Willow ptarmigan complicated changes in plumage several times a year and the differing colour and pattern of the summer plumage: The willow ptarmigan often hybridises with the black grouse ("Tetrao tetrix") and the hazel grouse ("Tetrastes bonasia") and occasionally with the western capercaillie ("Tetrao urogallus"), the spruce grouse ("Falcipennis canadensis") and the rock ptarmigan ("Lagopus muta"). During the Pleistocene, the willow ptarmigan widely occurred in continental Europe. Authors who recognize paleosubspecies have named the Pleistocene willow ptarmigan "L. l. noaillensis" (though the older name "medius" might be the correct one). These marginally different birds are said to have gradually changed from the
White-tailed ptarmigan to hatch. Males remain in the vicinity of the nest until the eggs hatch, a period of about 23 days. The precocial young leave the nest six to twelve hours after hatching. They are at first covered with down but become fully fledged in seven to ten days. Ptarmigan chicks begin their lives eating insects. Once the chicks' digestive tracts and ceca are more fully developed, their diets shift to one of flowers and leaves. Hens choose foraging patches where plant species containing proteins are abundant. The hens call their chicks to these plants, which are critical for the growth