{HEAD ~ That reference} discloses that {TAIL ~ ornithine decarboxylase activity} increased precedent to polyamine biosynthesis 
discloses that
{ HEAD ~ That reference } discloses that { TAIL ~ ornithine decarboxylase activity } increased precedent to polyamine biosynthesis
HEAD HEAD HEAD HEAD HEAD HEAD REL REL TAIL TAIL TAIL TAIL TAIL TAIL TAIL OTH OTH OTH OTH OTH