KNM-ER 1470 was much larger than the Olduvai remains, so the terms "H. habilis sensu lato" ("in the broad sense") and "H. habilis sensu stricto" ("in the strict sense") were used to include or exclude the larger morph, respectively. In 1986, English palaeoanthropologist Bernard Wood first suggested these remains represent a different "Homo" species, which coexisted with "H. habilis" and "H. ergaster"/"H. erectus". Coexisting "Homo" species conflicted with the predominant model of human evolution at the time which was that modern humans evolved in a straight line directly from "H. ergaster"/"H. erectus" which evolved directly from "H. habilis". In 1986, the remains were placed into a new species, "rudolfensis", by Russian anthropologist Valery Alekseyev (but he used the genus "Pithecanthropus", which was changed to "Homo" three years later by Groves). However, the validity of this species is debated, with some arguing that "H. habilis" was highly sexually dimorphic like modern non-human apes, with the larger skulls classified as ""H. rudolfensis"" actually representing male "H. habilis". In 1999, Wood and biological anthropologist Mark Collard recommended moving "rudolfensis" and "habilis" to "Australopithecus" based on the similarity of dental adaptations. However, they conceded that dental anatomy is highly variable among hominins and not always reliable when formulating family trees.