Book II contains his scientific determination of the nature of the soul, an element of his biology. By dividing substance into its three meanings (matter, form, and what is composed of both), he shows that the soul must be the first actuality of a naturally organised body. This is its form or essence. It cannot be matter because the soul is that in virtue of which things have life, and matter is only being in potency. The rest of the book is divided into a determination of the nature of the nutritive and sensitive souls. (1) All species of living things, plant or animal, must be able to nourish themselves and reproduce others of the same kind. (2) All animals have, in addition to the nutritive power, sense-perception, and thus they all have at least the sense of touch, which he argues is presupposed by all other senses, and the ability to feel pleasure and pain, which is the simplest kind of perception. If they can feel pleasure and pain they also have desire. Some animals in addition have other senses (sight, hearing, taste), and some have more subtle versions of each (the ability to distinguish objects in a complex way, beyond mere pleasure and pain.) He discusses how these function. Some animals have in addition the powers of memory, imagination, and self-motion.