The Big Lemon originally operated an express service numbered 42X which ran from Brighton station to Falmer station using elderly step entranced buses. However the service was dropped in December 2007. A relaunch of the service commenced in early 2008 numbered 42. In 2010 the company started two more services, 43 and 44. However just months after it began route 43 ended due to low passenger numbers. In 2011 The Big Lemon faced competition from Brighton & Hove Buses in that Brighton Buses lowered its fares to match fares charged by The Big Lemon. In January 2012 The Big Lemon stopped running route 42 and continued to run route 44 only, split into two shuttles numbered UB1 and UB2. The Big Lemon subsequently ceased operation of these services, and shuttle UB1 is operated by the University of Brighton.