In a 2013 study in Singapore, "Ocypode ceratophthalma" was also discovered to change color in response to the time of day. In a span of 24 hours, they were observed to alternate between lighter and darker coloration, being lightest at midday and darkest at night. However, when placed in a dark background, the crabs showed no significant changes in coloration. This suggests, at least for short-term color changes, they follow a day-night cycle to determine body coloration; basing it on their biological clocks rather than on the brightness or darkness of their environments. The researchers believe relying on the time of day rather than ambient light is more advantageous for survival in ghost crabs. It lets ghost crabs avoid changing color when in temporary shadow (for example within their burrows) and thus still remain inconspicuous when they are once again illuminated by daylight. However, this behavior is only exhibited by juveniles.