Map comparisons would involve {TAIL ~ hybridizations}, RFLP, PCR, SSR and sequencing, all of which are {HEAD ~ conventional techniques} 
Map comparisons would involve { TAIL ~ hybridizations } , RFLP , PCR , SSR and sequencing , all of which are { HEAD ~ conventional techniques }
OTH OTH OTH OTH TAIL TAIL TAIL TAIL TAIL OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH OTH HEAD HEAD HEAD HEAD HEAD HEAD