The origin of this diversity is debated; the break-up of Gondwana can explain vicariance older than 100 mya, but the diversity on the younger, smaller islands must have required a Cenozoic dispersal from the rims of the Indian Ocean to the islands. A "reverse colonisation", from islands to continents, apparently occurred more recently; the chameleons, for example, first diversified on Madagascar and then colonised Africa. Several species on the islands of the Indian Ocean are textbook cases of evolutionary processes; the dung beetles, day geckos, and lemurs are all examples of adaptive radiation.