Preserved fossilized eurypterid trackways tend to be large and heteropodous and often have an associated telson drag mark along the mid-line (as with the Scottish "Hibbertopterus" track). Such trackways have been discovered on every continent except for South America. In some places where eurypterid fossil remains are otherwise rare, such as in South Africa and the rest of the former supercontinent Gondwana, the discoveries of trackways both predate and outnumber eurypterid body fossils. Eurypterid trackways have been referred to several ichnogenera, most notably "Palmichnium" (defined as a series of four tracks often with an associated drag mark in the mid-line), wherein the holotype of the ichnospecies "P. kosinkiorum" preserves the largest eurypterid footprints known to date with the found tracks each being about in diameter. Other eurypterid ichnogenera include "Merostomichnites" (though it is likely that many specimens actually represent trackways of crustaceans) and "Arcuites" (which preserves grooves made by the swimming appendages).