One screen may cooperate with {HEAD ~ another screen} to provide {TAIL ~ shearing} of the tissue specimens 
to provide
One screen may cooperate with { HEAD ~ another screen } to provide { TAIL ~ shearing } of the tissue specimens
OTH OTH OTH OTH OTH HEAD HEAD HEAD HEAD HEAD HEAD REL REL TAIL TAIL TAIL TAIL TAIL OTH OTH OTH OTH