Arkhangelsk Oblast Arkhangelsk region were Poonezhye () along the Onega, the Dvina Land along the Northern Dvina, Pinezhye () along the Pinega, Mezen Lands along the Mezen, and Pomorye () on the White Sea coast. The main waterway was the Northern Dvina, and Novgorod merchants used the Volga and its tributary, the Sheksna, along the Slavyanka River into Lake Nikolskoye, then the boats were taken by land to Lake Blagoveshchenskoye, from there downstream along the Porozovitsa River into Lake Kubenskoye and further to the Sukhona and the Northern Dvina. Portages from the Northern Dvina Basin led further to the Mezen and the
Onega stepanovi placed in phylum Proarticulata, as the segments in recently discovered, exceptionally well-preserved fossils display the glide, or "staggered", symmetry characteristic of the majority of proarticulatans. Imprints of "Onega stepanovi" have been found in the Verkhovka and Zimnie Gory Formations of the Ediacaran (Vendian) rocks of the Arkhangelsk Region, Russia. All the fossil specimens are negative imprints on the bases of fine-grained sandstone beds with the "elephant skin" and tubercle texture diagnostic of microbial mats. The same bedding planes contain various other Ediacaran species: "Cyclomedusa", "Ediacaria", palaeopascichnids, "Eoporpita", "Yorgia", "Andiva", "Archaeaspinus", "Vendia", "Dickinsonia", "Anfesta", "Albumares", "Tribrachidium", "Kimberella", "Parvancorina", "Charniodiscus" and others.