Component of the replication protein A complex (RPA) required for DNA recombination, repair and replication. The activity of RPA is mediated by single-stranded DNA binding and protein interactions. Plays an essential role in meiotic and somatic DNA repair, but is dispensable for DNA replication and homologous recombination. Is essential for normal progression through meiosis in pollen mother cells. Is involved in repair of double-strand DNA breaks (DSBs) induced by genotoxic stresses. Heterotrimer of RPA1, RPA2 and RPA3 (canonical replication protein A complex) (By similarity). Interacts with RPA2B. Expressed in root tips, roots, shoot apical meristem (SAM), young leaves, flag leaves and ears, and at lower levels in mature leaves. Repressed by sucrose starvation. Normal phenotype during vegetative growth, but sterility at the reproductive stage. No formation of embryo sac in female meiocytes and abnormal chromosomal fragmentation after anaphase I in male meiocytes. Belongs to the replication factor A protein 1 family. Truncated N-terminus.