The most widely accepted theory of the population of the islands is the "Out-of-Taiwan" model that follows the Austronesian expansion during the Neolithic in a series of maritime migrations originating from Taiwan that spread to the islands of the Indo-Pacific; ultimately reaching as far as New Zealand, Easter Island, and Madagascar. Austronesians themselves originated from the Neolithic rice-cultivating pre-Austronesian civilizations of the Yangtze River delta in coastal southeastern China pre-dating the conquest of those regions by the Han Chinese. This includes civilizations like the Liangzhu culture, Hemudu culture, and the Majiabang culture. It connects speakers of the Austronesian languages in a common linguistic and genetic lineage, including the Taiwanese indigenous peoples, Islander Southeast Asians, Chams, Islander Melanesians, Micronesians, Polynesians, and the Malagasy people. Aside from language and genetics, they also share common cultural markers like multihull and outrigger boats, tattooing, rice cultivation, wetland agriculture, teeth blackening, jade carving, betel nut chewing, ancestor worship, and the same domesticated plants and animals (including dogs, pigs, chickens, yams, bananas, sugarcane, and coconuts).