Beta-D-glucosidase active on scopolin >> esculin >> 4-MU-glucoside > DIMBOA-glucoside. No activity with pNP-glucoside, oNP-glucoside and sinigrin as substrates. Hydrolysis of terminal, non-reducing beta-D-glucosyl residues with release of beta-D-glucose. Activated upon binding to PBP1 or PBP2. Component of the PYK10 complex, at least composed of PYK10/BGLU23, BGLU21, BGLU22, JAL22, JAL23, PBP1/JAL30, PBP2/JAL31, JAL32, JAL33, JAL34, JAL35, GLL22 and GLL23. Expressed exclusively in roots. Up-regulated by cold, 2,4-D, methyl jasmonate and phosphate starvation. May be due to a competing acceptor splice site. Belongs to the glycosyl hydrolase 1 family.