a computer programmed for analyzing {HEAD ~ said images} for determining {TAIL ~ the migration} of the cell over time; and 
for determining
a computer programmed for analyzing { HEAD ~ said images } for determining { TAIL ~ the migration } of the cell over time ; and
OTH OTH OTH OTH OTH HEAD HEAD HEAD HEAD HEAD HEAD REL REL TAIL TAIL TAIL TAIL TAIL TAIL OTH OTH OTH OTH OTH OTH OTH