Local resource competition (LRC) was first hypothesized by Anne Clark. She argued that the African bushbaby ("Otolemur crassicaudatus") demonstrated a male-biased sex ratio because daughters associated with mothers for longer periods of time than did sons. Since sons disperse further from the maternal territory than do daughters, they do not remain on the territories and do not act as competitors with mothers for resources. Clark predicted that the effect of the LRC on sex allocation resulted in a mother investing preferentially in male offspring to reduce competition between daughters and herself. By producing more male offspring that disperse and do not compete with her, the mother will have a greater fitness than she would if she had produced the ratio predicted by the equal investment theory.