White-tailed ptarmigan and development of the chicks. This suggests that the hens' food calls assist in enhancing survival rates of juvenile ptarmigans. The young birds remain with their mother throughout the summer and autumn season. The white-tailed ptarmigan is well-camouflaged when on the ground. In his pioneering 1909 book on the subject, "Concealing-Coloration in the Animal Kingdom", the American artist Abbott Thayer wrote: The white-tailed ptarmigan has feathers located on its feet to serve as protection from the extreme cold often experienced in the alpine tundra environment. In addition, these birds have feathers around their nostrils to warm the air prior to
Snow camouflage snow camouflage patterns. Charles Darwin mentioned the white winter coloration of the ptarmigan in his 1859 "Origin of Species": The white protective coloration of arctic animals was noted by an early student of camouflage, the naturalist Alfred Russel Wallace, in his 1889 book "Darwinism"; he listed the polar bear, the American polar hare, the snowy owl and the gyr falcon as remaining white all year, while the arctic fox, arctic hare, ermine and ptarmigan change their colour, and observed "the obvious explanation", that it was for concealment, at a time when Darwinism was at a low ebb. Later zoologists such