Pisces, or Pi (Freddie Prinze, Jr.), is a young orange wrasse that lives happily with his parents Pike and Piper in the polluted harbor of Boston, Massachusetts, until a fishing boat scoops them from the sea. Pi's parents manage to help him escape, but cannot escape themselves. Before they are taken away, Piper tells Pi to promise her he would go live with his aunt. Pi's porpoise friend Percy (Trent Ford) and Percy's mother Meg (Megahn Perry) agree to take him, but Pi refuses to leave in case his parents return, Meg sadly tells him that once anyone is taken by a net they are killed. Pi is heartbroken that his parents are gone forever.