Populations are known to fluctuate with cycles of small rodents and irruptions are known to occur in sub-boreal regions throughout the world. In Scandinavia, populations have been reported to vary from a few hundred birds in certain years to over 4000 birds in others and even up to 10 000 breeding pairs in optimal years. Irruptions can be used as indicators of small mammal abundance and in eastern North America, southern irruptions have been linked with low densities of red-backed voles in the high boreal forest.