Transcriptional corepressor involved in branch formation regulation, presumably by suppressing primary branch formation and promoting secondary branch formation (PubMed:22136599, Ref.7, PubMed:24336200). Required for the cell elongation in the first internode and pollen development (PubMed:22020753). Probable downstream regulator of strigolactones signaling important in axillary meristem maintenance (PubMed:22136599, Ref.7, PubMed:24336200). Acts in auxin signaling and is associated with the regulation of histone deacetylation (PubMed:22136599). Essential for the function of miR172 microRNA and its target genes in regulating panicle development (PubMed:26631749). Tetramer (PubMed:26601214). Interacts with D53, probably via the EAR motifs (PubMed:24336200). Binds to AP2-1/TOE1, AP2-3/SNB and AP2-2/IDS1 (PubMed:26631749). Interacts with WOX1 (PubMed:25697101, PubMed:24336200, PubMed:26631749, PubMed:26601214). Interacts with MOF1 (PubMed:32680975). Expressed in stems and panicles. Detected in roots, seeds, leaves and sheath (PubMed:22020753). Expressed in the meristem and lateral organ primordia (PubMed:22136599). Highest expression in endosperm at 7 days after pollination. The N-terminal TOPLESS domain (TPD) (1-210) binds directly to a 12-amino acid LxLxL EAR motif peptide. Semi-dwarf, with reduced male fertility and shorter panicles with arrested branches and abnormal spikelets.