The player plays as Dante through seven missions, taking over halfway through the game. His gameplay is like that in "Devil May Cry 3". He has access to multiple melee and ranged weapons which he gains after boss battles. He is able to cycle through them freely in combat and is no longer limited to equipping two weapons of each type as he was in the previous game. Dante also begins with his four styles, each of which grants him different abilities. But, unlike in "Devil May Cry 3", he may now switch them at will with buttons or pads on the controller. He also gains the Dark Slayer style near the end of his appearance, which only has one style level. Styles do not level up through experience as in the previous game, but instead must be upgraded like other skills in the shop screen in between missions or at statues. Dante can also enter Devil Trigger where he gains most of the benefits that Nero's Devil Trigger has. Since he does not have the Devil Bringer, he gets animation and property changes on some of his normal attacks instead.