According to the genetic study "The Genetic Legacy of Religious Diversity and Intolerance: Paternal Lineages of Christians, Jews, and Muslims in the Iberian Peninsula" at the University Pompeu Fabra of Barcelona and the University of Leicester, led by Briton Mark Jobling, Francesc Calafell, and Elena Bosch, published by the "American Journal of Human Genetics," genetic markers show that nearly 20% of Spaniards have Sephardic Jewish markers (direct male descent male for Y, equivalent weight for female mitochondria); residents of Catalonia have approximately 6%. This shows that there was historic intermarriage between ethnic Jews and other Spaniards, and essentially, that some Jews remained in Spain. Similarly, the study showed that some 11% of the population has DNA associated with the Moors.