Plays a role in the coordination of growth and proliferation. Required for entry into G0 phase under conditions of carbon limitation. Involved in the general stress response; acts together with PSR1 to activate stress response element (STRE)-mediated gene expression, possibly through dephosphorylation of MSN2. Interacts with MSN2 and PSR1. Negative mutants fail to enter the G0 phase under conditions of carbon limitation. Present with 1800 molecules/cell in log phase SD medium. Belongs to the WHI2 family.