Historically, the tightest limits were from direct production at colliders. The first mass limits for squarks and gluinos were made at CERN by the UA1 experiment and the UA2 experiment at the Super Proton Synchrotron. LEP later set very strong limits, which in 2006 were extended by the D0 experiment at the Tevatron. From 2003-2015, WMAP's and Planck's dark matter density measurements have strongly constrained supersymmetry models, which, if they explain dark matter, have to be tuned to invoke a particular mechanism to sufficiently reduce the neutralino density.