Rodents and shrews may enter the diet of juvenile crocodiles, i.e. , and become commonplace in subadult and small adult crocodiles. Species recorded include the Natal multimammate mouse ("Mastomys natalensis"), African marsh rat ("Dasymys incomtus"), common rufous-nosed rat ("Oenomys hypoxanthus"), and savanna swamp shrew ("Crocidura longipes"). In many areas, the cane rats are a particular favorite mammalian food for crocodiles, particularly the relatively large greater cane rat ("Thryonomys swinderianus"). In Uganda and Zambia, the latter species are the leading overall mammalian prey type for crocodiles and one Kenyan crocodile of in length had 40 greater cane rats in its stomach. Cape porcupines ("Hystrix africaeaustralis") are known to have been preyed on several times in Kruger National Park, their quills apparently being an insufficient defense against the tough jaws and digestive systems of crocodiles. Small carnivores are readily taken opportunistically, including both African clawless otters ("Aonyx capensis") and spotted-necked otters ("Hydrictis maculicollis"), as well as water mongoose ("Atilax paludinosus"), African wildcats ("Felis silvestris lybica") and servals ("Leptailurus serval").