Archaeopteryx or "Allosaurus", which had a crocodile-like anatomy of the brain and inner ear. Newer research shows that while the "Archaeopteryx" brain was more complex than that of more primitive theropods, it had a more generalized brain volume among Maniraptora dinosaurs, even smaller than that of other non-avian dinosaurs in several instances, which indicates the neurological development required for flight was already a common trait in the maniraptoran clade. Recent studies of flight feather barb geometry reveal that modern birds possess a larger barb angle in the trailing vane of the feather, whereas "Archaeopteryx" lacks this large barb angle, indicating potentially
Specimens of Archaeopteryx other "Archaeopteryx" specimens. The left wing preserves the better plumage and shows flight feathers firmly connected to the arms. Though the wings overlap each other enough that assigning feather attachment points to individual bone regions is not possible, it does appear, consistent with other specimens and modern relatives, that the long primary feathers are attached to the second manual digit. The feathers of the right wing are considerably more damaged; so many feathers are missing that only a few can be definitively linked to the ulna, designating them as secondary remiges. One small feather is isolated next to the right