The caves have been used by humans for around 45,000 years, demonstrated by the discovery of tools from the Palaeolithic period, along with fossilised animal remains. Evidence of Stone and Iron Age occupation continued into Roman Britain. A corn-grinding mill operated on the resurgent waters of the River Axe as early as the Domesday survey of 1086. The waters of the river are used in a handmade paper mill, the oldest extant in Britain, which began operations circa 1610. The low, constant temperature of the caves means that they can be used for maturing Cheddar cheese.