Catalytic component of the chromatin structure-remodeling complex (RSC), which is involved in transcription regulation and nucleosome positioning. Controls stem cell fate via the transcription regulation of WUS in the shoot apical meristem, by modulating its promoter. LFY-dependent repressor of the meristem identity switch from vegetative to reproductive development probably by modulating chromatin state. Involved in the regulation of floral homeotic gene expression in response to environmental stimuli. Required for carpel and ovule development, and for cotyledon separation via the regulation of CUC2 transcription. Regulates the promoters of several genes downstream of the jasmonate (JA) and ethylene (ET) signaling pathways. Required for resistance against the necrotrophic pathogen B.cinerea but not the biotrophic pathogen P.syringae. Interacts with LFY (PubMed:22323601). Binds to BARD1/ROW1 (PubMed:18591352). Mostly expressed in rapidly dividing cells in the vegetative, inflorescence, and root meristems, as well as in young leaf and flower primordia. Isoform 1 is predominantly found in seedlings whereas isoform 2 is present in both seedlings and inflorescences (at protein level). By wounding. Phosphorylated. Precocious transition from inflorescence to flower formation and impaired maintenance of the shoot apical meristem (SAM) during the reproductive phase. Abnormal flowers with splayed open first-whorl sepals due to outward bending of the pointy sepal tips. Reduced male fertility and reduced anther dehiscence. Partially unfused at the tip fourth-whorl carpels, with stigmatic tissue missing or placed internal to the carpel tip, leading to funnel shaped carpels. Female sterility ovule growth arrest at megagametogenesis. Fused cotyledons. Impaired expression of PDF1.2a, leading to reduced ethylene (ET) and jasmonic acid (JA) signaling. Reduced resistance toward B. cinerea. Belongs to the SNF2/RAD54 helicase family. Truncated N-terminus.