Willow ptarmigan complicated changes in plumage several times a year and the differing colour and pattern of the summer plumage: The willow ptarmigan often hybridises with the black grouse ("Tetrao tetrix") and the hazel grouse ("Tetrastes bonasia") and occasionally with the western capercaillie ("Tetrao urogallus"), the spruce grouse ("Falcipennis canadensis") and the rock ptarmigan ("Lagopus muta"). During the Pleistocene, the willow ptarmigan widely occurred in continental Europe. Authors who recognize paleosubspecies have named the Pleistocene willow ptarmigan "L. l. noaillensis" (though the older name "medius" might be the correct one). These marginally different birds are said to have gradually changed from the
Climate of the Arctic twice as much as the global mean temperature has. In 2009, NASA reported that 45 percent or more of the observed warming in the Arctic since 1976 was likely a result of changes in tiny airborne particles called aerosols. Climate models predict that the temperature increase in the Arctic over the next century will continue to be about twice the global average temperature increase. By the end of the 21st century, the annual average temperature in the Arctic is predicted to increase by , with more warming in winter ( than in summer. Decreases in sea-ice extent and thickness are