Archaeopteryx following Thomas Henry Huxley's lead in 1868, argued that birds evolved within theropod dinosaurs and "Archaeopteryx" was a critical piece of evidence for this argument; it had several avian features, such as a wishbone, flight feathers, wings, and a partially reversed first toe along with dinosaur and theropod features. For instance, it has a long ascending process of the ankle bone, interdental plates, an obturator process of the ischium, and long chevrons in the tail. In particular, Ostrom found that "Archaeopteryx" was remarkably similar to the theropod family Dromaeosauridae. Specimens of "Archaeopteryx" were most notable for their well-developed flight feathers.
Archaeopteryx be related particularly closely to "Anchiornis" and "Xiaotingia", which were recovered as basal troodontids instead. Agnolín and Novas (2013) found "Archaeopteryx" and (possibly synonymous) "Wellnhoferia" to be the basalmost avialans (Avialae being defined by the authors as including "Archaeopteryx lithographica" and "Passer", their most recent common ancestor and all of its descendants), with Microraptoria, Unenlagiinae, and the clade containing "Anchiornis" and "Xiaotingia" being successively closer outgroups to the Avialae. Another phylogenetic study by Godefroit, "et al.", using a more inclusive matrix than the one from the analysis in the description of "Eosinopteryx brevipenna", also found "Archaeopteryx" to be a member