Community range or territory size can vary widely, from while group size can range from a single pair to 31 individuals. Population density is also noticeably variable. These wide ranges can be attributed to differing levels of protection and degree of environmental degradation, with better protection and a less degraded environment resulting in higher population density and more moderately sized community ranges. (The duration and seasonality of the studies involved may also have contributed to low group size estimates and community ranges. A study at the Betampona Reserve, for instance, observed monogamous pairs with two to five infants maintaining ranges of .) Core areas at Ambatonikonilahy constituted approximately 10% of the overall community range and showed a close relationship with the location of the largest fruiting trees.