Feather a dromaeosaurid found in the Yixian formation (124.6 MYA). Previously, a temporal paradox existed in the evolution of feathers—theropods with highly derived bird-like characteristics occurred at a later time than "Archaeopteryx"—suggesting that the descendants of birds arose before the ancestor. However, the discovery of "Anchiornis huxleyi" in the Late Jurassic Tiaojishan Formation (160 MYA) in western Liaoning in 2009 resolved this paradox. By predating "Archaeopteryx", "Anchiornis" proves the existence of a modernly feathered theropod ancestor, providing insight into the dinosaur-bird transition. The specimen shows distribution of large pennaceous feathers on the forelimbs and tail, implying that pennaceous feathers spread to
Archaeopteryx common uses of that term. A more thorough analysis was published soon after to test this hypothesis, and failed to arrive at the same result; it found "Archaeopteryx" in its traditional position at the base of Avialae, while "Xiaotingia" was recovered as a basal dromaeosaurid or troodontid. The authors of the follow-up study noted that uncertainties still exist, and that it may not be possible to state confidently whether or not "Archaeopteryx" is a member of Avialae or not, barring new and better specimens of relevant species. Phylogenetic studies conducted by Senter, "et al." (2012) and Turner, Makovicky, and Norell