In 1939, Danny, an optimistic cat from in Kokomo, Indiana, travels to Hollywood in hopes of starting an acting career there. After meeting a new friend Pudge, Danny is selected by agent Farley Wink to feature in a film that is in production at Mammoth Pictures called "Li'l Ark Angel", alongside Wink's secretary: a white cat named Sawyer. Upon joining fellow animals; Tilly, Cranston, Frances, and T.W., Danny is dismayed on learning how minor his role is and tries to weasel his way into more time in the spotlight. Danny angers Darla Dimple, a popular yet spoiled child actress and star of the film, so she assigns her 36-foot tall butler Max to intimidate Danny into no longer trying to enlarge his part.