Reproduction is typically seasonal, with more young being born during the rainy season when there is an abundance of fruit. During the dry season, fewer babies are born due to the lack of available fruit. Litter sizes averaging between four and five young, with each female producing between two and four litters per year. Many young die while nursing in the mother's pouch. This death rate is especially high during the dry season. A major factor that determines survival of young is the mother's age; there are many deaths when the mother is less than 11 months.