Binds actin and actin filament bundles in a Ca(2+)-insensitive manner, but caps the barbed end of actin filaments and is able to sever them in a calcium-dependent manner. Involved in root hair growth through regulating actin organization in a Ca(2+)-dependent manner. Preferentially expressed in vegetative tissues. Detected in the whole seedling, hypocotyl, cotyledon, primary root, roots hair cells and trichomes. Expressed in flowers but not in the silique. No changes in the primary root length, but shorter root hairs. Increased sensitivity to latrunculin B (LatB) and instability of actin filaments. Belongs to the villin/gelsolin family.