Specimens of Archaeopteryx Specimens of Archaeopteryx Archaeopteryx fossils from the quarries of Solnhofen limestone represent the most famous and well-known fossils from this area. They are highly significant to paleontology and avian evolution in that they document the fossil record's oldest-known birds. Over the years, twelve body fossil specimens of "Archaeopteryx" and a feather that may belong to it have been found, though one of these specimens was reassigned to another genus by two researchers in 2017. All of the fossils come from the upper Jurassic lithographic limestone deposits, quarried for centuries, near Solnhofen, Germany. The initial discovery, a single feather, was unearthed
Archaeopteryx be related particularly closely to "Anchiornis" and "Xiaotingia", which were recovered as basal troodontids instead. Agnolín and Novas (2013) found "Archaeopteryx" and (possibly synonymous) "Wellnhoferia" to be the basalmost avialans (Avialae being defined by the authors as including "Archaeopteryx lithographica" and "Passer", their most recent common ancestor and all of its descendants), with Microraptoria, Unenlagiinae, and the clade containing "Anchiornis" and "Xiaotingia" being successively closer outgroups to the Avialae. Another phylogenetic study by Godefroit, "et al.", using a more inclusive matrix than the one from the analysis in the description of "Eosinopteryx brevipenna", also found "Archaeopteryx" to be a member