Tubulin is the major constituent of microtubules. The gamma chain is found at microtubule organizing centers (MTOC) such as the spindle poles or the centrosome, suggesting that it is involved in the minus-end nucleation of microtubule assembly. Required for oocyte activation and consequently for organization of the female meiotic spindle (PubMed:9155007, PubMed:9698447). Essential for centrosome organization and assembly of biastral mitotic spindles in embryos. Plays a role in stabilizing the augmin complex on the meiotic spindle (PubMed:23785300). Interacts with Ote. Embryonic mitotic spindle, female meiotic spindle. Expressed in nurse cells and oocytes of developing egg chambers. Expressed both maternally and zygotically. Zygotic expression is restricted to adult females. RNAi-mediated knockdown in oocytes results in spindle defects. Belongs to the tubulin family.