Sequentially catalyzes two glycosylation steps at the 5-OH and 3-OH positions of anthocyanidin. Unglycosylated anthocyanidin or anthocyanidin 5-O-glucoside, but not anthocyanidin 3-O-glucoside, can be used as glucosyl acceptor. Pigment biosynthesis; anthocyanin biosynthesis. In roses, anthocyanidin 3,5-O-diglucoside is the first stable anthocyanin, and therefore responsible for flower coloration, whereas this is usually anthocyanidin 3-O-glucoside in other plants. Belongs to the UDP-glycosyltransferase family. The color of the rose - Issue 63 of October 2005