Involved in peroxisome biogenesis and matrix protein import (PubMed:24510720). Required for pollen maturation and in vivo germination via its role in peroxisomal function, which partially involves jasmonic acid biosynthesis (PubMed:24510720). Transported to peroxisomes via the interaction with PEX19-1 (PubMed:21487094). Required for peroxisomal protein import by acting as an anchoring protein for the AAA ATPase complex, which consists of PEX1 and PEX6 (PubMed:21487094). Interacts with PEX6 and PEX19-1, but not with PEX1 (PubMed:21487094). Interacts (via N-terminus) with PEX13, and (via N-terminus and C-terminus) with PEX16 (PubMed:24510720). Expressed in roots, leaves, stems, flowers, buds and fruits. Not detected in microspores before and at stage of pollen mitosis I. Expressed in bicellular, tricellular and mature pollen grains. Embryo lethality, when homozygous.