Approximately during the Gelasian age (Late Pliocene or Early Pleistocene, around 2.5–2 mya), the main lineage of true kestrels emerged; this contains the species characterized by a malar stripe. This too seems to have evolved in Africa and subsequently spread across the Old World until they reached Australia some time during the Middle Pleistocene, less than one million years ago. This group contains several taxa found on Indian Ocean islands. A group of three predominantly grey species from Africa and Madagascar are usually considered kestrels due to their general shape and habits, but are probably distinct from the true kestrels as outlined above.