"Prestosuchus" has a "crocodile-normal" crurotarsal ankle, with a convex-concave suture between the astragalus and calcaneum. The astragalus has a large tibial facet and small fibular facet on its upper surface, a curved depression on its front surface, and facets for distal tarsal III and metatarsals I-II on its lower surface. The calcaneum is wide and has a convex fibular knob and calcaneal tuber, as with other crurotarsal archosaurs. The calcaneal tuber is wider than long and has a tall boxy expansion at its rear. The underside of the calcaneum has a large facet for distal tarsal IV and metatarsals III-IV, followed by a teardrop-shaped depression. The two distal tarsals (III and IV) were roughly triangular. Metatarsal III is the longest bone in the foot, while the thick and hooked metatarsal V is the shortest, followed by metatarsal I. All the metatarsals had strongly developed ginglymoidal joints. I-III and V are triangular in cross-section, while IV is hourglass shaped. The foot has a phalangeal formula of 2-3-4-3-X, indicating that the fourth toe was much smaller than the second and third, an unusual trait among early loricatans. The first toe had a large curved claw, the third toe was thick, and the second toe had both traits.