The ultimate goal of karyogamy is fusion of the two haploid nuclei. The first step in this process is the movement of the two pronuclei toward each other, which occurs directly after plasmogamy. Each pronucleus has a spindle pole body that is embedded in the nuclear envelope and serves as an attachment point for microtubules. Microtubules, an important fiber-like component of the cytoskeleton, emerge at the spindle pole body. The attachment point to the spindle pole body marks the minus end, and the plus end extends into the cytoplasm. The plus end has normal roles in mitotic division, but during nuclear congression, the plus ends are redirected. The microtubule plus ends attach to the opposite pronucleus, resulting in the pulling of the two pronuclei toward each other.