Honeycomb many feet, called travel stain by beekeepers when seen on frames of comb honey. Honeycomb in the "supers" that are not used for brood (e.g. by the placement of a queen excluder) stays light-colored. Numerous wasps, especially Polistinae and Vespinae, construct hexagonal prism-packed combs made of paper instead of wax; in some species (such as "Brachygastra mellifica"), honey is stored in the nest, thus technically forming a paper honeycomb. However, the term "honeycomb" is not often used for such structures. The axes of honeycomb cells are always quasihorizontal, and the nonangled rows of honeycomb cells are always horizontally (not vertically)
Honeycomb also an example of geometric efficiency, albeit three-dimensional. The ends are trihedral (i.e., composed of three planes) sections of rhombic dodecahedra, with the dihedral angles of all adjacent surfaces measuring 120°, the angle that minimizes surface area for a given volume. (The angle formed by the edges at the pyramidal apex, known as the tetrahedral angle, is approximately 109° 28' 16" ().) The shape of the cells is such that two opposing honeycomb layers nest into each other, with each facet of the closed ends being shared by opposing cells. Individual cells do not show this geometric perfection: in a