The development of more specialized methods due to the differences in cell density, water content and level of {TAIL ~ structural organization} of {HEAD ~ the full thickness LSE} was required 
The development of more specialized methods due to the differences in cell density , water content and level of { TAIL ~ structural organization } of { HEAD ~ the full thickness LSE } was required
OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH TAIL TAIL TAIL TAIL TAIL TAIL OTH HEAD HEAD HEAD HEAD HEAD HEAD HEAD HEAD OTH OTH