{HEAD ~ Map comparisons} would involve hybridizations, RFLP, {TAIL ~ PCR}, SSR and sequencing, all of which are conventional techniques 
involve
{ HEAD ~ Map comparisons } would involve hybridizations , RFLP , { TAIL ~ PCR } , SSR and sequencing , all of which are conventional techniques
HEAD HEAD HEAD HEAD HEAD HEAD OTH REL OTH OTH OTH OTH TAIL TAIL TAIL TAIL TAIL OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH