Light-harvesting complexes of green plants Light-harvesting complexes of green plants The light-harvesting complex (or antenna complex) is an array of protein and chlorophyll molecules embedded in the thylakoid membrane of plants, which transfer light energy to one chlorophyll "a" molecule at the reaction center of a photosystem. The antenna pigments are predominantly chlorophyll "b", xanthophylls, and carotenes. Chlorophyll "a" is known as the core pigment. Their absorption spectra are non-overlapping and broaden the range of light that can be absorbed in photosynthesis. The carotenoids have another role as an antioxidant to prevent photo-oxidative damage of chlorophyll molecules. Each antenna complex has between 250 and 400
Chlorophyll directly absorbing light and performing charge separation events without the assistance of other chlorophyll pigments, but the probability of that happening under a given light intensity is small. Thus, the other chlorophylls in the photosystem and antenna pigment proteins all cooperatively absorb and funnel light energy to the reaction center. Besides chlorophyll "a", there are other pigments, called accessory pigments, which occur in these pigment–protein antenna complexes. Chlorophylls are numerous in types, but all are defined by the presence of a fifth ring beyond the four pyrrole-like rings. Most chlorophylls are classified as chlorins, which are reduced relatives to porphyrins