Feathered dinosaur is possible that feathers first developed in an even earlier group, in light of the discovery of pycnofibers of pterosaurs. Crocodilians also possess beta keratin similar to those of birds, which suggests that they evolved from common ancestral genes. Shortly after the 1859 publication of Charles Darwin's "On the Origin of Species", British biologist Thomas Henry Huxley proposed that birds were descendants of dinosaurs. He compared the skeletal structure of "Compsognathus", a small theropod dinosaur, and the 'first bird' "Archaeopteryx lithographica" (both of which were found in the Upper Jurassic Bavarian limestone of Solnhofen). He showed that, apart from its
Specimens of Archaeopteryx its aerodynamic function as a flight feather. As the feather bears little resemblance to the retrices of the other full "Archaeopteryx" skeletons, it is generally thought to be a wing feather: Griffiths and others have concluded it was a remex, while Carney in 2012 interpreted it to be a covert feather. The feather has a relatively low degree of asymmetry, which Speakman and Thompson in 1994 concluded to indicate it as a secondary remex. If this is the case, then it would have originated from an animal smaller than even the Eichstätt specimen, which is the smallest "Archaeopteryx" specimen known