Virgin Trains retained one complete Mark 3 set. It was initially used to cover peak-time London Euston to Birmingham services while the Pendolinos underwent modifications, but later covering the loss of 390033, written off after the Grayrigg derailment in 2007. In July 2009, it was refurbished and repainted at Doncaster Works in the same style as the Pendolino and Voyager fleets, but with the British Rail seats and interior fittings retained. This set was nicknamed the "Pretendolino" by enthusiasts. It was transferred to Abellio Greater Anglia in November 2014. In 2016 it moved to TransPennine Express to be used as a crew trainer in the lead-up to the introduction of Mark 5s.