Specimens of Archaeopteryx feather's earlier discovery, von Meyer made the London fossil the type specimen of the genus ("Archaeopteryx") and species ("lithographica"), establishing it as the exemplar of the species' features. Its discovery and subsequent descriptions, a mere handful of years after Darwin's "On the Origin of Species", fulfilled the hope of a truly transitional form in its then-unique amalgamation of bird and reptilian characters. The specimen is preserved on two counter slabs broken originally into four pieces. The main, overlaying slab contains most of the skeletal elements, while only impressions and a few bone fragments are visible on the counter slab. This
Archaeopteryx common uses of that term. A more thorough analysis was published soon after to test this hypothesis, and failed to arrive at the same result; it found "Archaeopteryx" in its traditional position at the base of Avialae, while "Xiaotingia" was recovered as a basal dromaeosaurid or troodontid. The authors of the follow-up study noted that uncertainties still exist, and that it may not be possible to state confidently whether or not "Archaeopteryx" is a member of Avialae or not, barring new and better specimens of relevant species. Phylogenetic studies conducted by Senter, "et al." (2012) and Turner, Makovicky, and Norell