Willow ptarmigan complicated changes in plumage several times a year and the differing colour and pattern of the summer plumage: The willow ptarmigan often hybridises with the black grouse ("Tetrao tetrix") and the hazel grouse ("Tetrastes bonasia") and occasionally with the western capercaillie ("Tetrao urogallus"), the spruce grouse ("Falcipennis canadensis") and the rock ptarmigan ("Lagopus muta"). During the Pleistocene, the willow ptarmigan widely occurred in continental Europe. Authors who recognize paleosubspecies have named the Pleistocene willow ptarmigan "L. l. noaillensis" (though the older name "medius" might be the correct one). These marginally different birds are said to have gradually changed from the
Climate of the Arctic the northern and southern land areas receive their winter snow cover, which combined with the reduced solar radiation at the surface, ensures an end to the warm days those areas may experience in summer. By November, winter is in full swing in most of the Arctic, and the small amount of solar radiation still reaching the region does not play a significant role in its climate. The Arctic is often perceived as a region stuck in a permanent deep freeze. While much of the region does experience very low temperatures, there is considerable variability with both location and season. Winter