Part of ribonuclease P, a protein complex that generates mature tRNA molecules by cleaving their 5'-ends. Endonucleolytic cleavage of RNA, removing 5'-extranucleotides from tRNA precursor. Binds 1 zinc ion per subunit. Consists of a catalytic RNA component and at least 4-5 protein subunits. Belongs to the eukaryotic/archaeal RNase P protein component 4 family. Extended N-terminus.