Willow ptarmigan complicated changes in plumage several times a year and the differing colour and pattern of the summer plumage: The willow ptarmigan often hybridises with the black grouse ("Tetrao tetrix") and the hazel grouse ("Tetrastes bonasia") and occasionally with the western capercaillie ("Tetrao urogallus"), the spruce grouse ("Falcipennis canadensis") and the rock ptarmigan ("Lagopus muta"). During the Pleistocene, the willow ptarmigan widely occurred in continental Europe. Authors who recognize paleosubspecies have named the Pleistocene willow ptarmigan "L. l. noaillensis" (though the older name "medius" might be the correct one). These marginally different birds are said to have gradually changed from the
Rock ptarmigan the Russian High Arctic overwinters during the polar night and survives by feeding on rich vegetation on and underneath high cliffs where seabird colonies are located in summer.<ref name="FJL/">Dinets, V. (2001) Winter ecology of Willow Ptarmigan "Lagopus lagopus" and Rock Ptarmigan "L. mutus" at the northern limit of their range. "Ornitologia" 29: 326-327 (in Russian with English summary).</ref> During the last ice age, the species was far more widespread in continental Europe. The rock ptarmigan feeds primarily on birch and willow buds and catkins when available. It also eats various seeds, leaves, flowers, and berries of other plant species. Insects