Habitat requirements for the Indiana bat are not completely understood. Bottomland and floodplain forests were once thought to be the most important habitats during the summer, but subsequent study has shown that upland forest habitats may be equally important, especially in the southern portions of the species's range. Indiana bats are found in hardwood forests throughout most of their range and mixed hardwood-pine forests in the southeastern United States. Stone and Battle found a significantly greater proportion (p<0.05) of old-growth forest (greater than 100 years), more hardwoods, and fewer conifers in stands occupied by Indiana bats than in random stands in Alabama.