E3 ubiquitin-protein ligase that is required for the plant C/N response during seedling growth transition. May be involved in the early steps of the plant defense signaling pathway. S-ubiquitinyl-[E2 ubiquitin-conjugating enzyme]-L-cysteine + [acceptor protein]-L-lysine = [E2 ubiquitin-conjugating enzyme]-L-cysteine + N(6)-ubiquitinyl-[acceptor protein]-L-lysine. Protein modification; protein ubiquitination. Up-regulated by chitin. Down-regulated by abscisic acid (ABA). The RING-type zinc finger domain mediates binding to an E2 ubiquitin-conjugating enzyme. Hypersensitivity to C/N conditions during post-germinative growth. Belongs to the RING-type zinc finger family. ATL subfamily.