Honeycomb many feet, called travel stain by beekeepers when seen on frames of comb honey. Honeycomb in the "supers" that are not used for brood (e.g. by the placement of a queen excluder) stays light-colored. Numerous wasps, especially Polistinae and Vespinae, construct hexagonal prism-packed combs made of paper instead of wax; in some species (such as "Brachygastra mellifica"), honey is stored in the nest, thus technically forming a paper honeycomb. However, the term "honeycomb" is not often used for such structures. The axes of honeycomb cells are always quasihorizontal, and the nonangled rows of honeycomb cells are always horizontally (not vertically)
Honeycomb structure as far as we can see, is absolutely perfect in economizing labour and wax”. The first paper honeycomb structures might have been made by the Chinese 2000 years ago for ornaments, but no reference for this has been found. Paper honeycombs and the expansion production process has been invented in Halle/Saale in Germany by Hans Heilbrun in 1901 for decorative applications. First honeycomb structures from corrugated metal sheets had been proposed for bee keeping in 1890. For the same purpose, as foundation sheets to harvest more honey, a honeycomb moulding process using a paper paste glue mixture had been patented