The war involved multi-ethnic empires such as Great Britain, France, Germany, Russia, Austria-Hungary and Turkey. Many ethnic groups in these territories were conscripted for military service. The casualties listed by modern borders are also included in the above table of figures for the countries that existed in 1914. The casualty figures by 1924 post war borders are rough estimates by Russian historian Vadim Erlikman in a 2004 handbook of human losses in the 20th century, the sources of his figures were published in the Soviet era and in post-Soviet Russia. According to the 1914–1918 Online Encyclopedia "In addition to losses suffered by African military personnel and the laborers supporting their operations, very large, but unknown numbers of African civilians perished during the war." They made an estimate of civilian losses in Africa of 750,000 based on the study by the Vadim Erlikman. They noted that Erlikman's figures are based on the work of the Russian demographer Boris Urlanis, they noted that these estimates were "imprecise" and "could be used to provide a frame of reference for further inquiry". The "Oxford History of World War One" notes that "In east and central Africa the harshness of the war resulted in acute shortages of food with famine in some areas, a weakening of populations, and epidemic diseases which killed hundreds of thousands of people and also cattle."