The starch, {HEAD ~ a fungus nutrient}, can be replaced by {TAIL ~ PVA}, a formulation additive that does not serve as a fungus nutrient 
replaced by
The starch , { HEAD ~ a fungus nutrient } , can be replaced by { TAIL ~ PVA } , a formulation additive that does not serve as a fungus nutrient
OTH OTH OTH HEAD HEAD HEAD HEAD HEAD HEAD HEAD OTH OTH OTH REL REL TAIL TAIL TAIL TAIL TAIL OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH