In {HEAD ~ addition} to {TAIL ~ prokaryotes}, eukaryotic microbes, such as yeast cultures, may also be used 
to
In { HEAD ~ addition } to { TAIL ~ prokaryotes } , eukaryotic microbes , such as yeast cultures , may also be used
OTH HEAD HEAD HEAD HEAD HEAD REL TAIL TAIL TAIL TAIL TAIL OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH