In addition to {HEAD ~ prokaryotes}, {TAIL ~ eukaryotic microbes}, such as yeast cultures, may also be used 
In addition to { HEAD ~ prokaryotes } , { TAIL ~ eukaryotic microbes } , such as yeast cultures , may also be used
OTH OTH OTH HEAD HEAD HEAD HEAD HEAD OTH TAIL TAIL TAIL TAIL TAIL TAIL OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH