Transcription factor that mediates regulation of both acid- and alkaline-expressed genes in response to ambient pH. At alkaline ambient pH, activates transcription of alkaline-expressed genes (including pacC itself) and represses transcription of acid-expressed genes (By similarity). Not required for the pH-induced dimorphic transition. Activated by C-terminal proteolytic cleavage by signaling protease at neutral to alkaline ambient pH. Belongs to the pacC/RIM101 family.