Causes widespread changes in gene expression, and plays a direct role in the regulation of genes encoding ribosomal components. Associates with chromosomal DNA through interaction with RNA polymerase. Contributes to biofilm formation and secondary metabolite production. Important during transitions to and from the survival state. Interacts with RNA polymerase. Up-regulated at post-transcriptional level during survival and slow-growth conditions. Deletion mutant forms smaller biofilms, and produces more pyocyanine and less phenazine-1-carboxylic acid (PCA) than the wild-type. Affects expression of many genes and impacts fitness in fluctuating conditions.