The most recent study on Chechens, by Balanovsky "et al." in 2011 sampled a total of 330 Chechens from three sample locations (one in Malgobek, one in Achkhoy-Martan, and one from two sites in Dagestan) and found the following frequencies: A weak majority of Chechens belong to Haplogroup J2 (56.7%), which is associated with Mediterranean, Caucasian and Fertile Crescent populations. Other notable values were found among North Caucasian Turkic peoples (Kumyks (25%) and Balkars (24%)). It is notable that J2 suddenly collapses as one enters the territory of non-Nakh Northeast Caucasian peoples, dropping to very low values among Dagestani peoples. The overwhelming bulk of Chechen J2 is of the subclade J2a4b* (J2-M67), of which the highest frequencies by far are found among Nakh peoples: Chechens were 55.2% according to the Balanovsky study, while Ingush were 87.4%. Other notable haplogroups that appeared consistently appeared at high frequencies included J1 (20.9%), L (7.0%), G2 (5.5%), R1a (3.9%), Q-M242 (3%) and R1b-M269 (1.8%, but much higher in Chechnya itself as opposed to Dagestani or Ingushetian Chechens). Overall, tests have shown consistently that Chechens are most closely related to Ingush, Circassians and other North Caucasians, occasionally showing a kinship to other peoples in some tests. Balanovsky's study showed the Ingush to be the Chechens' closest relatives by far.