Plays a central role as a glucosyl donor in cellular metabolic pathways. alpha-D-glucose 1-phosphate + H(+) + UTP = diphosphate + UDP-alpha-D-glucose Homooctamer. Localizes to yeast but not hyphal cell surface. Expression is up-regulated during oropharyngeal candidiasis (OPC) infection and during stationary phase. Repressed by HOG1, BCR1 and hydrogen peroxide. Present with 17200 molecules/cell in log phase SD medium. Belongs to the UDPGP type 1 family.