Some affiliative behaviors are seasonal or gender-specific, such as the male squeal approach and anogenital inspections performed during the mating season. Another example is the female greeting behavior, where two females will use their anogenital scent glands to mark each other's backs, jump over one another, writhe together, and emit squealing vocalizations. This behavior is not seen during the end of the cool, dry season or around gestation. The frequency of other affiliative behaviors can be affected by age. All ruffed lemurs over five months of age allogroom, and, in captivity, subadults participate in play more frequently than adults.