Specimens of Archaeopteryx length. These hindlimb proportions agree roughly with those of the London specimen. The claw arc of the central pedal digit is about 120 degrees, which is close to the average for all "Archaeopteryx" specimens. This arc is also close to the average for that in perching birds and is well above the mean for ground-dwelling birds, despite conclusions against perching habits based on the short hallux and weak flexor tubercles. The remarkably preserved plumage is one of many features that sets the Berlin specimen head-and-coracoids above all other "Archaeopteryx" fossils to date. Clear feather impressions are visible on both wings,
Specimens of Archaeopteryx suggest that it had some black colouration which included the coverts (or possibly the secondary remiges). Carney pointed out that this is consistent with what we know of modern flight characteristics, in that black melanosomes have structural properties that strengthen feathers for flight. The preservation of the fossil feather is unusual. In contrast to the feather imprints found on the full skeletons of "Archaeopteryx", the isolated feather is preserved as a dark film, which could be composed either of organic matter (as in most fossil feathers) or as inorganic minerals. Though it is theoretically possible to determine which it is,