Coloration evidence for natural selection J. Bowler wrote that Darwin's theory In his 1889 book "Darwinism", the naturalist Alfred Russel Wallace considered the white coloration of Arctic animals. He recorded that the Arctic fox, Arctic hare, ermine and ptarmigan change their colour seasonally, and gave "the obvious explanation", that it was for concealment. The modern ornithologist W. L. N. Tickell, reviewing proposed explanations of white plumage in birds, writes that in the ptarmigan "it is difficult to escape the conclusion that cryptic brown summer plumage becomes a liability in snow, and white plumage is therefore another cryptic adaptation." All the same, he notes, "in spite
Climate of the Arctic noon, not much energy is reaching the surface. Furthermore, most of the small amount of solar radiation that reaches the surface is reflected away by the bright snow cover. Cold snow reflects between 70% and 90% of the solar radiation that reaches it, and snow covers most of the Arctic land and ice surface in winter. These factors result in a negligible input of solar energy to the Arctic in winter; the only things keeping the Arctic from continuously cooling all winter are the transport of warmer air and ocean water into the Arctic from the south and the transfer