Plays a major role in the promotion of the transition of the vegetative meristem to reproductive development (By similarity). Required for RNA-mediated chromatin silencing of a range of loci in the genome. Cotranscriptionally recognizes aberrant RNA and marks it for silencing. Controls alternative cleavage and polyadenylation on pre-mRNAs and antisense RNAs (By similarity). Regulates flowering time, seed size and cell volume, probably via the modulation of cell size (PubMed:17597396). Interacts with FY (By similarity). Binds to SF1, FIK, RPRD1B, OsI_31983 and MADS8 (By similarity).