Key receptor for sensing species-specific attractants in cooperation with other pollen receptor-like kinases (PubMed:26961657). Essential for pollen tube reorientation toward attractant peptides (PubMed:26961657, PubMed:29109411). Interacts with ROPGEF8, ROPGEF9, ROPGEF12, ROPGEF13, PRK3, LIP1 and LIP2 (PubMed:26961657). Binds to LURE peptides via its LRR repeats; interacts with LURE1.1, LURE1.2, LURE1.3 and LURE1.4 (PubMed:29109411). Re-localizes asymmetrically to direct tip growth direction toward the attractant peptides. Expressed specifically in the pollen tube, predominantly at the tip. The juxtamembrane domain (288-383) is required for interactions with ROPGEFs, while the kinase domain (384-659) is required for pollen tube growth. The protein kinase domain may be catalytically impaired due to the lack of the conserved Asp active site at position 512, which is replaced by a Asn residue. Impaired response to the ovular attractant LURE1.2. Belongs to the protein kinase superfamily. Ser/Thr protein kinase family.