Accessory pigment other accessory pigments is sometimes referred to as a "pigment bed". The different chlorophyll and non-chlorophyll pigments associated with the photosystems all have different absorption spectra, either because the spectra of the different chlorophyll pigments are modified by their local protein environment or because the accessory pigments have intrinsic structural differences. The result is that, in vivo, a composite absorption spectrum of all these pigments is broadened and flattened such that a wider range of visible and infrared radiation is absorbed by plants and algae. Most photosynthetic organisms do not absorb green light well, thus most remaining light under leaf
Light-harvesting complexes of green plants protons and electrons. Under changing light conditions, the reversible phosphorylation of light harvesting chlorophyll a/b binding proteins (LHCII) represents a system for balancing the excitation energy between the two photosystems. The N-terminus of the chlorophyll A-B binding protein extends into the stroma where it is involved with adhesion of granal membranes and photo-regulated by reversible phosphorylation of its threonine residues. Both these processes are believed to mediate the distribution of excitation energy between photosystems I and II. This family also includes the photosystem II protein PsbS, which plays a role in energy-dependent quenching that increases thermal dissipation of excess absorbed