Father Brian Finn (Edward Norton), dedicated to his calling as Catholic priest since childhood, shares the duties of his New York parish with the older Fr. Havel (Miloš Forman). Rabbi Jacob "Jake" Schram (Ben Stiller), best friends with Brian since they were children and the youngest rabbi at his synagogue, focuses on his work to the detriment of his private life, much to the chagrin of his mother, Ruth (Anne Bancroft). The two men show a close bond even in their professions, planning to open a jointly-sponsored community center. The pair reminisce about their childhood friend Anna Reilly (Jenna Elfman), meeting them in middle school after beating up a bully. The three were close friends until Anna's family moved to California and they ultimately lost touch.