A second species described in 1988 by Sereno and Zhao, along with two Chinese colleagues, was "P. meileyingensis" from the Jiufotang Formation, near the town of Meileyingzi, Liaoning Province, northeastern China. This species is known from four fossil skulls, one associated with some skeletal material, found in 1973 by Chinese scientists. The age of the Jiufotang in Liaoning is unknown, but in the neighbouring province of Inner Mongolia, it has been dated to about 110 Ma, in the Albian stage of the Early Cretaceous.