Cargo-specific adapter that is involved in nuclear migration during development and thereafter (PubMed:11748140, PubMed:19605495, PubMed:20005871, PubMed:20921138, PubMed:27697906). Component of the unc-83-unc-84 LINC (LInker of Nucleoskeleton and Cytoskeleton) complex where it interacts with unc-84 to form a bridge connecting the nuclear envelope to the cytoskeleton which allows for nuclear transport along microtubules (PubMed:11748140, PubMed:16481402, PubMed:25023515). Within the complex, connects the nuclear envelope to the microtubule cytoskeleton through the kinesin-1 light chain protein klc-2 (most likely within the Kinesin 1 motor complex) to regulate nuclear migrations (PubMed:19605495, PubMed:20921138). Moreover, within the complex, also recruits the large microtubule-associated bicd-1-dlc-1-egal-1 and lis-1-nud-2 complexes to the nuclear envelope to regulate both the bidirectional migration of nuclei and the extent of nuclear migrations (PubMed:20005871). Not required for centrosome attachment to the nucleus (PubMed:11748140). Component of the unc-83-unc-84 LINC complex which contains at least unc-83 and unc-84 (PubMed:11748140, PubMed:16481402). Within the unc-83-unc-84 LINC complex interacts with unc-84 (via C-terminus); the interaction is probably required to recruit unc-83 to the nuclear envelope where it then recruits dynein and kinesin-1 complexes to regulate nuclear migration (PubMed:11748140, PubMed:16481402). Interacts with bicd-1 and dlc-1 (PubMed:20005871). Interacts with nud-2 (via C-terminus); the interaction is direct, and is required for recruitment of nud-2 to the nuclear envelope (PubMed:20005871). Interacts with klc-2; the interaction is direct (PubMed:19605495). The transmembrane domain associates with the nuclear envelope (PubMed:11748140). Co-localizes with unc-84 and lmn-1 at the nuclear envelope (PubMed:11748140). Predominantly expressed in migratory nuclei (PubMed:11748140). Expressed in a variety of cell-types, including cells around the pharynx and in the uterus (PubMed:11748140). Expressed in embryos and adults (PubMed:11748140). First expressed at the nuclear envelope of migrating hyp7 nuclei, then, at the bean embryonic stage, it is expressed in hyp7 cells, P cells and intestinal cells (PubMed:11748140). The KASH domain, which contains a transmembrane domain, mediates nuclear envelope targeting. Viable, but with failed nuclear migrations in hyp7 hypodermal precursor cells (PubMed:11748140, PubMed:16481402, PubMed:19605495, PubMed:20005871, PubMed:20921138). Hyp7 nuclei are mislocalized to the dorsal cord of L1 stage larvae (PubMed:19605495).