The capelin or caplin ("Mallotus villosus") is a small forage fish of the smelt family found in the North Atlantic, North Pacific, and Arctic oceans. In summer, it grazes on dense swarms of plankton at the edge of the ice shelf. Larger capelin also eat a great deal of krill and other crustaceans. Among others, whales, seals, Atlantic cod, Atlantic mackerel, squid, and seabirds prey on capelin, in particular during the spawning season while the capelin migrate south. Capelin spawn on sand and gravel bottoms or sandy beaches at the age of two to six years. When spawning on beaches, capelin have an extremely high post-spawning mortality rate which, for males, is close to 100%.