Protoavis are well preserved, and they are astonishingly non-avian. The distal carpals, while long, are in no way similar to those observed in the urvogel or other archaic birds. There is no semilunate element, and the structure of the radiale and ulnare would have limited the flexibility of the wrist joint. The manus is not tridactyl, and metacarpal V is present. In even the most basal avialian, "Archaeopteryx", there is no vestige of the fifth metacarpal and its presence in "Protoavis" seems incongruous with the claim that it is a bird, let alone one more derived than "Archaeopteryx". Chatterjee claims that
Specimens of Archaeopteryx that "Archaeopteryx" could have raised its short second toe in a similar manner to its dromaeosaur cousin "Deinonychus", whose second toe bore an enlarged "sickle claw" for probable use in predation. It remains unclear whether "Archaeopteryx" also used its hyperextended toe-claw in a similar fashion or for some other purpose. Taken together, many of the unique features of the skull, feet and coracoid strengthen a close relationship of archaeopterygids to deinonychosaurian theropods. Well-preserved imprints of wing and tail plumage show details of the barbs, but not barbules, and the shafts of 11 primaries can be counted on the right wing.