Probable transcription factor (By similarity). Promotes cuticle development probably by modulating the expression of the downstream genes BDG and FDH, possibly repressed in a CFL1-dependent manner (PubMed:21954461). Involved, together with PDF2, in the regulation of flower organs development by promoting the expression of APETALA 3 (AP3) in the epidermis and internal cell layers of developing flowers (PubMed:23590515). In opposition to BBM, seems to promote cell differentiation and giant cell identity via transcriptionnal repression of meristem and cell proliferation genes (PubMed:25564655). Interacts with CFL1 (PubMed:21954461). Binds with BBM (PubMed:25564655). Expressed in trichomes forming at the base of young leaves, in endodermal cell lines around emergent lateral roots and in the epidermal layer of the stamen filament. During embryogenesis, first observed at low levels in the embryo protoderm starting at the late globular stage (PubMed:25564655). In primary and lateral roots, observed in the epidermis, the outer layer of columella cells and lateral root cap (PubMed:25564655). Restricted to the epidermal cell layer during floral organ formation (PubMed:23590515). Also present in flower meristems, shoot apical mersitems (SAM) and leaf primordia (PubMed:25564655). Defective cuticle phenotypes leading to curly flag leaves and organ fusion (PubMed:21954461). The double mutant pdf2-1 hdg1-1 exhibits abnormal flowers with sepaloid petals and carpelloid stamens in association with a reduced expression of APETALA 3 (AP3) in the epidermis and internal cell layers of developing flowers (PubMed:23590515). Increased cell division leading to cell overproliferation (PubMed:25564655). Belongs to the HD-ZIP homeobox family. Class IV subfamily.