Multiple jugals are known from "Europasaurus", which are more similar in morphology to basal sauropodomorphs than other macronarians. It forms part of the border of the orbit, infratemporal fenestra and the bottom edge of the skull, but does not reach the antorbital fenestra. The posterior process of the jugal are very fragile and narrow, showing a bone scar from the articulation with the quadratojugal. There are two prominences projecting from the back of the jugal body, which diverge at 75º and form the bottom and front edges of the infratemporal fenestra. Like in "Riojasaurus" and "Massospondylus", two non-sauropod sauropodomorphs, the jugal forms a large part of the orbit edge, from the back to the front bottom corner. This feature has been seen in embryos of titanosaurs, but no adult individuals. The quadratojugal bone is an elongate element that has two projecting arms, one anterior and one dorsal. Like in other sauropods, the anterior process is longer than the dorsal, but in "Europasaurus" the arms are more similar lengths. The horizontal process is parallel to the tooth row of "Europasaurus", similar to in "Camarasaurus" but unlike in "Giraffatitan" and "Abydosaurus". There is a prominent ventral flange on the anterior arm of the bone, which is a possibly synapomorphy of Brachiosauridae, although it is also found in some "Camarasaurus" individuals. The two quadratojugal processes diverge at a nearly right angle (90º), although the dorsal process curves as it follows the shape of the quadrate. Squamosals found from "Europasaurus" show the same approximate shape in lateral view as "Camarasaurus", that of a question mark. The squamosals articulate with many skull bones, including those of the skull roof, those of the ventral skull, and those of the braincase. Like the postorbitals, the squamosals are triradiate, with a ventral, anterior and medial