Coloration evidence for natural selection J. Bowler wrote that Darwin's theory In his 1889 book "Darwinism", the naturalist Alfred Russel Wallace considered the white coloration of Arctic animals. He recorded that the Arctic fox, Arctic hare, ermine and ptarmigan change their colour seasonally, and gave "the obvious explanation", that it was for concealment. The modern ornithologist W. L. N. Tickell, reviewing proposed explanations of white plumage in birds, writes that in the ptarmigan "it is difficult to escape the conclusion that cryptic brown summer plumage becomes a liability in snow, and white plumage is therefore another cryptic adaptation." All the same, he notes, "in spite
Snow camouflage to escape the conclusion that cryptic brown summer plumage becomes a liability in snow, and white plumage is therefore another cryptic adaptation." All the same, he notes, "in spite of winter plumage, many Ptarmigan in NE Iceland are killed by Gyrfalcons throughout the winter." Some animals of the far north, like the showshoe and Arctic hares, Arctic fox, stoat, and rock ptarmigan change their coat colour (by moulting and growing new fur or feathers) from brown or grey summer camouflage to white in the winter; the Arctic fox is the only species in the dog family to do so. However,