The palmette is related to a range of motifs in differing cultures and periods. In ancient Egypt palmette motifs existed both as a form of flower and as a stylized tree, often referred to as a Tree of life. Other examples from ancient Egypt are the alternating lotus flower and bud border designs, the winged disk of Horus with its pair of Uraeus serpents, the Eye of Horus and curve-topped commemorative stele. In later Assyrian versions of the Tree of Life, the feathered falcon wings of the Egyptian winged disk have become associated with the fronds of the palm tree. Similar lotus flower and bud borders, closely associated with palmettes and rosettes, also appeared in Mesopotamia. There appears to be an equivalence between the horns of horned creatures, the wings of winged beings including angels, griffins and sphinxes and both the fan and the volutes of the palmette; there is also an underlying 'V' shape in each of these forms that parallels the association of the palm itself with victory, energy and optimism.