Coloration evidence for natural selection J. Bowler wrote that Darwin's theory In his 1889 book "Darwinism", the naturalist Alfred Russel Wallace considered the white coloration of Arctic animals. He recorded that the Arctic fox, Arctic hare, ermine and ptarmigan change their colour seasonally, and gave "the obvious explanation", that it was for concealment. The modern ornithologist W. L. N. Tickell, reviewing proposed explanations of white plumage in birds, writes that in the ptarmigan "it is difficult to escape the conclusion that cryptic brown summer plumage becomes a liability in snow, and white plumage is therefore another cryptic adaptation." All the same, he notes, "in spite
Rock ptarmigan ptarmigan by about 10%. The male's "song" is a loud croaking. The rock ptarmigan is seasonally camouflaged; its feathers moult from white in winter to brown in spring or summer. The breeding male has greyish upper parts with white wings and under parts. In winter, its plumage becomes completely white except for the black tail. It can be distinguished from the winter willow ptarmigan by habitat—the rock ptarmigan prefers higher elevations and more barren habitat; it is also smaller with a more delicate bill. The rock ptarmigan is a sedentary species which breeds across arctic and subarctic Eurasia and North