During the Scottish Reformation, the abbey church had undergone a first Protestant ‘cleansing’ by September 1559, and was sacked in March 1560. By September 1563 the choir and feretory chapel were roofless, and it was said that the nave was also in a sorry state, with the walls so extensively damaged that it was a danger to enter. In 1672 parts of the east end collapsed, while in 1716 part of the central tower is said to have fallen, presumably destabilising much that still stood around its base, and the east gable tumbled in 1726. The final collapse of the central tower took place in 1753.