Binds actin monomers (G actin) and plays a role in the reorganization of the actin cytoskeleton and in formation of actin stress fibers. Interacts (via RPEL repeats) with ACTA1. Enriched at synapses (By similarity). Cytoplasmic in resting cells, and is imported into the nucleus upon serum stimulation. Interaction with actin prevents nuclear import (By similarity). Expressed in the gizzard, and in neurons from central and peripheral nervous systems. Binds three actin monomers via the three C-terminal RPEL repeats. Belongs to the phosphatase and actin regulator family.