The development is indirect and lecithotrophic, and the larvae are ciliated and pigmented, with a deep depression on the ventral side. Astogeny happens when the colony grows through asexual reproduction from the tip of a permanent terminal zooid, behind which the new zooids are budded from the stalk, a type of budding called monopodial. It is possible that in graptolite fossils the terminal zooid was not permanent because the new zooids formed from the tip of latest one, in other words, sympodial budding. These new organisms break a hole in the tubarium wall and start secreting their own tube.