In the game's present, Nagi's descendant and self-proclaimed greatest warrior, Susano, refuses to believe in Nagi's legend and frees Orochi, who escapes and curses the lands, sapping the life from Nippon. Sakuya, the wood sprite and guardian of Kamiki Village, calls forth Amaterasu, the sun goddess, known to the villagers as the reincarnation of the white wolf Shiranui, and asks her to remove the curse that covers the land. Accompanied by the artist Issun (an inch-high creature known as a Poncle), Amaterasu begins to restore the land to its normal state.