{TAIL ~ The epoxides} then spontaneously hydrolyze to {HEAD ~ intermediates} that can be further metabolized 
{ TAIL ~ The epoxides } then spontaneously hydrolyze to { HEAD ~ intermediates } that can be further metabolized
TAIL TAIL TAIL TAIL TAIL TAIL OTH OTH OTH OTH HEAD HEAD HEAD HEAD HEAD OTH OTH OTH OTH OTH