Cytoskeletal linker protein, which is essential for attachment of the centrosome to the nucleus (PubMed:14697201). Required for dynein localization to the nuclear envelope (PubMed:14697201). Forms a LINC (LInker of Nucleoskeleton and Cytoskeleton) complex together with unc-84, that may be involved in DNA damage repair (PubMed:27956467). Homodimer (PubMed:14697201). Interacts with the dynein subunit dli-1 via its N-terminus (PubMed:14697201). May interact with microtubules (PubMed:14697201). Interacts with sut-2 (PubMed:19273536). Interacts (via C-terminus) with unc-84 (via C-terminus); the interaction is direct (PubMed:27956467). Localizes to the minus end of microtubules, proximal to the centrosome. Centrosomal localization requires sun-1 and microtubules. Expressed in the syncytial gonad, oocytes, and in all cells during the development of the early embryo. Induced by DNA-damage. The large coiled coil domain is required for homodimerization. RNAi-mediated knockdown results in a reduced number of viable progeny following incubation with the DNA cross-linking agent cisplatin. In contrast to isoform a and isoform d, isoform b and isoform c contain a potential transmembrane domain which may mediate insertion into the nuclear membrane. Does not contain a transmembrane domain. Does not contain a transmembrane domain. Belongs to the hook family.