Involved in formate-dependent uric acid degradation under microaerobic and anaerobic conditions. May reduce the enzymes necessary for uric acid degradation. Binds 5 [4Fe-4S] clusters. Induced under anaerobic and microaerobic conditions. Expression increases in the presence of hypoxanthine, xanthine and uric acid. Expression is reduced in the presence of nitrite and nitrate under anaerobic conditions. Contains an N-terminal 4Fe-4S dicluster domain and a C-terminal pyridine nucleotide-disulfide oxidoreductase domain. Extended N-terminus.