The two modern parvorders of cetaceans – Mysticeti (baleen whales) and Odontoceti (toothed whales) – are thought to have separated from each other around 28-33 million years ago in a second cetacean radiation, the first occurring with the archaeocetes. The adaptation of animal echolocation in toothed whales distinguishes them from fully aquatic archaeocetes and early baleen whales. The presence of baleen in baleen whales occurred gradually, with earlier varieties having very little baleen, and their size is linked to baleen dependence (and subsequent increase in filter feeding).