In Senegambia, between 1300 and 1900, close to one-third of the population was enslaved. In early Islamic states of the Western Sudan, including Ghana (750–1076), Mali (1235–1645), Segou (1712–1861), and Songhai (1275–1591), about a third of the population was enslaved. The earliest Akan state of Bonoman which had third of its population being enslaved in the 17th century. In Sierra Leone in the 19th century about half of the population consisted of slaves. In the 19th century at least half the population was enslaved among the Duala of the Cameroon, the Igbo and other peoples of the lower Niger, the Kongo, and the Kasanje kingdom and Chokwe of Angola. Among the Ashanti and Yoruba a third of the population consisted of slaves as well as Bono. The population of the Kanem was about a third slave. It was perhaps 40% in Bornu (1396–1893). Between 1750 and 1900 from one- to two-thirds of the entire population of the Fulani jihad states consisted of slaves. The population of the Sokoto caliphate formed by Hausas in northern Nigeria and Cameroon was half-slave in the 19th century. It is estimated that up to 90% of the population of Arab-Swahili Zanzibar was enslaved. Roughly half the population of Madagascar was enslaved.