The paternal lineages of the Fula/Fulɓe/Fulani tend to vary depending on geographic location. According to a study by Cruciani et al. (2002), around 90% of Fulani individuals from Burkina Faso carried haplotype 24, which corresponds with the E-M2 that is common in West Africa. The remainder belonged to haplotype 42/haplogroup E-M132. Both of these clades are today most frequent among Niger–Congo-speaking populations, particularly those inhabiting Senegal. Similarly, 53% of the Fulani in northern Cameroon bore haplogroup E-M132, with the rest mainly carrying other African clades (12% haplogroup A and 6% haplogroup E1b1a). A minority carried the West Eurasian haplogroups T (18%) and R1 (12%). Mulcare et al. (2004) observed a similar frequency of haplogroup R1 subclades in their Fulani samples from Cameroon (18%).