Equus (genus) 54 million years ago (Mya). These animals had three toes on the hind feet and four on the front feet with small hooves in place of claws, but also had soft pads.<ref name="doi10.1126/science.1105458"></ref> Equids developed into larger, three-toed animals (e.g. "Mesohippus") during the Oligocene and Miocene. From there, the side toes became progressively smaller through the Pleistocene until the emergence of the single-toed "Equus". The genus "Equus", which includes all extant equines, is believed to have evolved from "Dinohippus", via the intermediate form "Plesippus". One of the oldest species is "Equus simplicidens", described as zebra-like with a donkey-like head shape.
Onager known as the Persian onager. Until this day, the species share the same name, "onager". The onager is a member of the subgenus "Asinus", belonging to the genus "Equus" and is classified under the family Equidae. The species was described and given its binomial name "Equus hemionus" by German zoologist Peter Simon Pallas in 1775. The Asiatic wild ass, among Old World equids, existed for more than 4 million years. The oldest divergence of "Equus" was the onager followed by the zebras and onwards. A new species called the kiang ("E. kiang"), a Tibetan relative, was previously considered to be