As a result, {HEAD ~ the determinable substance} in {TAIL ~ the cells} is liberated from the cell 
As a result , { HEAD ~ the determinable substance } in { TAIL ~ the cells } is liberated from the cell
OTH OTH OTH OTH HEAD HEAD HEAD HEAD HEAD HEAD HEAD REL TAIL TAIL TAIL TAIL TAIL TAIL OTH OTH OTH OTH OTH