During the Pleistocene interglacials, arctic animals that are no longer extant occupied Scotland, including the woolly rhinoceros, mammoth, polar bear, lemming, Arctic fox and the giant deer "Megaloceros giganteus". Other mammals that used to inhabit Scotland but became extinct in the wild during historic times include the Eurasian lynx, which lived in Britain until 1,500 years ago, the European brown bear, subspecies "Ursus arctos caledoniensis", which was taken to entertain the Roman circuses but died out in the 9th or 10th century, and the elk, which lasted until about 1300. The wild boar and wild ox or urus died out in the subsequent two centuries, although the former's domesticated cousin, the grice, lasted until 1930 in Shetland. The last known wolf was shot on Mackintosh land in Inverness-shire in 1743, and the walrus is now only an occasional vagrant. St Kilda also possessed an endemic subspecies of the house mouse, "Mus musculus muralis", which was longer, hairier, coloured differently and had a skull shape at variance to the norm. It became extinct in 1938, just eight years after the evacuation of the native St Kildans.