In the early 19th century, following independence, Peru, Ecuador, Colombia, and Brazil had overlapping claims to the Northwestern Amazon Basin, based on each country's interpretation of their colonial "de jure" titles. The disputed area was populated mostly by groups of nomadic Amerindian natives living in the Amazon jungle. In addition there were semi-assimilated sedentary Amerindians living with a handful of whites and mestizos, dedicated to trading in sparsely populated trading port villages that were found scattered along the river banks of the Amazon Basin. During the colonial era the disputed area known as Maynas had numerous missions administered by the Jesuits of Quito. After the Jesuits were expelled from South America, only a handful of missions survived in the 19th century as isolated trading villages. The Brazilians, by contrast, had a chain of villages along the Amazon River that stretched to its ports along the Atlantic Ocean.