Involved in resistance to abiotic stress. Heterodimers (PubMed:29272523). Interacts with CYSTM7 and WIH1/CYSTM13 (PubMed:29272523). Mostly expressed in stems and,at low levels, in stems, roots, flowers, siliques and leaves. Induced by heat and oxidation stress in shoots (PubMed:29272523). Induced in roots in response to cold and salt (PubMed:29272523). Belongs to the CYSTM1 family.