At the peak of its operational deployment, the Pup equipped only four RNAS squadrons (Nos. 3, 4, 8 and 9), and three RFC squadrons (Nos. 54, 46 and 66). By the spring of 1917, the Pup had been outclassed by the newest German fighters. The RNAS replaced their Pups, first with Sopwith Triplanes, and then with Sopwith Camels. The RFC soldiered on with Pups, in spite of increasing casualties, until it was possible to replace them with Camels in December 1917.