Meanwhile, finds from the Solnhofen had continued, accounting for the majority of complete high quality specimens discovered. They allowed to identify most new basal taxa, such as "Rhamphorhynchus", "Scaphognathus" and "Dorygnathus". This material gave birth to a German school of pterosaur research, which saw flying reptiles as the warm-blooded, furry and active Mesozoic counterparts of modern bats and birds. In 1882, Marsh and Karl Alfred Zittel published studies about the wing membranes of specimens of "Rhamphorhynchus". German studies continued well into the 1930s, describing new species such as "Anurognathus". In 1927, Ferdinand Broili discovered hair follicles in pterosaur skin, and paleoneurologist Tilly Edinger determined that the brains of pterosaurs more resembled those of birds than modern cold-blooded reptiles.