Accessory component of the DNA polymerase epsilon complex (By similarity). Participates in DNA repair and in chromosomal DNA replication (By similarity). Has a role in cell cycle progression (PubMed:19150446). Required for wing morphogenesis (PubMed:19150446). Component of the DNA polymerase epsilon complex consisting of four subunits: the catalytic subunit PolE1/DNApol-epsilon255 and the accessory subunits PolE2/DNApol-epsilon58, Chrac-14/DNApolE3 and PolE4/Mes4. Expressed throughout all developmental stages (PubMed:19150446). In the third instar larva expressed in fat bodies, brain lobes, eye-antenna imaginal disks, wing imaginal disks, salivary glands and ovaries (PubMed:19150446). RNAi-mediated knockdown results in an atrophied wing phenotype characterized by increased induction of cell proliferation in wing disks but failed cell cycle progression.