Willow ptarmigan complicated changes in plumage several times a year and the differing colour and pattern of the summer plumage: The willow ptarmigan often hybridises with the black grouse ("Tetrao tetrix") and the hazel grouse ("Tetrastes bonasia") and occasionally with the western capercaillie ("Tetrao urogallus"), the spruce grouse ("Falcipennis canadensis") and the rock ptarmigan ("Lagopus muta"). During the Pleistocene, the willow ptarmigan widely occurred in continental Europe. Authors who recognize paleosubspecies have named the Pleistocene willow ptarmigan "L. l. noaillensis" (though the older name "medius" might be the correct one). These marginally different birds are said to have gradually changed from the
Arctic Cordillera moss campion, bilberry, and Arctic white heather. The conditions here are far too severe for reptiles and amphibians to survive, and insects are also rare in the region. Muskoxen and barren-ground caribou are the only large herbivores in this environment, while polar bears and the Arctic wolf are the only large carnivores to be found in the region. Smaller herbivores include the Arctic hare and the collared lemming. Arctic foxes and stoats are some of the smaller carnivores found in the region. Marine mammals include narwhals, beluga whales, walrus, and ringed and bearded seals. The furry-legged rock ptarmigan is a