Officially established by Kujula Kadphises, the first Kushan emperor who united the Yuezhi tribes, the Kushan empire was a syncretic empire in central and southern Asia, including the regions of Gandhara and Mathura in northern India. From 127 to 151 CE, Gandharan reached its peak under the reign of Kanishka the Great. In this period, Kushan art inherited the Greco-Buddhist art. Mahayana Buddhism flourished, and the depictions of Buddha as a human form first appeared in art. Wearing a monk's robe and a long length of cloth draped over the left shoulder and around the body, the Buddha was depicted with 32 major lakshanas (distinguishing marks), including a golden-colored body, an ushnisha (a protuberance) on the top of his head, heavy earrings, elongated earlobes, long arms, the impression of a chakra (wheel) on the palms of his hands and the soles of his feet, and the urna (a mark between his eyebrows). One of the hallmarks of Gandharan art is its relation to naturalism of Hellenistic art. The naturalistic features found in Gandharan sculptures include the three-dimensional treatment of the drapery, with unregularized folds that are in realistic patterns of random shape and thickness. The physical form of the Buddha and his bodhisattvas are well-defined, solid, and muscular, with swelling chests, arms, and abdomens. Buddhism and Buddhism art spread to Central Asia and the far East across Bactria and Sogdia, where the Kushan Empire met the Han Dynasty of China.