Component of a chromatin regulatory complex involved in regulating chromatin structure in the nucleus (By similarity). Promotes flowering under long days (LD) via the regulation of bolting (PubMed:27968983). Expressed in the shoot apical meristem (SAM), embryos, seedlings, cotyledons, leaves primordia, young leaves and roots. During seed development, accumulates in the endosperm and in the developing embryo. Strongly expressed following germination throughout seedling growth, especially in aerial part. Present in cotyledons, including the vascular system, hydathodes and trichomes, as well as in young leaves and leaves primordia. In roots, mostly expressed 7 days after sowing. Localized in the central cylinder of roots and the lower hypocotyl area. Observed in the shoot apical meristem (SAM) and in the vascular system, particularly in the xylem parenchyma and phloem. During reproductive development, observed in flowers of stages 13-15, predominantly in the male reproductive tissues, mostly in the developing pollen grains, the anther tissues and the filaments, and, to a lower extent, in the vascular system of sepals and the gynoecium, and in petals. After fertilization, confined to the placenta and the abscission zone of the siliques. Altered shoot apical meristem (SAM) and increased growth rates, as well as prolonged vegetative phase and delayed bolting resulting in an impairment of flowering under long days (LD). Altered expression of FLC and SOC1. Belongs to the WD repeat SWD2 family.