Seeds germinate in spring and summer. Those that germinate in autumn produce plants that overwinter if they are large enough, while rosettes less than across die in winter. After flowering the entire plant usually dies at the end of its second year, but some individuals, especially in the northern parts of the range, require a longer growth period and flower in their third year. Under better growing conditions, some individuals flower in the first year. Triennial individuals have been found to produce fewer seeds than biennial and annual ones. While year of flowering and size are linked to the environment, most other characteristics appear to be genetic.