Actin-depolymerizing protein. Severs actin filaments (F-actin) and binds to actin monomers. In germinating pollen grains and elongating pollen tubes, associates with the subapical actin fringe. Specifically expressed in pollen. During male gametophyte development, expressed at the polarized microspore stage, bicellular stage, mature pollen stage, anthesis stage and open flower stage. Expressed in germinating pollen grains and elongating pollen tubes. Belongs to the actin-binding proteins ADF family.