Escherichia coli is an industrial workhorse organism with {HEAD ~ an unrivaled suite} of {TAIL ~ genetic tools} 
Escherichia coli is an industrial workhorse organism with { HEAD ~ an unrivaled suite } of { TAIL ~ genetic tools }
OTH OTH OTH OTH OTH OTH OTH OTH HEAD HEAD HEAD HEAD HEAD HEAD HEAD REL TAIL TAIL TAIL TAIL TAIL TAIL