Pterosaurs sported coats of hair-like filaments known as pycnofibers, which covered their bodies and parts of their wings. Pycnofibers grew in several forms, from simple filaments to branching down feathers. These are possibly homologous to the down feathers found on both avian and some non-avian dinosaurs, suggesting that early feathers evolved in the common ancestor of pterosaurs and dinosaurs, possibly as insulation. In life, pterosaurs would have had smooth or fluffy coats that did not resemble bird feathers. They were warm-blooded (endothermic) active animals. The respiratory system had efficient unidirectional "flow-through" breathing using air sacs, which hollowed out their bones to an extreme extent. Pterosaurs spanned a wide range of adult sizes, from the very small anurognathids to the largest known flying creatures of all time, including "Quetzalcoatlus" and "Hatzegopteryx", which reached wingspans of at least nine metres. The combination of endothermy, a good oxygen supply and strong muscles made pterosaurs powerful and capable flyers.