Shortly after gaining the power of Geass, Lelouch uses it to make Villetta relinquish her Knightmare to him, leaving her confused about what happened. Though she dismisses it at first, she begins to suspect Lelouch after Jeremiah experiences similar memory loss after odd behavior. When Jeremiah is supposedly killed during the Battle of Narita, Villetta steps up her attempts to find the connection between Lelouch and Zero with the help of Shirley Fenette. After discovering that Lelouch is Zero, Shirley shoots Villetta to protect Lelouch, giving Villetta amnesia. She is found by Kaname Ohgi, with whom she lives for a time. He names her since she cannot remember her real name. The two fall in love, but after unexpectedly regaining her memory at the end of the season, Villetta is disgusted at the thought that she ever loved an Eleven. She confronts him during the Black Rebellion and reveals her true identity, then shoots him in the stomach. This, however, does not kill him.