Ryll is a deformed Lyrinx male who has only stubby useless wings and is a very talented flesh former. During the course of the series, he creates many of the Lyrinx's most dangerous flesh formed creations. By the end of "Chimaera", he is appointed as the first Patriarch of the Lyrinx for three thousand years. Tiaan and Ryll have saved each other's lives several times and despite the war between their species, they come to regard each other as friends. He has romantic feelings towards Liett though he is often confused about her feelings towards him.