Later, there was a Roman villa at Preston Village, a Roman road from London ran nearby, and much physical evidence of Roman occupation has been discovered locally. From the 1st century AD, the Romans built a number of villas in Brighton and Romano-British Brythonic Celts formed farming settlements in the area. After the Romans left in the early 4th century AD, the Brighton area returned to the control of the native Celts. Anglo-Saxons then invaded in the late 5th century AD, and the region became part of the Kingdom of Sussex, founded in 477 AD by king Ælle.