The opening theme in C major is derived from a motif heard initially in the first movement (at rehearsal 1). Featuring a recurring two sixteenth–eighth note rhythm and a sprightly character, it is traded between strings and woodwinds in quick successive units, maintaining a sense of rhythmic unrest through offbeat accompaniment patterns and hemiolas. A second sweeping theme in C minor begins at rehearsal 93, similar in contour and rhythm to the first movement motive mentioned above. Marked "fortissimo" and "sonoramente" in the strings, it is occasionally supplemented by winds and horns and punctuated by accented off-beats in the brass and timpani. This theme is then repeated "piano" by the first violins with interjecting woodwind solos. The opening material returns in fragments, running through a series of harmonic sequences and building to the entrance of the third theme at rehearsal 100, which maintains the sweeping aspects of the C minor section with the added rhythmic punch of the opening. This is followed by a return to the opening thematic material in earnest.