Major component of the type IV fimbriae that plays an essential role in twitching motility, natural transformation, and protease secretion. By PilR (PilR/S system) and RNA polymerase sigma-54 factor/RpoN. Deletion mutants do not colonize the ovine hoof. Mutants show also altered secretion of extracellular proteases and twitching motility. Belongs to the N-Me-Phe pilin family.