Tertiary color if they exist at all, though indigo and scarlet are standard for blue–violet and red–vermilion. In another sense, a tertiary color is obtained by mixing secondary-colored pigments. These three colors are russet (orange–purple), slate (purple–green), and citron (green–orange), with the corresponding three quaternary colors plum (russet–slate), sage (slate–citron), buff (citron–russet) (with olive sometimes used for either slate or citron). Beyond that are shades of grey (blue grey and brown greys), which approach but never quite reach black. The RYB color terminology outlined above and in the color samples shown below is ultimately derived from the 1835 book "Chromatography", an analysis
Color vision fields that can compute local cone ratios. Such "double-opponent" cells were initially described in the goldfish retina by Nigel Daw; their existence in primates was suggested by David H. Hubel and Torsten Wiesel and subsequently proven by Bevil Conway. As Margaret Livingstone and David Hubel showed, double opponent cells are clustered within localized regions of V1 called blobs, and are thought to come in two flavors, red–green and blue–yellow. Red–green cells compare the relative amounts of red–green in one part of a scene with the amount of red–green in an adjacent part of the scene, responding best to local color