According to another study in 2008, the Native American contribution to present-day Cubans accounted for 33% of the maternal lineages, whereas Africa and Eurasia contributed 45% and 22% of the lineages, respectively. Haplogroup A2 is the main Native American haplogroup in Cuba (21.9% of the total sample), accounting for 67% of the Native American mtDNA gene pool. Regarding Y-chromosome haplogroups (male lineages), 78.8% of the sequences found in Cubans are of West Eurasian origin, 19.7% of African origin and 1.5% of East Asian origin. Among the West Eurasian fraction, the vast majority of individuals belong to West European haplogroup R1b. The African lineages found in Cubans have a Western (haplogroups E1, E2, E1b1a ) and Northern (E1b1b-M81 ) African origin. The "Berber" haplogroup E1b1b1b (E-M81), is found at a frequency of 6.1%.