National Express received the first two low-floor Spectras in 1998, allocating one to Travel West Midlands and one to Travel Dundee, with registrations of R1 NEG & R2 NEG, their original fleet numbers being 1 & 2. They entered service on 15 January 1998, with branding reading Britain's first "Low Floor Low Emissions Easy Access Double Decker" and Scotland's first "Low Floor Low Emissions Easy Access Double Decker" A batch of 20 buses were purchased in 1999, and allocated to Travel West Midlands, to upgrade route 50 (Birmingham - Druids Heath) from Scania N113DRB operation. R2 NEG was also transferred to Travel West Midlands in August 1999 so all the National Express' Spectras were operated by Travel West Midlands, where 1 & 2 were re-numbered 4001 & 4002 to give them the same fleet numbers as the rest of the Spectras. They were replaced on the 50 in late 2001 by Volvo B7TL/Alexander ALX400s. They were transferred to Wolverhampton in the summer of 2009, to replace the final MCW Metrobuses allocated there, and were operated on various routes across the city, with the new NXWM livery and digital destination blinds. All examples in the fleet have now been withdrawn (September 2016) with 4014 (T414 UON) destroyed by fire in 2001. After 17 years of service, R1 NEG (4001) was donated to the Transport Museum, Wythall on 19 June 2015.