Chromatin-associated protein which contributes to the modulation of chromatin structure (such as super-helical structure of DNA) and function (PubMed:25387881). Binds to chromatin of protein-coding genes throughout the genome to regulate nucleosome occupancy and chromatin accessibility, and to modulate the expression of target genes (PubMed:25387881). Negative regulator of stress tolerance (e.g. high salt) (PubMed:25387881). Found in a mRNA splicing-dependent exon junction complex (EJC) (PubMed:25387881). Binds specifically histones H3 and H4 (PubMed:25387881). Interacts with TOP1A, SCC3, At1g61730, At1g20940, At1g13930, DEK4, HDT1, NIT1, SHL, CYP19-1, GEBPL, HSP70-3, PDP2, PDP3, KIN2, RPL11A and PDS5A (PubMed:25387881). Associates with chromatin (PubMed:25387881). Enriched in regions where chromatin is decondensed or sparse in the interphase nuclei (By similarity). Highly expressed in young seedlings. Strongly and rapidly down-regulated by salt stress in shoots and roots. Increased expression of some target genes (e.g. TOP1A, EFS, MBD9, NUP160, DEK1, BIG, HB-1 and CMT3), but reduced expression of other target genes (e.g. HKL1 and PDS5) (PubMed:25387881). Increased salt stress tolerance leading to a better seed germination in high salt conditions (PubMed:25387881). Truncated N-terminus.