The Christian population of Muslim Spain (the Mozarabs) developed a style of Mozarabic art whose best known survivals are a series of illuminated manuscripts, several of the commentaries on the Book of Revelation by the Asturian Saint Beatus of Liébana (c. 730 – c. 800), which gave subject matter that allowed the brightly coloured primitivist style full scope to demonstrate its qualities in manuscripts of the 10th century like the Morgan Beatus, probably the earliest, the Gerona Beatus (illuminated by a female artist Ende), Escorial Beatus and the Saint-Sever Beatus, which was actually produced some distance from Muslim rule in France. Mozarabic elements, including a background of brightly coloured strips, can be seen in some later Romanesque frescos.