In 2002, phytochemical screening and analytical chromatography were used to study the presence of phenolic compounds and leucoanthocyanins in several naturally occurring hybrids and their putative parental species (including "N. rajah") from Sabah and Sarawak. The research was based on leaf material from nine dry herbarium specimens. Eight spots containing phenolic acids, flavonols, flavones, leucoanthocyanins and 'unknown flavonoid' 1 and 3 were identified from chromatographic profiles. The distributions of these in the hybrid "N. × alisaputrana" and its putative parental species "N. rajah" and "N. burbidgeae" are shown in the adjacent table. A specimen of "N. × alisaputrana" grown from tissue culture ("in vitro") was also tested.