Croats settled in the region from about 700 AD onwards. Conquered by King Pepin of Italy, son of Charlemagne, in 789, the Istrian peninsula up to the Kvarner Gulf was incorporated into the Carolingian March of Friuli by 803. In the east, it bordered on the medieval Kingdom of Croatia established by King Tomislav about 925. Having invaded Italy, King Otto of Germany made the Istrian lands part of the vast March of Verona and Aquileia; from the 11th century onwards, the Imperial estates were held by the Patriarchs of Aquileia.