Involved in oxygen transport from the gills to the various peripheral tissues. Heterotetramer of two alpha chains and two beta chains. Red blood cells. This fish has two types of hemoglobin: one cathodic Hb and two major anodic Hbs. The cathodic Hb displays a small normal Bohr effect and a reverse Bohr effect in the presence and absence of phosphate respectively. In addition, the cathodic HB displays a large phosphate effect. Belongs to the globin family.