It was used much in the same way by later scholars to mark differences between various translations or versions of the Bible and other manuscripts. The early Christian Alexandrian scholar Origen (c. 184–253 AD) used it to indicate differences between different versions of the Old Testament in his "Hexapla". Epiphanius of Salamis (c. 310–320 – 403) used both a horizontal slash or hook (with or without dots) and an upright and slightly slanting dagger to represent an obelus. St. Jerome (c. 347–420) used a simple horizontal slash for an obelus, but only for passages in the Old Testament. He describes the use of the asterisk and the dagger as: "an asterisk makes a light shine, the obelisk cuts and pierces".