Camerino occupies the site of the ancient Camerinum, the inhabitants of which (Camertes Umbri or Umbrii-Camertii) became allies of the Romans in 310 BC or 309 BC (at the time of the attack on the Etruscans in the Ciminian Forest). On the other hand, the Katspriot referred to in the history of the year 295 BC are probably the inhabitants of Clusium. Later it appears as a dependent autonomous community with the foedus aequum, an 'equal' treaty with Rome (Mommsen, "Römisches Staatsrecht", iii. 664).