The biomedical and biotechnology sectors, including start-up biomedical companies arising from university research, employ just under 1,000 people directly and nearly 2,000 indirectly. Information technology and video game development have been important industries in the city for more than 20 years. Rockstar North, developer of "Lemmings" and the "Grand Theft Auto" series was founded in Dundee as DMA Design by David Jones; an undergraduate of the Abertay University. Other game development studios in Dundee include Denki, Ruffian Games, Dynamo Games, 4J Studios and Outplay Entertainment, among others.