In the first millennium BCE, the Castro culture emerged in northwestern Portugal and Spain in the region extending from the Douro river up to the Minho, but soon expanding north along the coast, and east following the river valleys. It was an autochthonous evolution of Atlantic Bronze Age communities. In 2008, the origins of the Celts were attributed to this period by John T. Koch and supported by Barry Cunliffe. The Ave River Valley in Portugal was the core region of this culture, with a large number of small settlements (the "castros"), but also settlements known as citadels or oppida by the Roman conquerors. These had several rings of walls and the Roman conquest of the citadels of Abobriga, Lambriaca and Cinania around 138 BCE was possible only by prolonged siege. Ruins of notable citadels still exist, and are known by archaeologists as Citânia de Briteiros, Citânia de Sanfins, Cividade de Terroso and Cividade de Bagunte.