Involved in RNA editing. Catalyzes the specific deamination of adenosine-37 in the cytosolic tRNA-Ala. Generates inosine at the position 3'-adjacent to the anticodon tRNA-Ala. adenosine(37) in tRNA(Ala) + H(+) + H2O = inosine(37) in tRNA(Ala) + NH4(+) Binds 1 myo-inositol hexakisphosphate (IP6) per subunit. Homodimer. Highly expressed in siliques. Expressed at low levels in roots, rosette leaves, cauline leaves, stems and flowers. No visible phenotype under normal growth conditions, but mutant plants exhibit reduced growth rate under cold or heat stresses. Belongs to the ADAT1 family.