The Greenland shark has the longest known lifespan of all vertebrate species (estimated to be between 300 and 500 years), and is among the largest extant species of shark. It is a generalist feeder, consuming a variety of available foods. As an adaptation to living at depth, it has a high concentration of trimethylamine N-oxide in its tissues, which causes the meat to be toxic. Greenland shark flesh, treated to reduce toxin levels, is eaten in Iceland as a delicacy known as "kæstur hákarl".