Probable dehydrogenase required for replication of Brome mosaic virus. Involved in vacuolar processing and morphology. N-glycosylated. Exhibits lumenal vesicles within vacuoles suggestive of vacuole fusion/fission defects. Leads to internal accumulation of precursor CPY. Present with 892 molecules/cell in log phase SD medium. Belongs to the short-chain dehydrogenases/reductases (SDR) family.