{HEAD ~ The process} 3, wherein {TAIL ~ the mammalian cell} is nonneuronal in origin 
wherein
{ HEAD ~ The process } 3 , wherein { TAIL ~ the mammalian cell } is nonneuronal in origin
HEAD HEAD HEAD HEAD HEAD HEAD OTH OTH REL TAIL TAIL TAIL TAIL TAIL TAIL TAIL OTH OTH OTH OTH