Probable transcription factor involved in the development of floral organs. Acts as a C-class protein in association with MADS3. Involved in the control of lodicule number (whorl 2), stamen specification (whorl 3), floral meristem determinacy and regulation of the carpel morphogenesis (whorl 4). Plays a more predominant role in floral meristem determinacy than MADS3. Expressed in the lodicule, stamen carpel and ovule primordia. Expressed in lodicules, stamens, carpels and ovules during their development. Abnormal flowers with floral-like organs.