Archaeopteryx perhaps, to a cartilaginous sternum. The sideways orientation of the glenoid (shoulder) joint between scapula, coracoid, and humerus—instead of the dorsally angled arrangement found in modern birds—may indicate that "Archaeopteryx" was unable to lift its wings above its back, a requirement for the upstroke found in modern flapping flight. According to a study by Philip Senter in 2006, "Archaeopteryx" was indeed unable to use flapping flight as modern birds do, but it may well have used a downstroke-only flap-assisted gliding technique. "Archaeopteryx" wings were relatively large, which would have resulted in a low stall speed and reduced turning radius. The
Archaeopteryx be related particularly closely to "Anchiornis" and "Xiaotingia", which were recovered as basal troodontids instead. Agnolín and Novas (2013) found "Archaeopteryx" and (possibly synonymous) "Wellnhoferia" to be the basalmost avialans (Avialae being defined by the authors as including "Archaeopteryx lithographica" and "Passer", their most recent common ancestor and all of its descendants), with Microraptoria, Unenlagiinae, and the clade containing "Anchiornis" and "Xiaotingia" being successively closer outgroups to the Avialae. Another phylogenetic study by Godefroit, "et al.", using a more inclusive matrix than the one from the analysis in the description of "Eosinopteryx brevipenna", also found "Archaeopteryx" to be a member