In addition to {HEAD ~ prokaryotes}, eukaryotic microbes, such as {TAIL ~ yeast cultures}, may also be used 
In addition to { HEAD ~ prokaryotes } , eukaryotic microbes , such as { TAIL ~ yeast cultures } , may also be used
OTH OTH OTH HEAD HEAD HEAD HEAD HEAD OTH OTH OTH OTH OTH OTH TAIL TAIL TAIL TAIL TAIL TAIL OTH OTH OTH OTH OTH