Plays a role in nuclear pore complex (NPC) assembly and maintenance (PubMed:20547758). Required for nuclear import of Mad (PubMed:20547758). Mediates the association between the nuclear pore complex and a subset of active chromatin regions adjacent to lamin-associated domains (PubMed:31784359). Plays a role in double strand break repair by relocalizing the heterochromatic double strain breaks (DSBs) to the nuclear periphery as part of the homologous recombination (HR) repair process (PubMed:26502056). Regulates cytokinesis during spermatocyte meiosis by maintaining type-B lamin Lam localization to the spindle envelope (PubMed:27402967). Regulates female gonad development and oogenesis (PubMed:26485283). Part of the nuclear pore complex (NPC). Located on both the cytoplasmic and nuclear sides of the NPC core structure (By similarity). During syncytial embryo and larval neuroblast mitosis localizes to the nuclear envelope in interphase, accumulates in the nucleus in prometaphase, localizes to the spindle envelope in metaphase and then to condensing chromatin in telophase (PubMed:18562695). In spermatocytes, detected in the nuclear matrix by the nuclear envelope in metaphase I, and in the spindle envelope in premeiotic nuclei and during anaphase I (PubMed:27402967). Colocalizes with type-B lamin Lam throughout meiosis I (PubMed:27402967). Unlike in mammals, does not localize to kinetochore (PubMed:18562695, PubMed:27402967). Can localize to the nuclear lumen proximal to the inner nuclear membrane (PubMed:31784359). Expressed in spermatocytes (at protein level). Expressed in embryos, larvae and adults (at protein level) (PubMed:18562695, PubMed:31626769). Expressed in embryonal and larval neuroblasts (at protein level) (PubMed:18562695). Expressed during oogenesis (at protein level) (PubMed:31626769). Embryonic lethal (PubMed:18562695, PubMed:26485283). RNAi-mediated knockdown in somatic gonadal cells results in female, but not male, infertility associated with extensive disintegration of the egg chambers and cell death in nurse cells (PubMed:26485283). RNAi-mediated knockdown in spermatocytes causes abnormal localization of nuclear type-B lamin Lam, abnormal spindle envelope integrity and overall defective cytokinesis in meiosis (PubMed:27402967). Belongs to the nucleoporin Nup84/Nup107 family.