Probably catalyzes the NADPH-dependent reduction of acrylyl-CoA to propanoyl-CoA. Restores acrylate resistance when expressed in an E.coli strain K12 acuI deletion. NADP(+) + propanoyl-CoA = acryloyl-CoA + H(+) + NADPH Homodimer. By dimethylsulfonioproprionate (DMSP) and acrylate. 25-fold increased sensitivity to acrylate, growth is strongly inhibited by 20 mM dimethylsulfonioproprionate (DMSP). The zinc-binding residues of the alcohol dehydrogenase family are not conserved. Belongs to the zinc-containing alcohol dehydrogenase family. Acrylyl-CoA reductase subfamily.