The first use of aircraft registrations was based on the radio callsigns allocated at the London International Radiotelegraphic Conference in 1913. The format was a single letter prefix followed by four other letters (like A-BCDE). The major nations operating aircraft were allocated a single letter prefix. Smaller countries had to share a single letter prefix, but were allocated exclusive use of the first letter of the suffix. This was modified by agreement by the International Bureau at Berne and published on April 23, 1913. Although initial allocations were not specifically for aircraft but for any radio user, the International Air Navigation Convention held in Paris in 1919 (Paris Convention of 1919) made allocations specifically for aircraft registrations, based on the 1913 callsign list. The agreement stipulated that the nationality marks were to be followed by a hyphen then a group of four letters that must include a vowel (and for the convention Y was considered to be a vowel). This system operated until the adoption of the revised system in 1928.