In both the Nicoya and Atlantic-Watershed regions, metates are often made with saurian (specifically crocodile, alligator, or caiman) imagery. It is thought that the saurian represents the surface of the earth, which relates to agricultural fertility. One of the oldest and most prominent themes in Chibcha art is that of the Crocodile god. Depicted as an anthropomorphic being with a crocodile head, he has been carved into fly-panel metates, sometimes shown standing on a double-headed saurian and other times on a jaguar. As a symbol, the double-headed Saurian has the longest use and distribution of any iconographic element in the Isthmo-Columbian area.