Enzyme involved in the catabolism of H(2)CO(3) but that does not mediates the reversible hydration of carbon dioxide. Mediates complex I assembly in mitochondria and respiration (By similarity). Homotrimer (Probable). Component of the oxidoreductase respiratory chain complex I; element of the extra matrix-exposed domain, which is attached to the membrane arm of this complex. Probably integral to the membrane. Belongs to the gamma-class carbonic anhydrase family.