Equus (genus) 54 million years ago (Mya). These animals had three toes on the hind feet and four on the front feet with small hooves in place of claws, but also had soft pads.<ref name="doi10.1126/science.1105458"></ref> Equids developed into larger, three-toed animals (e.g. "Mesohippus") during the Oligocene and Miocene. From there, the side toes became progressively smaller through the Pleistocene until the emergence of the single-toed "Equus". The genus "Equus", which includes all extant equines, is believed to have evolved from "Dinohippus", via the intermediate form "Plesippus". One of the oldest species is "Equus simplicidens", described as zebra-like with a donkey-like head shape.
Equus lambei of the modern wild horse, including the domestic horse, "Equus caballus". Controversy still surrounds "E. lambei" and the divergence of other similar extinct horse species. Through examinations of the bones and teeth of "Equus lambei", many similarities can be seen with the modern horse "Equus caballus". There is also a strong resemblance to "E. lambei" in the metapodials of "E. caballus przewalskii" and the phalanges of "E. caballus lenensis" "." Metrical and morphological studies of horse teeth from the Bluefish Caves confirm the close similarity between "Equus lambei" with wild and domestic horses alive today. "E. lambei" is a "caballus"