Plays a role in the trafficking of mitochondria along microtubules. Regulates the kinesin-mediated axonal transport of mitochondria to nerve terminals along microtubules during hypoxia. Participates in the translocation of TRAK2/GRIF1 from the cytoplasm to the mitochondrion. Also plays a role in steroidogenesis through maintenance of mitochondrial abundance and morphology (By similarity). Plays an inhibitory role during neocortex development by regulating mitochondrial morphology, distribution and motility in neocortical neurons (By similarity). Interacts with RHOT1/Miro-1, RHOT2/Miro-2, TRAK1/OIP106 and TRAK2/GRIF1. Colocalizes with RHOT1, RHOT2, TRAK1 and TRAK2 at the mitochondrion. Extended N-terminus.