The mechanism of seed dispersal via animal link vectors (endozoochoric dispersal) may explain how most Burseraceae were able to expand their range so efficiently across the globe. "Beiselia, Boswellia", and "Triomma" have dry fruits better suited for wind dispersal, but most Burseraceae have fleshy, edible fruit that is eaten by many animal dispersers. The seeds may provide a high reward in fat (24–73%) and protein (2.7–25.9%) if digested, but many animals eat just the fleshy part of the fruit and either discard the endocarp right away or excrete it some time later. Some known Burseraceae fruit consumers include hornbills ("Buceros bicornis, Ceratogyma atrata, C. cylindricus, Penelopides panini"), oilbirds ("Steatnoris caripensis"), fruit pigeons, warblers, vireos, orioles, flycatchers, tanagers, woodpeckers, loeries, primates ("Cercopithecus" spp., "Lophocebus albigena"), lemurs ("Varecia variegate" subsp. "variegate"), and sun bears ("Helarctos malayanus"). The fruits may also have been water dispersed.