Component of the PAF1 complex which is a multifunctional complex involved in transcription initiation via genetic interactions with TATA-binding proteins, elongation and transcription-coupled histone modification. Component of the PAF1 complex which consists of at least cdc-73, ctr-9, leo-1, pafo-1 and rtfo-1. Nuclear localization depends on ctr-9, pafo-1 and cdc-73. Located in nuclei before mitosis. Located in the cytoplasm upon mitotic nuclear membrane breakdown. Located in nuclei when nuclear envelope is reassembled in telophase. Expressed in both somatic cells and germ cells from the one-cell stage onwards. RNAi-mediated knock-down is maternal effect embryonic lethal. Embryogenesis proceeds more slowly, with embryos displaying defects in the positioning and shape of epidermal cells. Belongs to the LEO1 family.