RNA-binding protein. Might be involved in alternative splicing regulation through an interaction with G-quartet RNA structure. When splicing or transcription are inhibited, accumulates in large, rounded speckles and in the nucleolus. Highly expressed in the hippocampus, the piriform cortex, Purkinje cells and the cingulate gyrus. Expressed before day 7 in the embryo and reached its highest levels at 10.5-11.5 days. In the embryo at day 11, expression is more specific in the roof of the hind brain and the lateral ventricle of the brain. Belongs to the AF4 family.