Required for meiotic recombination, synaptonemal complex formation and cell cycle progression. Recruited to meiosis recombination hotspots by MEI5 and SAE3. Double stacked ring-shaped homooctamer (By similarity). Forms a ternary complex with MEI5 and SAE3. Interacts with RDH54. Expressed 2.5 hours after induction of meiosis. Belongs to the RecA family. DMC1 subfamily.