R.A. Fisher developed an explanation, known as Fisher's principle, of why sex ratios in many animals are 1:1. If there were 10 times more females in a population than males, a male would on average be able to mate with more partners than a female would. Parents who preferentially invested in producing male offspring would have a fitness advantage over those who preferentially produced females. This strategy would result in increasing numbers of males in the population, thus eliminating the original advantage of males. The same would occur if there were originally more females than males in a population. The evolutionarily stable strategy (ESS) in this case would be for parents to produce a 1:1 ratio of males and females.