Archaeopteryx or "Allosaurus", which had a crocodile-like anatomy of the brain and inner ear. Newer research shows that while the "Archaeopteryx" brain was more complex than that of more primitive theropods, it had a more generalized brain volume among Maniraptora dinosaurs, even smaller than that of other non-avian dinosaurs in several instances, which indicates the neurological development required for flight was already a common trait in the maniraptoran clade. Recent studies of flight feather barb geometry reveal that modern birds possess a larger barb angle in the trailing vane of the feather, whereas "Archaeopteryx" lacks this large barb angle, indicating potentially
Specimens of Archaeopteryx skeleton of a feathered animal from the same lithographic shale deposits, which would later be known as the London specimen. Coincidentally, von Meyer proposed the name "Archaeopteryx lithographica" for the feather, but not for the skeleton. Therefore, the official name of the animal was originally linked to the single feather rather than any actual skeleton, and is formally considered the original holotype. Though 1860 is often the year named for the feather's discovery, there is no proof of this date, and some authors consider it more likely to have been found in 1861, as it seems reasonable that von Meyer's