Acts as a transcriptional activator. Binds to the DNA consensus sequence 5'-GACCACCCA-3' (By similarity). May regulate the transcription of specific genes during normal development (PubMed:8948590). May play a role in craniofacial development and digital development, as well as development of the central nervous system and gastrointestinal tract (PubMed:8948590). Mediates SHH signaling (PubMed:8948590). Plays a role in cell proliferation and differentiation via its role in SHH signaling. Belongs to the GLI C2H2-type zinc-finger protein family.