB-Septin-like GTPase superfamily. Septin GTPase family.