a computer programmed for analyzing said images for determining {HEAD ~ the migration} of {TAIL ~ the cell} over time; and 
of
a computer programmed for analyzing said images for determining { HEAD ~ the migration } of { TAIL ~ the cell } over time ; and
OTH OTH OTH OTH OTH OTH OTH OTH OTH HEAD HEAD HEAD HEAD HEAD HEAD REL TAIL TAIL TAIL TAIL TAIL TAIL OTH OTH OTH OTH