The predicted sphingid was discovered 21 years later and described as a subspecies of the one African species studied by Wallace: "Xanthopan morganii praedicta", for which, the subspecific name "praedicta" ("the predicted one") was given. The Madagascan individuals had a pink, rather than white, breast and abdomen and a black apical line on the forewing, broader than in mainland specimens. Molecular clock models using either rate- or fossil-based calibrations imply that the Madagascan subspecies "X. morgani praedicta" and the African subspecies "morgani" diverged 7.4 ± 2.8 Mya (million years ago), which overlaps the divergence of "A. sesquipedale" from its sister, "A. sororium", namely 7.5 ± 5.2 Mya. Since both these orchids have extremely long spurs, long spurs likely existed before that and were exploited by long-tongued moths similar to "Xanthopan morganii praedicta". The long geological separation of subspecies "morgani" and "praedicta" matches their morphological differences in the color of breast and abdomen.