In insects the central complex plays an important role in higher-order brain function. The neuropil in "Drosophila" Ellipsoid is composed of four substructures. Each section has been observed in several insects as well as the influence it has on behavior, however the exact function of this neuropil has proven elusive. Abnormal walking behavior and flight behavior are controlled primarily by the central complex and genetic mutations that disrupt the structure support the hypothesis that the central complex neuropil is a site of behavioral control. However, only specific components of the behavior were affected with the genetic mutations. For example, basic leg coordination of walking was normal, whereas speed, activity, and turning were affected. These observations suggest that the central complex not only plays a role in locomotor behavior, but fine tuning as well. There is also additional evidence that the neuropil may function in olfactory associative learning and memory.