Negative regulator of pollen germination (PubMed:27468890). Prevents pollination in moist environments by inhibiting jasmonic acid synthesis (PubMed:27468890). Stabilizes pollen tube growth (PubMed:27468890). Interacts with TCP4. The nuclear protein is functional. Expressed specifically in pollen (PubMed:27468890). Not detected in roots, stems or leaves (PubMed:27468890). Detected first in bicellular pollen and then most abundant in tricellular and mature pollen and growing pollen tubes. No visible phenotype under normal growth conditions, but hyperactive pollen germination.