Yellowstone cutthroat trout generally declined in the second half of the 20th century due to angler overharvest, competition with exotic fishes, and overzealous egg collection. Populations rebounded in the park after the advent of catch-and-release-only fishing rules in the 1970s, but new and aggressive invaders are causing an increasing threat to these native fish and alarming park fisheries biologists. Nonnative lake trout, an effective fish predator, were discovered in Yellowstone Lake in 1994. Throughout the west cutthroat trout populations preyed upon by introduced lake trout have typically declined, exhibited lower growth, or have disappeared. Aggressive lake trout control efforts by the National Park Service and no harvest limits have resulted in removing thousands of lake trout from Yellowstone Lake since 1994, including more than 12,000 in 2000. Still, the number of Yellowstone cutthroat trout monitored during the annual fall count in Yellowstone Lake was lower in recent years than at any other time in the 25-year history of the monitoring effort. Whirling disease, which has been implicated in recent years in the decline of trout populations in many western states, was discovered in Yellowstone Lake in 1998. So far, it is unclear which of these two nonnative invaders has been the greater factor in the decline of Yellowstone cutthroat trout, but there is no question they are causing it.