Among pterodactyloids, a greater variation in diet is present. Pteranodontia contained many piscivorous taxa, such as the Ornithocheirae, Boreopteridae, Pteranodontidae and Nyctosauridae. Niche partitioning caused ornithocheirs and the later nyctosaurids to be aerial dip-feeders like today's frigatebirds (with the exception of the plunge-diving adapted "Alcione elainus"), while boreopterids were freshwater diving animals similar to cormorants, and pteranodonts pelagic plunge-divers akin to boobies and gannets. The istiodactylids were likely primarily scavengers. Archaeopterodactyloidea obtained food in coastal or freshwater habitats. "Germanodactylus" and "Pterodactylus" were piscivores, while the Ctenochasmatidae were suspension feeders, using their numerous fine teeth to filter small organisms from shallow water. "Pterodaustro" was adaptated for flamingo-like filter-feeding.