After Petar Kružić's death, and the lack of water supply, the defenders of Klis finally surrendered to the Ottomans in exchange for their freedom. On March 12, 1537, the town and fortress was given up to the Ottomans, many of the citizens left the town while the Uskoks went to the city of Senj on the Adriatic coast, where they continued fighting the Ottomans. They may have started to gather around Senj as early as 1520 (Bracewell 1992). The Ottoman raids and destruction brought Senj natives together with those from the Habsburg lands, Dalmatians, Ragusans and Italians. At Senj, the Klis Uskoks were soon joined by other refugees from Novi Vinodolski in northwestern Croatia, from Otočac on the Gacka River, and from other Croatian towns and villages.