The mature boreal forest pattern in the south shows balsam fir dominant on well-drained sites in eastern Canada changing centrally and westward to a prominence of white spruce, with black spruce and tamarack forming the forests on peats, and with jack pine usually present on dry sites except in the extreme east, where it is absent. The effects of fires are inextricably woven into the patterns of vegetation on the landscape, which in the east favour black spruce, paper birch, and jack pine over balsam fir, and in the west give the advantage to aspen, jack pine, black spruce, and birch over white spruce. Many investigators have reported the ubiquity of charcoal under the forest floor and in the upper soil profile. Charcoal in soils provided Bryson et al. (1965) with clues about the forest history of an area 280 km north of the then current tree line at Ennadai Lake, District Keewatin, Northwest Territories.