Ethology also the discontinuance of undesirable behaviours. The term "ethology" derives from the Greek language: ἦθος, "ethos" meaning "character" and , "-logia" meaning "the study of". The term was first popularized by American myrmecologist (a person who studies ants) William Morton Wheeler in 1902. Because ethology is considered a topic of biology, ethologists have been concerned particularly with the evolution of behaviour and its understanding in terms of natural selection. In one sense, the first modern ethologist was Charles Darwin, whose 1872 book "The Expression of the Emotions in Man and Animals" influenced many ethologists. He pursued his interest in behaviour
The Expression of the Emotions in Man and Animals with "ill-temper", sulkiness and determination), Chapter 10 on hatred and anger, Chapter 11 on "disdain, contempt, disgust, guilt, pride, helplessness, patience and affirmation" and Chapter 12 on "surprise, astonishment, fear and horror". In his discussion of the emotion of disgust, Darwin notes its close links with the sense of smell, and conjectures an association with excretory products. In Chapter 13, Darwin discusses complex emotional states including self-attention, shame, shyness, modesty and blushing. Darwin describes blushing as "the most peculiar and most human of the expressions". Darwin closes the book with Chapter 14 in which he recapitulates his main argument: he