Evidence for an erect stance can be found in trackways attributed to diadectids. The most well-preserved of these trackways are present in the Tambach Formation in central Germany. A 2007 study identified two different ichnospecies, "Ichniotherium cottae" and "I. sphaerodactylum", as footprints of the diadectids "Silvadectes absitus" and "Orobates pabsti", respectively. This was the first species-level identification of trackmakers of Paleozoic-era trackways, making the footprints the oldest yet associated with specific animal species. The close positioning of the footprints attributed to the more advanced diadectides suggests that the animals held their feet almost underneath their bodies, giving them a more efficient gait and to some degree paralleling the stance of mammals more than that of the sprawling amphibians and reptiles.