Transcription factor involved in the expression of a broad class of genes including snRNAs. Required for sporulation and DNA-damage repair. Prevents the spreading of SIR silencing at telomeres and protects histone H4, but not H3, from deacetylation (By similarity). Expression is regulated by FKH2 and repressed upon adherence to polystyrene and interaction with macrophages. Truncated N-terminus.