With only a few exceptions, primates retain plesiomorphic (original, "primitive") hands with five digits, each equipped with either a nail or a claw. For example, nearly all living strepsirrhine primates have nails on all digits except the second toe which is equipped with a grooming claw. Tarsiers have a grooming claw on second and third toes. Less commonly known, a grooming claw is also found on the second pedal digit of owl monkeys ("Aotus"), titis ("Callicebus"), and possibly other New World monkeys. The needle-clawed bushbaby ("Euoticus") has keeled nails (the thumb and the first and the second toes have claws) featuring a central ridge that ends in a needle-like tip.