Coloration evidence for natural selection J. Bowler wrote that Darwin's theory In his 1889 book "Darwinism", the naturalist Alfred Russel Wallace considered the white coloration of Arctic animals. He recorded that the Arctic fox, Arctic hare, ermine and ptarmigan change their colour seasonally, and gave "the obvious explanation", that it was for concealment. The modern ornithologist W. L. N. Tickell, reviewing proposed explanations of white plumage in birds, writes that in the ptarmigan "it is difficult to escape the conclusion that cryptic brown summer plumage becomes a liability in snow, and white plumage is therefore another cryptic adaptation." All the same, he notes, "in spite
White-tailed ptarmigan to hatch. Males remain in the vicinity of the nest until the eggs hatch, a period of about 23 days. The precocial young leave the nest six to twelve hours after hatching. They are at first covered with down but become fully fledged in seven to ten days. Ptarmigan chicks begin their lives eating insects. Once the chicks' digestive tracts and ceca are more fully developed, their diets shift to one of flowers and leaves. Hens choose foraging patches where plant species containing proteins are abundant. The hens call their chicks to these plants, which are critical for the growth