Influences germ cell fate in hermaphrodites (PubMed:24098152). Acts downstream of tra-2 and tra-3 and through the Tip60 histone acetyltransferase complex to regulate germ cell fate decisions (By similarity). Required for spermatogenesis and embryonic development (By similarity). Acts with tra-2 to promote expression of fog-3 and control male tail development (By similarity). Involved in the negative regulation of vulval development (PubMed:15068795). Localized to condensed chromosomes during pachytene and diakinesis stages of meiosis I in germ cells. Expressed in germ cells and somatic cells. Expressed throughout development from one-cell stage embryos onwards. RNAi-mediated knockdown results in partial sterility and the production of oocytes by 14% of males at 20 degees Celsius and by 23% of males at 25 degrees Celsius (PubMed:15068795, PubMed:24098152). At 25 degrees Celsius, 8% of males produce no mature germ cells and hermaphrodites develop into females (PubMed:24098152). Belongs to the PI3/PI4-kinase family. TRA1 subfamily.