Influences germ cell fate in hermaphrodites (PubMed:24098152). Acts downstream of tra-2 and tra-3 and through the Tip60 histone acetyltransferase complex to regulate germ cell fate decisions (PubMed:24098152). Required for spermatogenesis and embryonic development (PubMed:24098152). Acts with tra-2 to promote expression of fog-3 and control male tail development (PubMed:24098152). Involved in the negative regulation of vulval development (By similarity). Localized to condensed chromosomes during pachytene and diakinesis stages of meiosis I in germ cells. Null mutants display a feminization of the germ line (Fog) phenotype producing oocytes instead of sperm. Belongs to the PI3/PI4-kinase family. TRA1 subfamily.