The terrestrial fossil record of the Permian is patchy and temporally discontinuous. Early Permian records are dominated by equatorial Europe and North America, while those of the Middle and Late Permian are dominated by temperate Karoo Supergroup sediments of South Africa and the Ural region of European Russia. Early Permian terrestrial faunas of North America and Europe were dominated by pelycosaurs including the herbivorous edaphosaurids, and carnivorous sphenacodontids, diadectids and amphibians, An extinction, dubbed "Olson's Extinction" is thought to have occurred during the Early-Middle Permian transition, with the most notable effect being the decline of amphibian taxa. The Middle Permian faunas of South Africa and Russia are dominated by primitive therapsids, most abundantly by the diverse Dinocephalia. Dinocephalians become extinct at the end of the Middle Permian, during the Capitanian mass extinction event. Late Permian faunas are dominated by advanced therapsids such as the predatory gorgonopsians and herbivorous dicynodonts, alongside large herbivorous pareiasaur parareptiles. Towards the very end of the Permian the first archosauriforms appeared, a group that would give rise to the pseudosuchians, dinosaurs, and pterosaurs in the following period. Also appearing at the end of the Permian were the first cynodonts, which would go on to evolve into mammals during the Triassic. Another group of therapsids, the therocephalians (such as "Lycosuchus"), arose in the Middle Permian. There were no flying vertebrates (though a family of gliding reptiles known as weigeltisaurs was present in the Late Permian).