Over the following century, both the Spanish and French established settlements in Florida, with varying degrees of success. In 1559, Spanish Pensacola was established by Don Tristán de Luna y Arellano as the first European settlement in the continental United States, but it had become abandoned by 1561 and would not be reinhabited until the 1690s. French Huguenots founded Fort Caroline in modern-day Jacksonville in 1564, but this fort was conquered by forces from the new Spanish colony of St. Augustine the following year. After Huguenot leader Jean Ribault had learned of the new Spanish threat, he launched an expedition to sack the Spanish settlement; en route, however, severe storms at sea waylaid the expedition, which consisted of most of the colony's men, allowing St. Augustine founder Pedro Menéndez de Avilés time to march his men over land and conquer Fort Caroline. Most of the Huguenots were slaughtered, and Menéndez de Avilés marched south and captured the survivors of the wrecked French fleet, ordering all but a few Catholics executed beside a river subsequently called "Matanzas" (Spanish for 'killings'). When St. Augustine, Florida was established by Pedro Menéndez de Avilés, and hundreds of Spanish-Cuban soldiers and their families moved from Cuba to St. Augustine to establish a new life in 1565. St. Augustine came to serve as the capitals of the British and Spanish colonies of East and West Florida, respectively. The Spanish never had a firm hold on Florida, and maintained tenuous control over the region by converting the local tribes, briefly with Jesuits and later with Franciscan friars. The local leaders (caciques) demonstrated their loyalty to the Spanish by converting to Roman Catholicism and welcoming the Franciscan priests into their villages.