Respiration occurs by both plants and animals throughout the water column, resulting in the destruction, or usage, of organic matter, but photosynthesis can only take place via photosynthetic algae in the presence of light, nutrients and CO2. In well-mixed water columns plankton are evenly distributed, but a net production only occurs above the compensation depth. Below the compensation depth there is a net loss of organic matter. The total population of photosynthetic organisms cannot increase if the loss exceeds the net production.