A few fossil traces tentatively associated with the Coelurosauria date back as far as the late Triassic. What has been found between then and the earliest Middle Jurassic is fragmentary. The oldest known unambiguous members of Coelurosauria are the proceratosaurid tyrannosauroids "Proceratosaurus" and "Kileskus" from the late Middle Jurassic. Many nearly complete fossil coelurosaurians are known from the Late Jurassic. "Archaeopteryx" (incl. "Wellnhoferia") is known from Bavaria at 155-150 Ma. "Ornitholestes", the troodontid "Hesperornithoides", "Coelurus fragilis" and "Tanycolagreus topwilsoni" are all known from the Morrison Formation in Wyoming at about 150 Ma. "Epidendrosaurus" and "Pedopenna" are known from the Daohugou Beds in China at about 165-163 Ma. The wide range of fossils in the late Jurassic and morphological evidence shows that coelurosaurian differentiation was virtually complete before the end of the Jurassic.