Born in Pereslavl-Zalessky, Alexander was the second son of Prince Yaroslav Vsevolodovich and Feodosia Igorevna of Ryazan. His maternal grandfather was Igor Glebovich, the second son of Gleb Rostislavich, Prince of Ryazan (d. 1178). His maternal grandmother was Agrafena of Kiev, daughter of Rostislav I of Kiev. Alexander seemed to have no chance of claiming the throne of Vladimir. In 1236, however, the Novgorodians summoned him to become knyaz (or prince) of Novgorod and, as their military leader, to defend their northwest lands from Swedish and German invaders.