Punctuation developed dramatically when large numbers of copies of the Bible started to be produced. These were designed to be read aloud, so the copyists began to introduce a range of marks to aid the reader, including indentation, various punctuation marks (diple, , ), and an early version of initial capitals (). Jerome and his colleagues, who made a translation of the Bible into Latin, the "Vulgate" (), employed a layout system based on established practices for teaching the speeches of Demosthenes and Cicero. Under his layout every sense-unit was indented and given its own line. This layout was solely used for biblical manuscripts during the 5th–9th centuries but was abandoned in favor of punctuation.