Regulates the mitotic activity in roots. Plays a role with HSP70-1 in facilitating WIT1 nuclear envelope targeting. Binds to FPP proteins (By similarity). Interacts with WAP, WIP1, WIP2 and WIP3 through its WPP domain. Interacts with HSP70-1, HSP70-3 and WIT1. Component of a ternary complex composed of WPP1, HSP70-1 and WIT1. Associated to the nuclear envelope (NE) in undifferentiated cells of the root tip. Associated with the outer NE and the nuclear pores in interphase cells and with the immature cell plate during cytokinesis. In differentiated cells, localized in both cytoplasm and nucleus. Accumulate in speckles of the cytoplasm belonging to the Golgi apparatus (By similarity). Appears at the NE as cells reenter the cell cycle during dedifferentiation. Expressed in roots, stems and leaves. The WPP domain is required for the nuclear envelope localization.