During periods with less disturbance kauri tends to lose ground to broadleaf competitors which can better tolerate shaded environments. In the complete absence of disturbance, kauri tends to become rare as it is excluded by its competitors. Kauri biomass tends to decrease during such times, as more biomass becomes concentrated in angiosperm species like towai. Kauri trees also tend to become more randomly distributed in age, with each tree dying at a different point in time, and regeneration gaps becoming rare and sporadic. Over thousands of years these varying regeneration strategies produce a tug of war effect where kauri retreats uphill during periods of calm, then takes over lower areas briefly during mass disturbances. Although such trends cannot be observed in a human lifetime, research into current patterns of distribution, behavior of species in experimental conditions, and study of pollen sediments (see palynology) have helped shed light on the life history of kauri.