Polycomb group (PcG) protein. PcG proteins act by forming multiprotein complexes, which are required to maintain the transcriptionally repressive state of homeotic genes throughout development. PcG proteins are not required to initiate repression, but to maintain it during later stages of development. They probably act via the methylation of histones, rendering chromatin heritably changed in its expressibility. Required to prevent the proliferation of the central cell by repressing unknown target genes before fertilization. Regulates the anteroposterior organization of the endosperm. Probably indirectly associated with FIE and/or MEA. In plants, PcG complexes are probably composed of a member of the EZ family (CLF or MEA), FIE, and a member of the VEFS family (FIS2, VRN2 or EMF2). Weakly expressed. Expressed in late siliques. Expressed maternally and zygotically. Expressed in the central cell before fertilization, and in the endosperm after fertilization, then decreases before the time of endosperm cellularization but continues in the chalazal cyst. Belongs to the VEFS (VRN2-EMF2-FIS2-SU(Z)12) family. In cv. Columbia (AC P0DKJ8), the sequence differs from that shown due to a deletion in the genomic sequence that remove 60 residues after the Asn-562. Truncated N-terminus.