Component of the large ribosomal subunit. The ribosome is a large ribonucleoprotein complex responsible for the synthesis of proteins in the cell. Component of the large ribosomal subunit (PubMed:23636399, PubMed:32669547). May bind IPO9 with low affinity (PubMed:11823430). Interacts with RBM3 (By similarity). Citrullinated by PADI4. Belongs to the universal ribosomal protein uL4 family.