Males are characterized as weakly territorial prior to mating, however, there has been some evidence on territory features correlating with female settlement, thus female reproductive success. Studies have shown that shade is an important resource for female reproductive success. As a fitness-limiting feature, it would be reasonable to expect this could account for any existing tendencies for males being territorial. However, levels of male aggression do not change, which lends to the conclusion that female choice corresponds with changes in fitness-indicator traits. Extra-pair mating has been corresponded with nesting site quality, adding further characterization to male aggression. Many studies have showed territory characteristics as important for mate acquisition, however a recent study shows evidence of its decreased role comparatively to female choice.