Adult females, subadults, and juveniles are moderate in build. Distinguishing the sexes is difficult until about age five. The body is modest in size and the neck, chest, and shoulders are sized in proportion with the torso. Adult females and subadults have more complex and variable coloration than adult males. They are dark silver-gray to charcoal above. The flanks, chest, sides, and underside of the neck, often forming a chevron pattern in this area, are cream to tan with rusty tones. Variable cream to rust-colored areas are on the sides and top of the muzzle, chin, and as a "brush stroke" running backwards under the eye. In contrast, adult males are medium gray to black, or reddish to dark brown all over. Their manes can have variable amounts of silver-gray or yellowish tinting on the guard hairs. Pups are blackish at birth, with variable oval areas of buff on the sides, in the axillary area, and on the chin and sides of the muzzle. After three to four months, pups molt to the color of adult females and subadults.