The Conowingo Dam, and to a lesser extent the Holtwood and Safe Harbor Dams further upstream, stopped migratory fish species, especially American shad, from swimming further up the Susquehanna River to spawn. In 1984, a fish capture feature was added at Conowingo and shad were trucked upstream above all three dams and released. This program ended in 1999. A fish lift was installed in 1991. All three dams completed installation of fish lifts in time for the 2000 season. During the 2000 migration season, 153,000 American shad passed through the Conowingo fish lift. "However, passage rates of shad from Conowingo to Holtwood have been only 30 to 50 percent, suggesting that fish are having difficulty moving upstream in the waters of the Conowingo pool."