Consider the following strategy profiles: the leader plays Cournot; the follower plays Cournot if the leader plays Cournot and the follower plays Stackelberg if the leader plays Stackelberg and if the leader plays something else, the follower plays an arbitrary strategy (hence this actually describes several profiles). This profile is a Nash equilibrium. As argued above, on the equilibrium path play is a best response to a best response. However, playing Cournot would not have been the best response of the leader were it that the follower would play Stackelberg if it (the leader) played Stackelberg. In this case, the best response of the leader would be to play Stackelberg. Hence, what makes this profile (or rather, these profiles) a Nash equilibrium (or rather, Nash equilibria) is the fact that the follower would play non-Stackelberg if the leader were to play Stackelberg.