Similarly, when {HEAD ~ the pH} of {TAIL ~ the environment} falls below the pH of the protein, the protein has a net positive charge 
of
Similarly , when { HEAD ~ the pH } of { TAIL ~ the environment } falls below the pH of the protein , the protein has a net positive charge
OTH OTH OTH HEAD HEAD HEAD HEAD HEAD HEAD REL TAIL TAIL TAIL TAIL TAIL TAIL OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH