a computer programmed for analyzing said images for determining {TAIL ~ the migration} of {HEAD ~ the cell} over time; and 
a computer programmed for analyzing said images for determining { TAIL ~ the migration } of { HEAD ~ the cell } over time ; and
OTH OTH OTH OTH OTH OTH OTH OTH OTH TAIL TAIL TAIL TAIL TAIL TAIL OTH HEAD HEAD HEAD HEAD HEAD HEAD OTH OTH OTH OTH