Required for normal embryogenesis and seed viability. Involved in the first asymmetrical cell division of the zygote. Regulates the number and planes of cell divisions required for generating the normal embryo proper and suspensor, apical-basal axis, cotyledons and meristem. Part of the nuclear pore complex (NPC). The NPC has an eight-fold symmetrical structure comprising a central transport channel and two rings, the cytoplasmic and nuclear rings, to which eight filaments are attached. The cytoplasmic filaments have loose ends, while the nuclear filaments are joined in a distal ring, forming a nuclear basket. NPCs are highly dynamic in configuration and composition, and can be devided in 3 subcomplexes, the NUP62 subcomplex, the NUP107-160 subcomplex and the NUP93 subcomplex, containing approximately 30 different nucleoporin proteins Interacts (via N-terminus) with RH38. Highly expressed in reproductive tissues, such as flowers and early developing seeds. Detected at low levels in vegetative tissues, such as leaves, shoots, stems or roots. Expressed in anthers, stigma papilla, embryo and endosperm. Contains FG repeats mediating the translocation through the NPC by interacting with transport factors. Lethal when homozygous. Knockdown alleles have no obvious defect in growth and development. Named LONO after an Hawaiian god associated with fertility, agriculture, rainfall and music.