Holding a desirable territory increases the male's likelihood of finding a mate and therefore increases his reproductive success. However, each individual needs to weigh the benefits of mating with the costs of defending a territory. "Aglais io" exhibits this type of territorial behaviour, and must defend a desirable territory from other males. If only one of the males knows the territory well, he will successfully chase off any intruders. On the other hand, if both males are familiar with the territory, there will be a contest between the two to determine which of them stays in the territory. The most desirable sites are those that will increase the male's quota of females. These sites are generally feeding and oviposition sites, which are sought after by females. This territorial behaviour is reinforced by the fact that these sites are all concentrated. If the valuable resources were dispersed, there would be less observable territorial behaviour.