Willow ptarmigan complicated changes in plumage several times a year and the differing colour and pattern of the summer plumage: The willow ptarmigan often hybridises with the black grouse ("Tetrao tetrix") and the hazel grouse ("Tetrastes bonasia") and occasionally with the western capercaillie ("Tetrao urogallus"), the spruce grouse ("Falcipennis canadensis") and the rock ptarmigan ("Lagopus muta"). During the Pleistocene, the willow ptarmigan widely occurred in continental Europe. Authors who recognize paleosubspecies have named the Pleistocene willow ptarmigan "L. l. noaillensis" (though the older name "medius" might be the correct one). These marginally different birds are said to have gradually changed from the
White-tailed ptarmigan season, while willow catkins, mountain avens flowers, and chickweed blooms, other flowers and leaves, lichens and berries form the majority of the ptarmigan's diet in the summer. Once fall and winter arrive in the region, the ptarmigan feeds on pine needles, seeds, willow and alder buds and twigs. Winter food sources have a much higher cellulose content than does summer forage, so the ptarmigan relies on bacteria-aided digestion in the cecum to extract essential nutrients. During the summer, the ptarmigan eats grit to assist in digesting plant material. White-tailed ptarmigan males are usually monogamous and remain with the same mate