Component of the LINC (LInker of Nucleoskeleton and Cytoskeleton) complex involved in the connection between the nuclear lamina and the cytoskeleton (By similarity). Is required to nuclear migration in eye and to anchor klar in the nuclear membrane (PubMed:18820457). Core component of the LINC complex which is composed of inner nuclear membrane SUN domain-containing proteins coupled to outer nuclear membrane KASH domain-containing nesprins. Lam is required for perinuclear localization of Koi. Expressed in all cells in the eye disk. Homozygous mutant flies live to adulthood and are fertile and exhibit rough and irregular eyes. Flies have an aberrant R-cell rhabdomeres morphology and R-cell and cone cell nuclei are mispositioned.