It was in 2000 that the historian Philippe Delorme arranged for DNA testing of the heart as well as bone samples from one of the many historical claimants to Louis-Charles's identity, namely, Karl Wilhelm Naundorff, a German clockmaker (see below). Ernst Brinkmann of Münster University and Belgian genetics professor Jean-Jacques Cassiman of the Katholieke Universiteit Leuven, conducted mitochondrial DNA tests using a strand of the hair of the boy's mother, Marie-Antoinette, and other samples from her sisters Maria Johanna Gabriela and Maria Josepha, their mother, Empress Maria Theresa, and two living direct descendants in the strict maternal line of Maria Theresa, namely Queen Anne of Romania and her brother, Prince André de Bourbon Parme, maternal relatives of Louis XVII. The tests proved both that Naundorff was not the dauphin, and the heart was that of Louis-Charles.