Stands occupied by this species can vary greatly. A Virginia pine roost was in a stand with a density of only 367 trees/ha, while in Kentucky, a shagbark hickory roost was in a closed-canopy stand with 1,210 trees/ha. Overall tree density in Great Smoky Mountain National Park was higher around primary roosts than at alternate roosts. At the landscape level, the basal area for stands with roosts was 30% lower than basal area of random stands in Alabama. Tree density in southern Iowa varied between different habitats. In a forested floodplain, tree density was lowest at 229 trees/ha, while a riparian strip had the highest tree density at 493 trees/ha.