Escherichia coli is {HEAD ~ an industrial workhorse organism} with an unrivaled suite of {TAIL ~ genetic tools} 
Escherichia coli is { HEAD ~ an industrial workhorse organism } with an unrivaled suite of { TAIL ~ genetic tools }
OTH OTH OTH HEAD HEAD HEAD HEAD HEAD HEAD HEAD HEAD OTH OTH OTH OTH OTH TAIL TAIL TAIL TAIL TAIL TAIL