The links include {HEAD ~ a middle connecting portion} that extends laterally between {TAIL ~ the first and second opposing ends} 
extends laterally between
The links include { HEAD ~ a middle connecting portion } that extends laterally between { TAIL ~ the first and second opposing ends }
OTH OTH OTH HEAD HEAD HEAD HEAD HEAD HEAD HEAD HEAD OTH REL REL REL TAIL TAIL TAIL TAIL TAIL TAIL TAIL TAIL TAIL TAIL