High-affinity permease involved in the transport of the cationic amino acids (e.g. arginine, lysine, histidine, citrulline, valine, and glutamate). Transport mostly basic amino acids, and, to a lower extent neutral and acidic amino acids. May function as a proton symporter. Inhibited by the protonophore 2,4-dinitrophenol. Optimal transport of histidine at pH 4.5-5. Expressed in roots, stems, flowers, petioles, seeds, siliques, and leaves. Mostly present in major veins. Belongs to the amino acid-polyamine-organocation (APC) superfamily. Cationic amino acid transporter (CAT) (TC 2.A.3.3) family. Truncated N-terminus. Truncated N-terminus.