First, algal spores and a source of {TAIL ~ water} for {HEAD ~ treatment} are provided 
First , algal spores and a source of { TAIL ~ water } for { HEAD ~ treatment } are provided
OTH OTH OTH OTH OTH OTH OTH OTH TAIL TAIL TAIL TAIL TAIL OTH HEAD HEAD HEAD HEAD HEAD OTH OTH