Constantinople was founded by the Roman Emperor Constantine I (272–337) in 324 on the site of an already-existing city, Byzantium, which was settled in the early days of Greek colonial expansion, in around 657 BC, by colonists of the city-state of Megara. This is the first major settlement that would develop on the site of later Constantinople, but the first known settlements was that of "Lygos", referred to in Pliny's Natural Histories. Apart from this, little is known about this initial settlement. The site, according to the founding myth of the city, was abandoned by the time Greek settlers from the city-state of Megara founded Byzantium () in around 657 BC, across from the town of Chalcedon on the Asiatic side of the Bosphorus.