Microtubule-binding protein required for ciliogenesis. May function in ciliogenesis by mediating the transport of proteins like BBS4 to the cilium, but also through the organization of the centriolar satellites (By similarity). Plays a role in retina morphogenesis and/or homeostasis (By similarity). Homodimer; disulfide-linked (By similarity). Interacts with CEP290 (By similarity). Interacts with PCM1 (By similarity). Interacts with ARMC9, TOGARAM1, CSPP1 and CEP104 (By similarity). Restricted to the centrosomes and the spindle microtubules during mitosis. Enriched in the inner segment of the photoreceptor.