Probable transcription factor involved in the regulation of time of flowering through the photoperiod flowering pathway. May repress FT. Interacts with FT. Expressed in siliques, ovules and germinating seeds. Detected in siliques from 3 days after pollination until maturity of seeds. Under epigenetic control. In wild-type plant, FWA is silenced in the sporophyte. The epi-allele fwa mutants (which do not have a change in the nucleotide sequence of FWA) cause a late-flowering phenotype due to ectopic FWA expression in sporophytic tissues. Repression of FWA is dependent on histone H3 'Lys-4' methylation and cytosine methylation which are partly controlled by the lysine-specific demthylase 1 (LSD1) homologs, LDL1 and LDL2. Belongs to the HD-ZIP homeobox family. Class IV subfamily.