In the mid-16th century the Tsardom of Russia conquered the Tatar khanates of Kazan and Astrakhan, thus annexing the entire Volga Region and making the way to the Ural Mountains open. The colonisation of the new easternmost lands of Russia and further onslaught eastward was led by the rich merchants Stroganovs. Tsar Ivan IV granted large estates near the Urals as well as tax privileges to Anikey Stroganov, who organized large scale migration to these lands. Stroganovs developed farming, hunting, saltworks, fishing, and ore mining on the Urals and established trade with Siberian tribes.