Wave-formed ripple Wave-formed ripple In sedimentology, wave-formed ripples or wave-formed ripple marks are a feature of sediments (sandstones, limestones, siltstones) and dunes. These ripple marks are often characterised (and thus distinguished from current ripples) by symmetric cross sections and long relatively straight crests, which may commonly bifurcate. Commonly, these crests can be truncated by subsequent flows. Their wavelength (periodicity) depends on the sediment grain size, water depth and water-particle orbits in the waves. On tidal flats the pattern of wave-formed ripples may be complicated, as a product of changing depth and wind and tidal runoff directions. Symmetrical ripples are commonly found in
Genetic distance and Reynolds, Weir and Cockerham's genetic distance, listed below. In all the formulae in this section, formula_1 and formula_2 represent two different populations for which formula_3 loci have been studied. Let formula_4 represent the formula_5 allele at the formula_6 locus. In 1972, Masatoshi Nei published what came to be known as Nei's standard genetic distance. This distance has the nice property that if the rate of genetic change (amino acid substitution) is constant per year or generation then Nei's standard genetic distance ("D") increases in proportion to divergence time. This measure assumes that genetic differences are caused by mutation and