The Hariyanka dynasty king Bimbisara was responsible for expanding the boundaries of his kingdom through matrimonial alliances and conquest. The land of Kosala fell to Magadha in this way. Estimates place the territory ruled by this early dynasty at 300 leagues in diameter, and encompassing 80,000 small settlements. Bimbisara is contemporary with the Buddha, and is recorded as a lay disciple. Bimbisara (543–493 BCE) was imprisoned and killed by his own son who became his successor, Ajatashatru (491–461 BCE), under whose rule, the dynasty reached its largest extent.