Required for the production of double-strand breaks (DSBs) in meiotic recombination initiation; not required for mitosis. Plays a role in activation of the DSB nuclease SPO11 at recombination hotspots. Prevents accumulation of meiotic double strand break intermediates and is needed for the proper timing of the first meiotic division. Component of the MER2-MEI4-REC114 complex which seems to be required for meiotic double-strand break (DSB) formation. Interacts with MEI4, MER2 and REC104. Component of the MER2-MEI4-REC114 complex. Localizes to chromosomes. Meiosis-specific. Belongs to the REC114 family.