Inactive ADP-ribosyltransferase that functions with SRO1 to regulate oxidative stress, hormonal and developmental responses. Required for embryogenesis, vegetative and reproductive development, and abiotic stress responses. May regulate several stress-responsive genes. Seems to play a larger developmental role than SRO1. Does not bind NAD in vitro. Interacts with the transcription factors NAC013/NTL1 and NAC046 (PubMed:25348421). Interacts with dehydration-responsive DREB2 proteins and a number of transcription factors belonging to several protein families. Interacts with turnip crinkle virus (TCV) movement protein P8. Speckle-like pattern. Expressed in young developing tissues, such as young leaves and flowers and root tips. In mature plants, expressed in vasculature of leaves and roots, and guard cells. Expressed in the embryo proper at the globular stage. Expressed in the embryo until the torpedo stage, after which expression within the procambial strands becomes pronounced. By ozone, salt stress and cadmium. Small plants, altered leaf shape and early flowering. Plants lacking RCD1 show enhanced resistance to methyl viologen, tolerance to freezing and supplementary UV-B irradiation, reduced sensitivity to abscisic acid, ethylene and jasmonate, increased sensitivity to ozone and up-regulation of reactive oxygen species scavenging enzymes. Plants overexpressing RCD1 show a weak rcd1 mutant phenotype. May be due to a competing acceptor splice site. Lacks the conserved catalytic triad His-Tyr-Glu of the active site. Sequencing errors.