Young dunes are called yellow dunes and dunes which have high humus content are called grey dunes. Leaching occurs on the dunes, washing humus into the slacks, and the slacks may be much more developed than the exposed tops of the dunes. It is usually in the slacks that more rare species are developed and there is a tendency for the dune slacks' soil to be waterlogged where only marsh plants can survive. In Europe these plants include: creeping willow, cotton grass, yellow iris, reeds, and rushes. As for vertebrates in European dunes, natterjack toads sometimes breed here.