Alosa to detect ultrasound (at frequencies above 20 kHz, which is the limit of human hearing). This was first discovered by fisheries biologists studying a type of shad known as blueback herring, and was later verified in laboratory studies of hearing in American shad. This ability is thought to help them avoid dolphins that find prey using echolocation. "Alosa" species are generally pelagic. They are mostly anadromous or semianadromous with the exception of strictly freshwater landlocked species. "Alosa" species are generally migratory and schooling fish. Males usually mature about a year before females; they spawn in the late spring to summer.
Alosa lifespan of "Alosa" species can be up to 10 years, but this is generally uncommon, as many die after spawning. The systematics and distribution of "Alosa" shads are complex. The genus inhabits a wide range of habitats, and many taxa are migratory. A few forms are landlocked, including one from Killarney in Ireland, two from lakes in northern Italy, and two in Greece. Several species are native to the Black Sea and Caspian Sea. "Alosa" species of the Caspian are systemically characterized by the number of rakers on the first gill arch. They are classified as being "multirakered", "medium-rakered", or