Environment is an important factor in how innate behavior has evolved. A hypothesis of Michael McCollough, a positive psychologist, explains that environment plays a key role in human behaviors such as forgiveness and revenge. This hypothesis theorizes that various social environments cause either forgiveness or revenge to prevail. McCollough relates his theory to game theory. In a tit-for-tat strategy, cooperation and retaliation are comparable to forgiveness and revenge. The choice between the two can be beneficial or detrimental, depending on what the partner-organism chooses. Though this psychological example of game theory does not have such directly measurable results, it provides an interesting theory of unique thought. From a more biological standpoint, the brain's limbic system operates as the main control-area for response to certain stimuli, including a variety of instinctual behavior. The limbic system processes external stimuli related to emotions, social activity, and motivation, which propagates a behavioral response. Some behaviors include maternal care, aggression, defense, and social hierarchy. These behaviors are influenced by sensory input — sight, sound, touch, and smell.