The first American Consulate in Japan was opened at the temple of Gyokusen-ji under Consul General Townsend Harris. Harris negotiated the Treaty of Amity and Commerce between the two countries, which was signed at nearby Ryōsen-ji in 1858. Gyokusen-ji is also the location of a small number of foreign graves dating from as early as 1854 marking the final resting place of US forces personnel that died while serving as a part of the Perry Expedition. Japan's relations with Imperial Russia were also negotiated in Shimoda, and in 1855 the Treaty of Shimoda was signed at Chōraku-ji. In June 1859, with the opening of the port of Yokohama to foreign trade, the port of Shimoda was again closed and the American consulate was relocated to Zenpuku-ji in Edo.