In 2021, a study of another 4 Late Pleistocene northeast Siberian wolf sequences showed that they are genetically similar to the Taimyr and Yana wolves. These 6 extinct wolves sequentially branched off from the lineage that leads to the modern wolf and dog. The 50,000 YBP Tirekhtyakh River, 48,000 YBP Bunge-Toll site, and 32,000 YBP Yana RHS specimens were separate lineages not related to each other. The 16,800 YBP Ulakhan Sular and the 14,100 YBP Tumat specimens both cluster with a modern wolf from Ellesmere Island, indicating that these 2 specimens derive from the same lineage as the North American wolves. All 6 Late Pleistocene wolves share alleles with Arctic dogs: Greenland dogs, Siberian and Alaskan huskies, Alaskan malamutes, the extinct Zhokhov dog and the extinct pre-European contact dogs of North America. It is possible that another population of extinct wolves, that were related to all six specimens, may have contributed to the ancestry of the Arctic dogs. There was evidence that 4 of the extinct Siberian wolves had contributed to the ancestry of modern wolf populations in Shanxi, western China, and possibly Chukotka and Inner Mongolia.