Required for DNA double-strand breaks (DSBs) formation in unsynapsed regions during meiotic recombination (PubMed:20551173, PubMed:27723721). Probably acts by forming a complex with IHO1 and MEI4, which activates DSBs formation in unsynapsed regions, an essential step to ensure completion of synapsis (PubMed:27723721). Interacts with MEI4 (PubMed:20551173). Interacts with IHO1 (PubMed:27723721). Part of the MCD recombinosome complex, at least composed of IHO1, REC114 and MEI4 (PubMed:27723721). Interacts with ANKRD31; the interaction is direct (PubMed:31000436, PubMed:31003867). Expressed in adult testis and embryonic ovary. Also expressed at low levels in liver. In the testis, expression is detected at 4 days postpartum (dpp) with a peak at day 10. Levels decrease between 14-18 dpp with an increase in the adult. Belongs to the REC114 family.