Transcriptional regulator of sporocyte development (PubMed:10465788). Acts as an adapter-like transcriptional repressor recruiting TPL/TPR corepressors to inhibit TCP transcription factors (PubMed:25378179). Required for nucellus and embryo sac development (PubMed:10500234). Plays a central role in patterning both the proximal-distal and the adaxial-abaxial axes during ovule development (PubMed:12183381). Involved in establishing the prospective chalaza of the ovule and in controlling the cell number and the length of the funiculus, and is required for the development of the integuments (PubMed:10976054). Required, with BEL1, for cytokinin-induced PIN1 expression in ovules (PubMed:22786869). Involved in controlling stamen identity (PubMed:19726570). May also regulate the morphology of lateral organs by repressing auxin production (PubMed:18557819). Homodimer and heterodimer with SPEARs (PubMed:25527103). Interacts in vitro with YAB1, YAB3 and YAB4 (PubMed:15299139). Interacts (via EAR motif) with TPL, TPR1, TPR2, TPR3 and TPR4 (PubMed:25527103, PubMed:25378179). Interacts with SPEAR1, SPEAR2, SPEAR3, SPEAR4, TCP1, TCP6, TCP8, TCP9, TCP11, TCP15, TCP20, TCP21 and TCP23 (PubMed:25527103). Interacts with TCP2, TCP3, TCP4, TCP5, TCP10, TCP13, TCP17 and TCP24 (PubMed:25527103, PubMed:25378179). Expressed in flower buds (PubMed:10465788, PubMed:10500234). Not found in leaves, siliques and stems (PubMed:10465788). Detected in rosette leaves, stem tissue and seedlings (PubMed:10500234). Expressed during microsporogenesis and megasoprogenesis (PubMed:10465788). Expressed during ovule development (PubMed:25378179). Up-regulated by the transcription factor AGAMOUS (PubMed:15254538). Regulated by ATXR3/SDG2 via chromatin methylation (PubMed:21037105). Triggered by TGA9 AND TGA10 in anthers (PubMed:20805327). Contains 1 EAR motif required for the interaction with TPL and TPRs (PubMed:25378179). Complete male and female sterility. Belongs to the NOZZLE family. Extended C-terminus.