In England of 950, King Athelstan imposed an annual tribute of 300 wolf skins on Welsh king Hywel Dda, an imposition which was maintained until the Norman conquest of England. At the time, several criminals, rather than being put to death, would be ordered to provide a certain number of wolf tongues annually. The Norman kings (reigning from 1066 to 1154) employed servants as wolf hunters and many held lands granted on condition they fulfilled this duty. William the Conqueror granted the lordship of Riddesdale in Northumberland to Robert de Umfraville on condition that he defend that land from enemies and wolves. There were no restrictions or penalties in the hunting of wolves, except in royal game reserves, under the reasoning that the temptation for a commoner to shoot a deer there was too great. King John gave a premium of 10 shillings for the capture of two wolves. King Edward VI who reigned from 1272 to 1307 ordered the total extermination of all wolves in his kingdom and personally employed one Peter Corbet, with instructions to destroy wolves in the counties of Gloucestershire, Herefordshire, Worcestershire, Shropshire and Staffordshire, areas near the Welsh Marches where wolves were more common than in the southern areas of England. James I of Scotland passed a law in 1427 requiring three wolf hunts a year between April 25 to August 1, coinciding with the wolf's cubbing season. The wolf became extinct in England during the reign of Henry VII (1485–1509).