Developed from the VS-300, Sikorsky's R-4 was the first large-scale mass-produced helicopter, with a production order for 100 aircraft. The R-4 was the only Allied helicopter to serve in World War II, primarily for search and rescue (by the USAAF 1st Air Commando Group) in the Burma campaign; in Alaska; and in other areas with harsh terrain. Total production reached 131 helicopters before the R-4 was replaced by other Sikorsky helicopters such as the R-5 and the R-6. In all, Sikorsky produced over 400 helicopters before the end of World War II.