The bulk and mass of individual crocodiles can be fairly variable, some animals being relatively slender, while others being very robust; females are often bulkier than males of a similar length. As an example of the body mass increase undergone by mature crocodiles, one of the larger crocodiles handled firsthand by Cott (1961) was and weighed , while the largest specimen measured by Graham and Beard (1973) was and weighed more than . In attempts to parse the mean male and female lengths across the species, the mean adult length was estimated to be reportedly in males, at which males would average about in weight, while that of the female is , at which females would average about . This gives the Nile crocodile somewhat of a size advantage over the next largest non-marine predator on the African continent, the lion ("Panthera leo"), which averages in males and in females, and attains a maximum known weight of , far less than that of large male crocodiles.