Nevertheless, in certain cases, {HEAD ~ iron exhibits} stronger antipathogenic characteristics, particularly antimicrobial characteristics, compared to {TAIL ~ manganese} 
compared to
Nevertheless , in certain cases , { HEAD ~ iron exhibits } stronger antipathogenic characteristics , particularly antimicrobial characteristics , compared to { TAIL ~ manganese }
OTH OTH OTH OTH OTH OTH HEAD HEAD HEAD HEAD HEAD HEAD OTH OTH OTH OTH OTH OTH OTH OTH REL REL TAIL TAIL TAIL TAIL TAIL