Another important brackish water habitat is the mangrove swamp or mangal. Many, though not all, mangrove swamps fringe estuaries and lagoons where the salinity changes with each tide. Among the most specialised residents of mangrove forests are mudskippers, fish that forage for food on land, and archer fish, perch-like fish that "spit" at insects and other small animals living in the trees, knocking them into the water where they can be eaten. Like estuaries, mangrove swamps are extremely important breeding grounds for many fish, with species such as snappers, halfbeaks, and tarpon spawning or maturing among them. Besides fish, numerous other animals use mangroves, including such species as the saltwater crocodile, American crocodile, proboscis monkey, diamondback terrapin, and the crab-eating frog, "Fejervarya cancrivora" (formerly "Rana cancrivora"). Mangroves represent important nesting site for numerous birds groups such as herons, storks, spoonbills, ibises, kingfishers, shorebirds and seabirds.