In 1992, the estimated elephant population size in Sabah ranged from 500-2,000 individuals, based on survey work conducted in the Tabin Wildlife Reserve, in Lower Kinabatangan District and in the Deramakot Forest Reserve. An elephant population census was conducted in Sabah between July 2007 and December 2008, counting dung piles along 216 line transects in five main elephant managed ranges, covering a total distance of . Results of this survey suggest an elephant population of 1,184–3,652 individuals inhabiting the ranges of Tabin, Lower Kinabatangan, North Kinabatangan, Ulu Kalumpang Forest Reserve and the central forest of Sabah. The elephant density and population size varied throughout the five key ranges affected by (i) conversion of lowland forest, (ii) fragmentation of habitat, and (iii) existing land use activities such as logging. The upper catchment of Ulu Segama Forest Reserve had the highest density of elephants with 3.69 elephants per . Only the unprotected central forest area supported an elephant population of more than 1,000 individuals.