The number of available budding locations has a large effect on colony fragmentation. A large number of bud nests results in small colony fragments, indicating that the colony has the ability to control size and caste ratios. However, a minimum group size of 469 individuals appears preferred by the species. Amount of fragmentation does not have an effect on food distribution. After budding, nest units do not compete for resources, but rather act cooperatively. This is evolutionarily explained by the high amount of genetic relatedness among these nest units. In addition, major disturbances to the central nest cause the colony to abandon it and flee to a bud nest. Thus, nest units may exchange individuals after budding occurs, further explaining their cooperative behavior.