Some Rhamphorhynchidae, such as "Rhamphorhynchus" itself or "Dorygnathus", were fish-eaters with long, slender wings, needle-like dentition and long, thin jaws. "Sericipterus", "Scaphognathus" and "Harpactognathus" had more robust jaws and teeth (which were ziphodont, dagger-shaped, in " Sericipterus"), and shorter, broader wings. These were either terrestrial/aerial predators of vertebrates or corvid-like generalists. Wukongopteridae like "Darwinopterus" were first considered aerial predators. Lacking a robust jaw structure or powerful flying muscles, they are now seen as arboreal or semiterrestrial insectivores. "Darwinopterus robustidens", in particular, seems to have been a beetle specialist.