Willow ptarmigan complicated changes in plumage several times a year and the differing colour and pattern of the summer plumage: The willow ptarmigan often hybridises with the black grouse ("Tetrao tetrix") and the hazel grouse ("Tetrastes bonasia") and occasionally with the western capercaillie ("Tetrao urogallus"), the spruce grouse ("Falcipennis canadensis") and the rock ptarmigan ("Lagopus muta"). During the Pleistocene, the willow ptarmigan widely occurred in continental Europe. Authors who recognize paleosubspecies have named the Pleistocene willow ptarmigan "L. l. noaillensis" (though the older name "medius" might be the correct one). These marginally different birds are said to have gradually changed from the
Climate of the Arctic of heat from the subsurface land and ocean (both of which gain heat in summer and release it in winter) to the surface and atmosphere. Arctic days lengthen rapidly in March and April, and the sun rises higher in the sky, both bringing more solar radiation to the Arctic than in winter. During these early months of Northern Hemisphere spring most of the Arctic is still experiencing winter conditions, but with the addition of sunlight. The continued low temperatures, and the persisting white snow cover, mean that this additional energy reaching the Arctic from the sun is slow to have