Some montane species apparently evolved within their current range, adapting to their local environment, such as the Mount Lyell shrew. However, it has also been noted that some isolated mountain ecosystems have a tendency to lose species over time, perhaps because small, insularized populations are vulnerable to the forces of extinction, and the isolation of the habitat reduces the possibility of colonization by new species. Furthermore, some species, such as the grizzly bear, require a range of habitats. These bears historically made use of the forests and meadows found in the Madrean sky islands, as well as lower-elevation habitats such as riparian zones. (Grizzlies were extirpated from the region in the 20th century.) Seasonal movements between highland and lowland habitats can be a kind of migration, such as that undertaken by the mountain quail of the Great Basin mountains. These birds live in high elevations when free of snow, and instead of migrating south for the winter, they migrate "down".