Alosa macedonica Alosa macedonica Alosa macedonica, or the Macedonian shad (also known as "liparia"), is a landlocked species of clupeid fish endemic to Greece. Its single natural occurrence is the freshwater Lake Volvi in northern Greece. It is threatened by habitat loss. "Alosa macedonica" is a member of the "Alosa" genus, whose other species are often anadromous migrating between marine and freshwater. Research suggests that the ancestors of "Alosa macedonica" inhabited marine regions of the Aegean Sea. "Alosa macedonica" have teeth in the palatine and vomer. They have approximately 50 vertebrae and 106-128 gill rakers. They are about 181-230mm in length and
Alosa lifespan of "Alosa" species can be up to 10 years, but this is generally uncommon, as many die after spawning. The systematics and distribution of "Alosa" shads are complex. The genus inhabits a wide range of habitats, and many taxa are migratory. A few forms are landlocked, including one from Killarney in Ireland, two from lakes in northern Italy, and two in Greece. Several species are native to the Black Sea and Caspian Sea. "Alosa" species of the Caspian are systemically characterized by the number of rakers on the first gill arch. They are classified as being "multirakered", "medium-rakered", or