Expressed in the shoot apex and leaves (PubMed:18315698). Expressed in dry seeds (PubMed:20490919). Expressed in roots and leaves (PubMed:24062085). Induced by chilling and mannitol (PubMed:7948880, PubMed:9517002). Induced by drought stress (PubMed:9517002, PubMed:20490919, PubMed:24062085). Induced by salt stress (PubMed:9517002, PubMed:20490919, PubMed:24062085). Induced by abscisic acid (ABA) (PubMed:20490919, PubMed:24062085). Induced by D-allose and D-glucose (PubMed:23397192). Belongs to the LEA type 4 family.