Less fiber is deposited in those areas, resulting in {HEAD ~ the discrete areas} being comprised of {TAIL ~ a thinner mat} 
comprised of
Less fiber is deposited in those areas , resulting in { HEAD ~ the discrete areas } being comprised of { TAIL ~ a thinner mat }
OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH HEAD HEAD HEAD HEAD HEAD HEAD HEAD OTH REL REL TAIL TAIL TAIL TAIL TAIL TAIL TAIL