In general, the mixing of the soil by the activities of animals, sometimes called pedoturbation, tends to undo or counteract the tendency of other soil-forming processes that create distinct horizons. Termites and ants may also retard soil profile development by denuding large areas of soil around their nests, leading to increased loss of soil by erosion. Large animals such as gophers, moles, and prairie dogs bore into the lower soil horizons, bringing materials to the surface. Their tunnels are often open to the surface, encouraging the movement of water and air into the subsurface layers. In localized areas, they enhance mixing of the lower and upper horizons by creating, and later refilling the tunnels. Old animal burrows in the lower horizons often become filled with soil material from the overlying A horizon, creating profile features known as crotovinas.