{HEAD ~ The nucleic acid} 1, having {TAIL ~ C terminal mutations} at positions 1010 and 1011 
{ HEAD ~ The nucleic acid } 1 , having { TAIL ~ C terminal mutations } at positions 1010 and 1011
HEAD HEAD HEAD HEAD HEAD HEAD HEAD OTH OTH REL TAIL TAIL TAIL TAIL TAIL TAIL TAIL OTH OTH OTH OTH OTH