In {TAIL ~ addition} to prokaryotes, {HEAD ~ eukaryotic microbes}, such as yeast cultures, may also be used 
In
In { TAIL ~ addition } to prokaryotes , { HEAD ~ eukaryotic microbes } , such as yeast cultures , may also be used
REL TAIL TAIL TAIL TAIL TAIL OTH OTH OTH HEAD HEAD HEAD HEAD HEAD HEAD OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH