In 1663 Swammerdam moved to France to continue his studies. He studied one year at the Protestant University of Saumur, under the guidance of Tanaquil Faber. Subsequently, he studied in Paris at the scientific academy of Melchisédech Thévenot. 1665 he returned to the Dutch Republic and joined a group of physicians who performed dissections and published their findings. Between 1666 and 1667 Swammerdam concluded his study of medicine at the University of Leiden. Together with van Horne he researched the anatomy of the uterus. He used waxen injection techniques and a single-lens microscope made by Johannes Hudde. The result of this research was published under the title "Miraculum naturae sive uteri muliebris fabrica" in 1672. Swammerdam received his doctorate in medicine in 1667 under van Horne for his dissertation on the mechanism of respiration, published under the title "De respiratione usuque pulmonum".