The defence of the Scottish Border was carried out by two Wardens– that of the East March (based at Berwick-upon-Tweed) and that of the West March at Carlisle. Both offices had been held by the Percy family in the fourteenth century, and their support of King Henry IV seemed to have paid off in 1399, when Henry Percy, 1st Earl of Northumberland was appointed Warden of the West March and his son Henry Percy ("Hotspur") as Warden of the East March. But Hotspur rebelled, and his father was held to be complicit in his treason. After Hotspur was killed at the Battle of Shrewsbury in 1403, Ralph Neville was employed by King Henry V to capture the elder Percy. His reward was to succeed the Percys as Warden of both Marches. Under King Henry V, the Percys were restored to their lands, and eventually in 1417, to the East March. The West March, however, was to become an almost hereditary Neville appointment.