Required for chromosome segregation during meiosis (PubMed:16394097). During diakinesis and prometaphase I, essential for the formation of class I meiotic crossovers and homologous recombination (PubMed:16394097, PubMed:21771883, PubMed:24656236). Interacts with SHOC1. Expressed in roots, stems, leaves, siliques, flowers, inflorescence and anthers. Strongly expressed in meiocytes. Expressed in the shoot apical meristem (SAM) and young flower buds. In mature flowers, restricted to the reproductive organs stamens and carpels. In the stamen, accumulates in male meiocytes and in tapetal cells, as well as in mature pollen grains. In the carpel, present in developing ovules and in developing embryos. Reduction in the number of class I crossovers (COs) during meiosis (PubMed:21771883, PubMed:16394097, PubMed:24656236). Delay in the formation of synaptonemal complexes (SC) as well as a few unsynapsed axial elements in early pachytene nuclei in some meiocytes. Early nodules (ENs) are observed on the central element of SC. Reduced fertility due to altered male and female meiosis (PubMed:16394097). Belongs to the ERCC1/RAD10/SWI10 family.