All dinoflagellates contain red algal plastids or remnant (nonphotosynthetic) organelles of red algal origin. The parasitic dinoflagellate "Hematodinium" however lacks a plastid entirely. Some groups that have lost the photosynthetic properties of their original red algae plastids has obtained new photosynthetic plastids (chloroplasts) through so-called serial endosymbiosis, both secondary and tertiary. Like their original plastids, the new chloroplasts in these groups can be traced back to red algae, except from those in the members of the genus Lepidodinium, which possess plastids derived from green algae, possibly Trebouxiophyceae or Ulvophyceae. Lineages with tertiary endosymbiosis are Dinophysis, with plastids from a cryptomonad, the Karenia, Karlodinium, and Takayama, which possess plastids of haptophyte origin, and the Peridiniaceae, Durinskia and Kryptoperidinium, which has plastids derived from diatoms Some species also perform kleptoplasty.