Escherichia coli is {HEAD ~ an industrial workhorse organism} with {TAIL ~ an unrivaled suite} of genetic tools 
with
Escherichia coli is { HEAD ~ an industrial workhorse organism } with { TAIL ~ an unrivaled suite } of genetic tools
OTH OTH OTH HEAD HEAD HEAD HEAD HEAD HEAD HEAD HEAD REL TAIL TAIL TAIL TAIL TAIL TAIL TAIL OTH OTH OTH