In some embodiments, {HEAD ~ the substrate} includes a soil substrate, {TAIL ~ a topsoil substrate}, a subsoil substrate, a clay substrate, a sand substrate, a rock substrate, or a stone substrate 
includes
In some embodiments , { HEAD ~ the substrate } includes a soil substrate , { TAIL ~ a topsoil substrate } , a subsoil substrate , a clay substrate , a sand substrate , a rock substrate , or a stone substrate
OTH OTH OTH OTH HEAD HEAD HEAD HEAD HEAD HEAD REL OTH OTH OTH OTH TAIL TAIL TAIL TAIL TAIL TAIL TAIL OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH