The various fossil beds of the Green River Formation span a 5 million year period, dating to between 53.5 and 48.5 million years old. This span of time includes the transition between the moist early Eocene climate and the slightly drier mid-Eocene. The climate was moist and mild enough to support crocodiles, which do not tolerate frost, and the lakes were surrounded by sycamore ( e.g. "Platanus wyomingensis" ) forests. As the lake configurations shifted, each Green River location is distinct in character and time. The lake system formed over underlying river deltas and shifted in the flat landscape with slight tectonic movements, receiving sediments from the Uinta highland and the Rocky Mountains to the east and north. The lagerstätten formed in anoxic conditions in the fine carbonate muds that formed in the lakebeds. Lack of oxygen slowed bacterial decomposition and kept scavengers away, so leaves of palms, ferns and sycamores, some showing the insect damage they had sustained during their growth, were covered with fine-grained sediment and preserved. Insects were preserved whole, even delicate wing membranes and spider spinnerets.