Willow ptarmigan complicated changes in plumage several times a year and the differing colour and pattern of the summer plumage: The willow ptarmigan often hybridises with the black grouse ("Tetrao tetrix") and the hazel grouse ("Tetrastes bonasia") and occasionally with the western capercaillie ("Tetrao urogallus"), the spruce grouse ("Falcipennis canadensis") and the rock ptarmigan ("Lagopus muta"). During the Pleistocene, the willow ptarmigan widely occurred in continental Europe. Authors who recognize paleosubspecies have named the Pleistocene willow ptarmigan "L. l. noaillensis" (though the older name "medius" might be the correct one). These marginally different birds are said to have gradually changed from the
Climate of the Arctic Climate of the Arctic The climate of the Arctic is characterized by long, cold winters and short, cool summers. There is a large amount of variability in climate across the Arctic, but all regions experience extremes of solar radiation in both summer and winter. Some parts of the Arctic are covered by ice (sea ice, glacial ice, or snow) year-round, and nearly all parts of the Arctic experience long periods with some form of ice on the surface. Average January temperatures range from about −34 °C to 0 °C (−29 to +32 °F), and winter temperatures can drop below over