When the pH rises above {HEAD ~ the pI} of {TAIL ~ the protein}, the protein assumes a net negative charge 
of
When the pH rises above { HEAD ~ the pI } of { TAIL ~ the protein } , the protein assumes a net negative charge
OTH OTH OTH OTH OTH HEAD HEAD HEAD HEAD HEAD HEAD REL TAIL TAIL TAIL TAIL TAIL TAIL OTH OTH OTH OTH OTH OTH OTH OTH