Matt Helders of the group, his vocals have been featured on many of the band's songs. He often sings in response to or in harmony with lead singer Alex Turner, and sings lead vocals on "D Is for Dangerous", "Brick by Brick", and "I.D.S.T.", as well as on parts of "You Probably Couldn't See for the Lights but You Were Staring Straight at Me", "Who the Fuck Are Arctic Monkeys?" and "Teddy Picker". Also, during two shows of the North American leg of the band's "Humbug" Tour, he sang an interpretation of Wham!'s "Last Christmas" during an encore performance of "Fluorescent Adolescent".
Matt Helders featured on Toddla T's album "Skanky Skanky". He remixed "Again & Again" for Roots Manuva, which gained airplay from Zane Lowe on BBC Radio 1. In August 2008, it was announced that Helders would compile the latest release in the LateNightTales DJ mix CD series. The set, entitled "", was released on 27 October 2008 and included a spoken work track performed and written by bandmate Alex Turner. Helders recorded the drum tracks on "Post Pop Depression", written by Iggy Pop and Josh Homme, which was released on 18 March 2016. Helders will tour with them to support the album.