However, the increasing numerical strength of Allied bomber units, along with inadequate anti-aircraft systems, imposed crippling losses on Japanese units. Approximately 100 out of 130 Japanese aircraft based in the Wewak area were lost during the attacks of August 17–21 1943. By the end of the campaign, nearly 2,000 Japanese aircraft had been lost in air attacks from up to 200 Allied aircraft at a time, around half of which were Consolidated B-24 Liberators and North American B-25 Mitchells armed with fragmentation bombs. After the Japanese retreat, over 340 aircraft wrecks were later found at Hollandia.