On Henry's death in 1252 his three sons partitioned the principality, founding the lines of Aschersleben, Bernburg and Zerbst. The family ruling in Aschersleben became extinct in 1315, and this district was subsequently incorporated into the neighbouring Bishopric of Halberstadt, thus dividing the territory of Anhalt-Bernburg into two separate pieces. The last prince of the original line of Anhalt-Bernburg died in 1468 and his lands were inherited by the princes of the sole remaining line, that of Anhalt-Zerbst. The territory belonging to this branch of the family had been divided in 1396, and after the acquisition of Bernburg, Prince George I made a further partition of Zerbst (Zerbst and Dessau). Early in the 16th century, owing to the death or abdication of several princes, the family had narrowed down to the two branches of Anhalt-Köthen and Anhalt-Dessau (both issued from Anhalt-Dessau in 1471).