May be involved in gonadal development. Isoform 1 and isoform 3 are expressed in ovary, testis, brain and heart. Isoform 4 and isoform 5 are expressed in brain. Isoform 1, isoform 3, isoform 4 and isoform 5 are expressed in the gonad/mesonephros complex at stage 25. Isoform 1 is expressed in somatic cells of the gonad mesonephros and somites of a tadpole at stage 25.