May play an important role in formation and fusion of Golgi-derived vesicles during acrosome biogenesis. Interacts (via N-terminus) with TES (via LIM domain 2). Heterodimer with TES; the heterodimer interacts with ENAH to form a heterotrimer. Interacts with ACTL9. Detected at the Golgi apparatus during acrosome biogenesis (PubMed:21278383). Detected at the subacrosomal layer in round spermatids (PubMed:21278383). Detected in sperm head and tail (PubMed:21278383). Detected in testis. Detected at the acrosome of round spermatids (at protein level). Detected in adult and embryonic testis. Detected in developing male germ cells. Belongs to the actin family.