Many animals have sensory organs to detect acceleration, especially gravity. In these, the proof mass is usually one or more crystals of calcium carbonate otoliths (Latin for "ear stone") or statoconia, acting against a bed of hairs connected to neurons. The hairs form the springs, with the neurons as sensors. The damping is usually by a fluid. Many vertebrates, including humans, have these structures in their inner ears. Most invertebrates have similar organs, but not as part of their hearing organs. These are called statocysts.