These conditions are responsible for what has been called the Sahara pump theory. During periods of a wet or "Green Sahara", the Sahara becomes a savanna grassland and various flora and fauna become more common. Following inter-pluvial arid periods, the Sahara area then reverts to desert conditions and the flora and fauna are forced to retreat northwards to the Atlas Mountains, southwards into West Africa, or eastwards into the Nile Valley. This separates populations of some of the species in areas with different climates, forcing them to adapt, possibly giving rise to allopatric speciation.