The subordinate theme has a complex structure, a three-part form of its own (a-b-a'), which leads into the closing theme in measure 86 (Animato), which concludes the exposition in measure 107. The development (beginning in measure 108) begins with the same material as the exposition, except the cello is in the low register, and the key is changed from D major to F major. After some contrapuntal work, the development reaches a dominant pedal point (measure 167), which resolves in the main key of the piece, D major, in the recapitulation in measure 180. The recapitulation follows the broad outlines of the exposition, except the subordinate theme (measure 224) begins in E-flat major instead of the customary D major. The three-part structure of the subordinate theme, though, allows Borodin to reach the expected D major in the a' part of the subordinate theme (measure 257), and the closing theme (measure 266) concludes the movement.