Catalyzes the transfer of short (four to five carbons) branched acyl chains to the furanose ring of di-acylsucrose acceptors to produce tri-acylsucroses such as S3:15 (5,5,5), S4:17 (2,5,5,5) and S4:24 (2,5,5,12) acylsucroses. Monomer. Expressed in tip cells of type I trichomes of stems and petioles, sites of acylsugars production. Reduced total acylsugar levels, with abnormal accumulation of di-acylsucrose, S2:17 (5,12), and two triacylsucroses, S3:19 (2,5,12) and S3:22 (5,5,12) lacking furanose ring acylation, but impaired accumulation of S3:15 (5,5,5), S4:17 (2,5,5,5) and S4:24 (2,5,5,12) acylsucroses. Belongs to the plant acyltransferase family.