Has double-stranded DNA-stimulated ATPase and ATP-dependent DNA helicase (5' to 3') activity suggesting a role in nuclear processes such as recombination and transcription. Proposed core component of the chromatin remodeling Ino80 complex which is involved in transcriptional regulation, DNA replication and probably DNA repair. ATP + H2O = ADP + H(+) + phosphate Forms homohexameric rings. May form a dodecamer with rvb1 made of two stacked hexameric rings. Component of the chromatin remodeling Ino80 complex. Component of the RNA polymerase II holoenzyme complex (By similarity). Belongs to the RuvB family.