As a result, {HEAD ~ the determinable substance} in the cells is liberated from {TAIL ~ the cell} 
is liberated from
As a result , { HEAD ~ the determinable substance } in the cells is liberated from { TAIL ~ the cell }
OTH OTH OTH OTH HEAD HEAD HEAD HEAD HEAD HEAD HEAD OTH OTH OTH REL REL REL TAIL TAIL TAIL TAIL TAIL TAIL