Willow ptarmigan complicated changes in plumage several times a year and the differing colour and pattern of the summer plumage: The willow ptarmigan often hybridises with the black grouse ("Tetrao tetrix") and the hazel grouse ("Tetrastes bonasia") and occasionally with the western capercaillie ("Tetrao urogallus"), the spruce grouse ("Falcipennis canadensis") and the rock ptarmigan ("Lagopus muta"). During the Pleistocene, the willow ptarmigan widely occurred in continental Europe. Authors who recognize paleosubspecies have named the Pleistocene willow ptarmigan "L. l. noaillensis" (though the older name "medius" might be the correct one). These marginally different birds are said to have gradually changed from the
Arctic vegetation in the spring, and flower and set seed much sooner than plants that grow in warmer conditions. Their peak metabolic rate also occurs at a much lower temperature than plants from farther south. Compact cushions of vegetation keep the plants close to the warm soil and shield the tender central growing shoot. The height of Arctic plants is also governed by snow depth. Plants that protrude above the snow are subject to strong winds, blowing snow, and being eaten by caribou, muskox, or ptarmigan. Mosses and lichens are common in the Arctic. These plants have the ability to stop growth