Acts as a hyperosmolarity-gated non-selective cation channel that permeates Ca(2+) ions (PubMed:25162526, PubMed:30190597). Shows the following permeability sequence: K(+) > Ba(2+) = Ca(2+) > Na(+) = Mg(2+) = Cs(+) (PubMed:25162526). Mechanosensitive ion channel that converts mechanical stimuli into a flow of ions (PubMed:30190597). Activated by mechanical pressure. Homodimer. Expressed in leaves, flowers, roots and guard cells. Impaired osmotic Ca(2+) signaling in guard cells and root cells, and attenuated water transpiration regulation and root growth in response to osmotic stress. Belongs to the CSC1 (TC 1.A.17) family.