Willow ptarmigan complicated changes in plumage several times a year and the differing colour and pattern of the summer plumage: The willow ptarmigan often hybridises with the black grouse ("Tetrao tetrix") and the hazel grouse ("Tetrastes bonasia") and occasionally with the western capercaillie ("Tetrao urogallus"), the spruce grouse ("Falcipennis canadensis") and the rock ptarmigan ("Lagopus muta"). During the Pleistocene, the willow ptarmigan widely occurred in continental Europe. Authors who recognize paleosubspecies have named the Pleistocene willow ptarmigan "L. l. noaillensis" (though the older name "medius" might be the correct one). These marginally different birds are said to have gradually changed from the
Climate of the Arctic noon, not much energy is reaching the surface. Furthermore, most of the small amount of solar radiation that reaches the surface is reflected away by the bright snow cover. Cold snow reflects between 70% and 90% of the solar radiation that reaches it, and snow covers most of the Arctic land and ice surface in winter. These factors result in a negligible input of solar energy to the Arctic in winter; the only things keeping the Arctic from continuously cooling all winter are the transport of warmer air and ocean water into the Arctic from the south and the transfer