Florida entered the game with a three-game winning streak started during Florida head coach Urban Meyer's first season at Florida. Tennessee led the series 19-18 all-time against the Gators dating back to 1916. Coming off of an SEC East title the previous season, Tennessee opened SEC play against the Gators after having lost early to UCLA out of conference on the road. The Gators and Volunteers last met in the Swamp in 2007 where the Gators won 59–20. The previous time the two met at Neyland Stadium, the Gators were able to win by a slim one point and then went on to win the BCS National Championship. The Volunteers would be the first of four regular season teams the Gators faced that were coached by a championship-winning head coach (the other three being Les Miles of LSU, Steve Spurrier of South Carolina, and Bobby Bowden of FSU).