Regional population control has been used on species that cannot be excluded from the airport environment. A nesting colony of laughing gulls at Jamaica Bay Wildlife Refuge contributed to 98–315 bird strikes per year, in 1979–1992, at adjacent John F. Kennedy International Airport (JFK). Though JFK had an active bird management program that precluded birds from feeding and loafing on the airport, it did not stop them from overflying the airport to other feeding sites. U.S. Department of Agriculture Wildlife Services personnel began shooting all gulls that flew over the airport, hypothesizing that eventually, the gulls would alter their flight patterns. They shot 28,352 gulls in two years (approximately half of the population at Jamaica Bay and 5–6% of the nationwide population per year). Strikes with laughing gulls decreased by 89% by 1992. However this was more a function of the population reduction than the gulls altering their flight pattern (Dolbeer et al. 1993, Dolbeer et al. 2003, DeVault et al. 2013).