In 1800, the Nez Perce had more than 100 permanent villages, ranging from 50 to 600 individuals, depending on the season and social grouping. Archeologists have identified a total of about 300 related sites including camps and villages, mostly in the Salmon River Canyon. In 1805, the Nez Perce were the largest tribe on the Columbia River Plateau, with a population of about 6,000. By the beginning of the 20th century, the Nez Perce had declined to about 1,800 due to epidemics, conflicts with non-Indians, and other factors. A total of 3,499 Nez Perce were counted in the 2010 Census.