Elephant dung piles collected in Kahuzi-Biéga National Park contained seeds and fruit remains of "Omphalocarpum mortehanii", junglesop ("Anonidium mannii"), "Antrocaryon nannanii", "Klainedoxa gabonensis", "Treculia africana", "Tetrapleura tetraptera", "Uapaca guineensis", "Autranella congolensis", "Gambeya africana" and "G. lacourtiana", "Mammea africana", "Cissus dinklagei", and "Grewia midlbrandii". Dung piles collected in a lowland rain forest in the northern Republic of Congo contained seeds of at least 96 plant species, with a minimum of 30 intact seeds and up to 1102 large seeds of more than in a single pile. Based on the analysis of 855 dung piles, it has been estimated that African forest elephants disperse a daily mean of 346 large seeds per of at least 73 tree species; they transport about a third of the large seeds for more than .