Dormancy caused by an impermeable seed coat is known as physical dormancy. Physical dormancy is the result of impermeable layer(s) that develops during maturation and drying of the seed or fruit. This impermeable layer prevents the seed from taking up water or gases. As a result, the seed is prevented from germinating until dormancy is broken. In natural systems, physical dormancy is broken by several factors including high temperatures, fluctuating temperatures, fire, freezing/thawing, drying or passage through the digestive tracts of animals. Physical dormancy is believed to have developed >100 mya.