Catalyzes the key reaction of chlorophyll catabolism, porphyrin macrocycle cleavage of pheophorbide a (pheide a) to a primary fluorescent catabolite (pFCC). Works in a two-step reaction with pheophorbide a oxygenase (PaO) by reducing the C20/C1 double bond of the intermediate, RCC. Belongs to the chlorophyll catabolic enzymes (CCEs) (By similarity). May play a role in senescence and response to wounding (PubMed:21807436). 2 oxidized [2Fe-2S]-[ferredoxin] + primary fluorescent chlorophyll catabolite = 3 H(+) + red chlorophyll catabolite + 2 reduced [2Fe-2S]-[ferredoxin] Porphyrin-containing compound metabolism; chlorophyll degradation. Expressed in leaves (PubMed:21807436). Expressed at low levels in roots, stems, panicles and seeds (PubMed:21807436). Induced by wounding and senescence in leaves. Plants silencing RCCR1 exhibit lesion-mimic spots on leaves. Truncated N-terminus.