Adapter protein, which is found in various multiprotein chromatin complexes with histone acetyltransferase activity (HAT), which gives a specific tag for epigenetic transcription activation. May be required for the mitotic checkpoint and normal cell cycle progression (By similarity). May play a role in the formation and maintenance of the auditory system (PubMed:31231791). Morpholino TRRAP knockdown results in defects in the inner ear. Morphant larvae show reduced lateral line neuromasts, decreased number of hair cells per neuromast, and fewer and thinner stereocilia on the hair cells. In addition, the acoustic startle response is decreased and sound-induced fast escape reflex is impaired. Belongs to the PI3/PI4-kinase family. TRA1 subfamily.