Primorsky District, Arkhangelsk Oblast Novgorod Republic. Novgorod merchants reached the White Sea in the 13th century; the Pomors, who inhabit the White Sea coast now, are the descendants of the original Novgorod population. After the fall of Novgorod, the area became a part of the Grand Duchy of Moscow. Until the 1700s, Arkhangelsk was the main trading harbor for the sea trade of Russia and Western Europe, and the Northern Dvina was the main trading route connecting central Russia with Arkhangelsk. The traditional occupation of the population of the Dvina Bay was fishery and trade. Peter the Great drastically changed the situation, by founding
Onega stepanovi placed in phylum Proarticulata, as the segments in recently discovered, exceptionally well-preserved fossils display the glide, or "staggered", symmetry characteristic of the majority of proarticulatans. Imprints of "Onega stepanovi" have been found in the Verkhovka and Zimnie Gory Formations of the Ediacaran (Vendian) rocks of the Arkhangelsk Region, Russia. All the fossil specimens are negative imprints on the bases of fine-grained sandstone beds with the "elephant skin" and tubercle texture diagnostic of microbial mats. The same bedding planes contain various other Ediacaran species: "Cyclomedusa", "Ediacaria", palaeopascichnids, "Eoporpita", "Yorgia", "Andiva", "Archaeaspinus", "Vendia", "Dickinsonia", "Anfesta", "Albumares", "Tribrachidium", "Kimberella", "Parvancorina", "Charniodiscus" and others.