Willow ptarmigan complicated changes in plumage several times a year and the differing colour and pattern of the summer plumage: The willow ptarmigan often hybridises with the black grouse ("Tetrao tetrix") and the hazel grouse ("Tetrastes bonasia") and occasionally with the western capercaillie ("Tetrao urogallus"), the spruce grouse ("Falcipennis canadensis") and the rock ptarmigan ("Lagopus muta"). During the Pleistocene, the willow ptarmigan widely occurred in continental Europe. Authors who recognize paleosubspecies have named the Pleistocene willow ptarmigan "L. l. noaillensis" (though the older name "medius" might be the correct one). These marginally different birds are said to have gradually changed from the
Ptarmigan (sternwheeler) surface). Captain Armstrong was able to raise "Ptarmigan" and return her to service. However "Ptarmigan" caught fire later in 1907, and her upper works were destroyed. Again, "Ptarmigan" was salvaged, her upper works were rebuilt and she was returned to service. The gasoline engine was a new development in the early 1900s. Captain Northcote Cantlie, who has gone down in history as "an eccentric Scot" purchased a gasoline-engined launch, "Gian", and placed her on the upper Columbia. Unlike Armstrong, Cantlie came from a wealthy background, drank champagne for breakfast, and kept a bagpiper as his personal attendant. Cantlie, knowing that