Putative transcription factor that acts as a key negative regulator of cell accumulation in shoot and floral meristems. Negatively regulates the size of the WUSCHEL (WUS)-expressing organizing center in inflorescence meristems. May act by down-regulating expression of WUS. Can compensate for mutant ULT1 protein when overexpressed. Expressed in influorescence, pollen and siliques, with a higher expression in influorescence. Specifically expressed during the reproductive developmental stage. Expressed in embryonic shoot apical meristems, in inflorescence and floral meristems, and in developing stamens, carpels and ovules.