Protein-lysine acyltransferase that catalyzes fatty acylation of the protoxin, thereby converting it to the active toxin. a fatty acyl-[ACP] + L-lysyl-[protein] = H(+) + holo-[ACP] + N(6)-(fatty acyl)-L-lysyl-[protein] Homodimer. Belongs to the RTX toxin acyltransferase family.