Throughout UCR's history, researchers have developed more than 40 new citrus varieties and invented new techniques to help the $960 million-a-year California citrus industry fight pests and diseases. In 1927, entomologists at the CES introduced two wasps from Australia as natural enemies of a major citrus pest, the citrophilus mealybug, saving growers in Orange County $1 million in annual losses. This event was pivotal in establishing biological control as a practical means of reducing pest populations. In 1963, plant physiologist Charles Coggins proved that application of gibberellic acid allows fruit to remain on citrus trees for extended periods. The ultimate result of his work, which continued through the 1980s, was the extension of the citrus-growing season in California from four to nine months. In 1980, UC Riverside released the Oroblanco grapefruit, its first patented citrus variety. Since then, the citrus breeding program has released other varieties such as the Melogold grapefruit, the Gold Nugget mandarin (or tangerine), and others that have yet to be given trademark names.