Component of the large ribosomal subunit. The ribosome is a large ribonucleoprotein complex responsible for the synthesis of proteins in the cell. Component of the large ribosomal subunit. May bind IPO9 with low affinity (By similarity). Interacts with RBM3 (By similarity). Citrullinated by PADI4. Belongs to the universal ribosomal protein uL4 family.