Early telephone exchanges signaled the switchboard operator when a subscriber picked up the telephone handset to make a call. The operator answered requesting the destination of the call. When manual exchanges were replaced with automated switching systems, the exchange generated a tone to the caller when the telephone set was picked up, indicating that the system was ready to accept dialed digits. Each digit was transmitted as it was dialed which caused the switching system to select the desired destination circuit. Modern electronic telephones may store the digits as they are entered, and only switch off-hook to complete the dialing when the subscriber presses a button.