Another species analysed in Gulden's 2001 study, "Galerina pseudomycenopsis", also could not be distinguished from "G. marginata" based on ribosomal DNA sequences and restriction fragment length polymorphism analyses. Because of differences in ecology, fruit body color and spore size combined with inadequate sampling, the authors preferred to maintain "G. pseudomycenopsis" as a distinct species. A 2005 study again failed to separate the two species using molecular methods, but reported that the incompatibility demonstrated in mating experiments suggests that the species are distinct.