Colonies comprise socially monogamous, extended family groups with overlapping generations, known as "clans", which exhibit cooperative breeding. Non-breeding individuals become helpers to relatives and assist to raise their brood. In white-fronted bee-eaters, this helping behavior is particularly well developed with helpers assisting in half of all nesting attempts. These helpers may contribute to all aspects of the reproductive attempt, from digging the roosting or nesting chamber, to allofeeding the female, incubating and feeding the young; and have a large effect on increasing the number of young produced.