The influence of Greek art, however, led to "heroic" nude portrayals of Roman men and gods, a practice that began in the 2nd century BC. When statues of Roman generals nude in the manner of Hellenistic kings first began to be displayed, they were shocking not simply because they exposed the male figure, but because they evoked concepts of royalty and divinity that were contrary to Republican ideals of citizenship as embodied by the toga. The god Mars is presented as a mature, bearded man in the attire of a Roman general when he is conceived of as the dignified father of the Roman people, while depictions of Mars as youthful, beardless, and nude show the influence of the Greek Ares. In art produced under Augustus, the programmatic adoption of Hellenistic and Neo-Attic style led to more complex signification of the male body shown nude, partially nude, or costumed in a muscle cuirass.