A typical forest tree breeding program starts with selection of superior phenotypes (plus trees) in a natural or planted forest, often based on growth rate, tree form and site adaptation traits. This application of mass selection improves the mean performance of the forest. Offspring is obtained from selected trees and grown in test plantations that act as genetic trials. Based on such tests the best genotypes among the parents can be selected. Selected trees are typically multiplied by either seeds or grafting and seed orchards are established when the preferred output is improved seed. Alternatively, the best genotypes can be directly propagated by cuttings or in-vitro methods and used directly in clonal plantations. The first system is frequently used in pines and other conifers, while the second is typical in some broadleaves (poplars, eucalypts and others). The objectives of a tree breeding program range from yield improvement and adaptation to particular conditions, to pest- and disease-resistance, wood properties, etc. Currently, tree breeding is starting to take advantage of the fast development in plant genetics and genomics.