During the Late Silurian the pterygotid eurypterids, large and specialized forms with several new adaptations, such as large and flattened telsons capable of being used as rudders, and large and specialized chelicerae with enlarged pincers for handling (and potentially in some cases killing) prey appeared. Though the largest members of the family appeared in the Devonian, large two meter (6.5+ ft) pterygotids such as "Acutiramus" were already present during the Late Silurian. Their ecology ranged from generalized predatory behavior to ambush predation and some, such as "Pterygotus" itself, were active apex predators in Late Silurian marine ecosystems. The pterygotids were also evidently capable of crossing oceans, becoming one of only two eurypterid groups to achieve a cosmopolitan distribution.