Twistorial formulae for interactions beyond the self-dual sector first arose from Witten's twistor string theory. This is a quantum theory of holomorphic maps of a Riemann surface into twistor space. It gave rise to the remarkably compact RSV (Roiban, Spradlin & Volovich) formulae for tree-level S-matrices of Yang–Mills theories, but its gravity degrees of freedom gave rise to a version of conformal supergravity limiting its applicability; conformal gravity is an unphysical theory containing ghosts, but its interactions are combined with those of Yang–Mills theory in loop amplitudes calculated via twistor string theory.