Anthropologist C. Loring Brace (1993) cross-analysed the craniometric traits of Natufian specimens with those of various ancient and modern groups from the Near East, Africa and Europe. The Late Pleistocene Epipalaeolithic Natufian sample was described as problematic due to its small size (consisting of only three males and one female), as well as the lack of a comparative sample from the Natufians' putative descendants in the Neolithic Near East. Brace observed that the Natufian fossils lay between those of the Niger-Congo-speaking populations and the other samples, which he suggested may point to a Sub-Saharan influence in their constitution. Subsequent ancient DNA analysis of Natufian skeletal remains by Lazaridis et al. (2016) found that the specimens instead were a mix of 50% Basal Eurasian ancestral component (see genetics) and 50% Western Eurasian Unknown Hunter Gatherer (UHG) population related to European Western Hunter-Gatherers.