The subfamily Homininae can be further subdivided into three branches: the tribe Gorillini (gorillas), the tribe Hominini with subtribes Panina (chimpanzees) and Hominina (humans and their extinct relatives), and the extinct tribe Dryopithecini. The Late Miocene fossil "Nakalipithecus nakayamai", described in 2007, is a basal member of this clade, as is, perhaps, its contemporary "Ouranopithecus"; that is, they are not assignable to any of the three extant branches. Their existence suggests that the Homininae tribes diverged not earlier than about 8 million years ago (see Human evolutionary genetics).