Transcriptional corepressor. Negative regulator of jasmonate responses (By similarity). Tetramer (PubMed:26601214). Interacts with NINJA/AFPH2 (PubMed:20360743). Interacts with SMXL6 (PubMed:22065421). Interacts with SPL (via EAR motif) (PubMed:25527103, PubMed:25378179). Interacts with SPEAR3/TIE1 (PubMed:23444332). A number of isoforms are produced. According to EST sequences. The N-terminal TOPLESS domain (TPD) (1-209) binds directly to a 12-amino acid LxLxL EAR motif peptide.