A structure-specific endonuclease that resolves Holliday junction (HJ) intermediates during genetic recombination. Acts only on 4-way DNA junctions in a sequence non-specific manner; introduces paired nicks in opposing strands 2 bases 3' of the point of strand exchange only on continuous strands of 4-way junction DNA. Cleaves both mobile and immobile junctions. Endonucleolytic cleavage at a junction such as a reciprocal single-stranded crossover between two homologous DNA duplexes (Holliday junction). Binds 1 Mg(2+) ion per subunit. kcat is 4.8 min(-1) and 10 min (-1) at 5 mM and 15 mM MgCl2 respectively. Optimum temperature is 60 degrees Celsius. Could be higher with a more thermally stable substrate. Homodimer. A second Holliday junction resolving enzyme, Hjc, with different substrate specificity exists in this organism. Belongs to the Holliday junction resolvase Hjc family. Hje subfamily.