May play a key role in egg organization. May be a transcriptional regulator having a role in chromatin remodeling in concert with Hira, a histone chaperone. Involved in chromosome segregation by affecting kinetochores function in the first meiotic division. Abundant in the nucleoplasm. Only a fraction of the protein is associated with the chromosomes. Localizes to the chromosome arms and coexpresses with cid at the centromere. Oocyte specific. Expressed both maternally and zygotically. Expressed at all oogenic stages. The N-terminus is blocked. Females exhibit disrupted chromosome segregation in the first meiotic division and produce very low numbers of viable progeny. This female sterility is partially suppressed when some oocytes undergo precocious anaphase. Analysis of the X chromosome of these progeny demonstrates they have inherited the two maternal X-chromosome homologs and have no paternal chromosome markers. This suggests that they developed from diploid gametes that underwent gynogenesis, a form of parthenogenesis that requires fertilization.