Feathered dinosaur is possible that feathers first developed in an even earlier group, in light of the discovery of pycnofibers of pterosaurs. Crocodilians also possess beta keratin similar to those of birds, which suggests that they evolved from common ancestral genes. Shortly after the 1859 publication of Charles Darwin's "On the Origin of Species", British biologist Thomas Henry Huxley proposed that birds were descendants of dinosaurs. He compared the skeletal structure of "Compsognathus", a small theropod dinosaur, and the 'first bird' "Archaeopteryx lithographica" (both of which were found in the Upper Jurassic Bavarian limestone of Solnhofen). He showed that, apart from its
Specimens of Archaeopteryx other "Archaeopteryx" specimens. The left wing preserves the better plumage and shows flight feathers firmly connected to the arms. Though the wings overlap each other enough that assigning feather attachment points to individual bone regions is not possible, it does appear, consistent with other specimens and modern relatives, that the long primary feathers are attached to the second manual digit. The feathers of the right wing are considerably more damaged; so many feathers are missing that only a few can be definitively linked to the ulna, designating them as secondary remiges. One small feather is isolated next to the right