Traditionally, the "crocodile-normal" and "advanced mesotarsal" ankle arrangements have been considered as a dichotomy among archosaurs: early archosaurs and pseudosuchians possess the more mobile "crocodile-normal" configuration, while pterosaurs and dinosauromorphs (including birds) possess the stiffer "advanced mesotarsal" configuration. The presence of the "crocodile-normal" ankle in "Teleocrater" (convex joint with the astragalus, presence of a tuber, and the convexity of the fibular facet on the calcaneum) indicates that this configuration was probably plesiomorphic for archosaurs, including avemetatarsalians, supported by reconstructions of character state evolution using the two datasets. At the same time, features associated with the "advanced mesotarsal" ankle (lack of a tuber and the concavity of the fibular facet on the calcaneum) were reconstructed as having appeared at least two different times among ornithodirans, with basal dinosaurs also possessing a mixture of "crocodile-normal" and "advanced mesotarsal" characteristics. This demonstrates that the evolution of ankle morphology in avemetatarsalians is more complex than previously thought, and led Nesbitt "et al." to conclude that the strict "crocodile-normal"/"advanced mesotarsal" dichotomy is reductionist.