Transcription factor that targets gene promoters containing 2 conserved CGAA repeat sequences (PubMed:30874562). Positively regulates the expression of ergosterol biosynthesis genes including CYP51A and CYP51B encoding the sterol 14-alpha demethylase, and ERG6A and ERG6B encoding the sterol 24-C-methyltransferase (PubMed:30874562). Interacts with HOG1. Phosphorylation at Thr-82, Ser-92, Ser-102, thr-117 and ser-305 by HOG1 is required for regulating expression of ergosterol biosynthesis genes. Reduces the production of ergosterol (PubMed:30874562). Leads to increased sensitivity to azole compounds, but not to iprodione and fludioxonil that target the high osmolarity glycerol (HOG) pathway (PubMed:30874562).