Probably catalyzes the NADPH-dependent reduction of acrylyl-CoA to propanoyl-CoA. NADP(+) + propanoyl-CoA = acryloyl-CoA + H(+) + NADPH Homodimer. 100-fold increased sensitivity to acrylate, about 8-fold increased sensitivity to 3-hydroxypropionate. Acrylate is bacteriostatic, not bacteriocidal. Can be complemented by acuI from a number of other bacteria, including Rhodobacter sphaeroides strain 2.4.1 (AC Q3J6K9) and Ruegeria pomeyroi (AC Q5LS56). The zinc-binding residues of the alcohol dehydrogenase family are not conserved. Belongs to the zinc-containing alcohol dehydrogenase family. Acrylyl-CoA reductase subfamily.