Component of the anaphase promoting complex/cyclosome (APC/C), a cell cycle-regulated E3 ubiquitin-protein ligase complex that controls progression through mitosis and the G1 phase of the cell cycle. Protein modification; protein ubiquitination. The APC/C is composed of at least 11 subunits that stay tightly associated throughout the cell cycle (By similarity). Interacts with APC10, FZR1, FZR2, FZR3 (PubMed:21687678). Binds to GIG1 and PYM (PubMed:22167058, PubMed:22844260). Part of the mitotic checkpoint complex (MCC); interacts with MAD2, BUB3.1, BUBR1 and BUB1 (PubMed:21687678). Binds to cyclins CYCA1-2, CYCB2-1 and CYCB2-2 (PubMed:21687678). Interacts with PANS1 (PubMed:24206843). Expressed in meristems and organ primordia. Present in flowers, leaves, stems, roots, pollen grains and developing seeds. Expression starts to rise in the S-phase and peaks in the M-phase with a non-negligible basic expression level also in the other cell cycle phases. Delay in plant development due to reduced cell number and male sterility. Belongs to the WD repeat CDC20/Fizzy family.