However, {TAIL ~ sham operated animals} showed {HEAD ~ little increase} in the expression of RHAMM  
However , { TAIL ~ sham operated animals } showed { HEAD ~ little increase } in the expression of RHAMM
OTH OTH TAIL TAIL TAIL TAIL TAIL TAIL TAIL OTH HEAD HEAD HEAD HEAD HEAD HEAD OTH OTH OTH OTH OTH OTH