A bone unique to pterosaurs, known as the pteroid, connected to the wrist and helped to support the forward membrane (the propatagium) between the wrist and shoulder. Evidence of webbing between the three free fingers of the pterosaur forelimb suggests that this forward membrane may have been more extensive than the simple pteroid-to-shoulder connection traditionally depicted in life restorations. The position of the pteroid bone itself has been controversial. Some scientists, notably Matthew Wilkinson, have argued that the pteroid pointed forward, extending the forward membrane and allowing it to function as an adjustable flap. This view was contradicted in a 2007 paper by Chris Bennett, who showed that the pteroid did not articulate as previously thought and could not have pointed forward, but rather was directed inward toward the body as traditionally interpreted. David Peters in 2009 proposed that the pteroid articulated with the 'saddle' of the radiale (proximal syncarpal) and both the pteroid and preaxial carpal were migrated centralia. This seems to be confirmed by specimens of "Changchengopterus pani" and "Darwinopterus linglongtaensis" showing the pteroid in articulation with the proximal syncarpal.