These localities are the coast of Burnmouth, the banks of the Whiteadder Water near Chirnside, the River Tweed near Coldstream, and the rocks near Tantallon Castle alongside the Firth of Forth. Fossils of both aquatic and terrestrial tetrapods are known from these localities, providing an important record of the transition between life in water and life on land and filling some of the lacunae in Romer's gap. These new localities may represent a larger fauna, as all lie within a short distance of each other and share many fishes with the nearby and contemporary Foulden fish bed locality (which has not produced tetrapods thus far). As with East Kirkton Quarry, tetrapods at these sites were discovered through the long-term efforts of Stan Wood and colleagues.