Females begin to exhibit signs of senescence around July. The indicative behavior includes resting in flowers, remaining in the nest, or even just falling to the ground from flight. Older individuals also crawl, avoid taking flight, and do not struggle when handled by humans. The old bees die by early August, the same time that juveniles emerge from brood cells. Due to the simultaneous nature of expiration of old bees and emergence of new ones, there is little overlap between generations, except for some females that survive a second winter.