The glabella was inflated, and globular-shaped, and covered in small wart-like bumps. If it was filled with fat, or oil, the glabellum would have helped to have made the creature positively buoyant. On the other hand, trilobites with large glabellae are often suspected of being predatory, as the volume of glabella would be filled with digestive organs, or used to store captured/swallowed prey. The free cheeks of the cephalon formed a pair of long, curved spines, and the segments of the pleural lobes were separated and elongated to form rib-like struts. These modifications, along with the "V" shaped pygidium give these trilobites a cartoon "fish-skeleton" appearance. The defensive value of these highly elongate spines is also apparent, as they would have stuck in the throats of vertebrate predators, such as the Silurian acanthodian "Nostolepis".