Willow ptarmigan a range of lower plant species. It is also believed that the greening of parts of the Arctic is affecting Willow Ptarmigan populations by altering the shape and size of the shrubs they are able to feed on. Male willow ptarmigans are territorial birds. Males arrive in the breeding areas and set up territories in April and May, aggressively defending them against male interlopers. When the females arrive a few weeks later, the male performs courtship displays such as aerial manoeuvres, strutting and tail-fanning. When she has chosen a mate and a nesting site, the female lays a clutch of
Snow camouflage snow camouflage patterns. Charles Darwin mentioned the white winter coloration of the ptarmigan in his 1859 "Origin of Species": The white protective coloration of arctic animals was noted by an early student of camouflage, the naturalist Alfred Russel Wallace, in his 1889 book "Darwinism"; he listed the polar bear, the American polar hare, the snowy owl and the gyr falcon as remaining white all year, while the arctic fox, arctic hare, ermine and ptarmigan change their colour, and observed "the obvious explanation", that it was for concealment, at a time when Darwinism was at a low ebb. Later zoologists such