In contrast, Azhdarchoidea mostly were terrestrial pterosaurs. Tapejaridae were arboreal omnivores, supplementing seeds and fruits with small insects and vertebrates. Dsungaripteridae were specialist molluscivores, using their powerful jaws to crush the shells of molluscs and crustaceans. Thalassodromidae were likely terrestrial carnivores. "Thalassodromeus" itself was named after a fishing method known as "skim-feeding", later understood to be biomechanically impossible. Perhaps it pursued relatively large prey, in view of its reinforced jaw joints and relatively high bite force. Azhdarchidae are now understood to be terrestrial predators akin to ground hornbills or some storks, eating any prey item they could swallow whole. "Hatzegopteryx" was a robustly built predator of relatively large prey, including medium-sized dinosaurs. "Alanqa" may have been a specialist molluscivore. An astonishing study was published in the journal Cretaceous Research. Paleontologists were analyzing the jaw of a pterosaur species Lonchodraco giganteus and found foramina near the tip of its beak and were in a similar pattern like extant birds which would suggest a tactile feeding habit in pterosaurs.