Component of the MIS complex, a complex that mediates N6-methyladenosine (m6A) methylation on some mRNAs during meiosis and is required for sporulation. In the complex, mediates the entry of the MIS complex in nucleolus, where methylation takes place. Component of the MIS complex, composed of SLZ1, MUM2 and IME4. Mediates localization of the MIS complex to the nucleolus during the period of m6A mRNA accumulation. During sporulation. Expression is activated by IME1 during meiosis. Impairs sporulation.