The first known organized states to have ruled over the region of present-day Uşak were the Phrygians in the eastern portion and the Lydians in the west during the seventh century BC. The Karun Treasure, discovered by clandestine treasure hunters in Uşak in 1965, and whose smuggling outside Turkey and subsequent retrieval decades later from New York City's Metropolitan Museum of Art made international headlines, gives an indication of the high degree of civilization attained by these Anatolian states. The region of Lydia was later taken over by the Persian Empire in the 6th century BC and by Alexander the Great and his successors as of the 4th century. Thereafter, Uşak was ruled successively by the Roman Empire, the Byzantine Empire, the Germiyanids and finally the Ottoman Empire (as of 1429).