In another embodiment the mutagen is {TAIL ~ γ irradation}, {HEAD ~ ultraviolet irradiation}, or X irradiation 
In another embodiment the mutagen is { TAIL ~ γ irradation } , { HEAD ~ ultraviolet irradiation } , or X irradiation
OTH OTH OTH OTH OTH OTH TAIL TAIL TAIL TAIL TAIL TAIL OTH HEAD HEAD HEAD HEAD HEAD HEAD OTH OTH OTH OTH