In {TAIL ~ some embodiments}, {HEAD ~ the substrate} includes a soil substrate, a topsoil substrate, a subsoil substrate, a clay substrate, a sand substrate, a rock substrate, or a stone substrate 
In
In { TAIL ~ some embodiments } , { HEAD ~ the substrate } includes a soil substrate , a topsoil substrate , a subsoil substrate , a clay substrate , a sand substrate , a rock substrate , or a stone substrate
REL TAIL TAIL TAIL TAIL TAIL TAIL OTH HEAD HEAD HEAD HEAD HEAD HEAD OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH