Most of Caria surrendered to the Persians in aftermath of Lade, although some strongholds had to be captured through force. The Persian fleet and army wintered at Miletus, before setting out in 493 BC to finally stamp out the last embers of the revolt. They attacked and captured the islands of Chios, Lesbos and Tenedos and then moved over to the mainland, and captured each of the remaining cities of Ionia. Although the cities of Ionia were undoubtedly harrowed in the aftermath, none seems to have suffered quite the fate of Miletus. The Persian army then re-conquered the settlements on the Asian side of the Propontis, while the fleet sailed up the European coast of the Hellespont, taking each settlement in turn. With all of Asia Minor now firmly returned to Persian rule, the revolt was finally over.