Archaeopteryx perhaps, to a cartilaginous sternum. The sideways orientation of the glenoid (shoulder) joint between scapula, coracoid, and humerus—instead of the dorsally angled arrangement found in modern birds—may indicate that "Archaeopteryx" was unable to lift its wings above its back, a requirement for the upstroke found in modern flapping flight. According to a study by Philip Senter in 2006, "Archaeopteryx" was indeed unable to use flapping flight as modern birds do, but it may well have used a downstroke-only flap-assisted gliding technique. "Archaeopteryx" wings were relatively large, which would have resulted in a low stall speed and reduced turning radius. The
Specimens of Archaeopteryx that "Archaeopteryx" could have raised its short second toe in a similar manner to its dromaeosaur cousin "Deinonychus", whose second toe bore an enlarged "sickle claw" for probable use in predation. It remains unclear whether "Archaeopteryx" also used its hyperextended toe-claw in a similar fashion or for some other purpose. Taken together, many of the unique features of the skull, feet and coracoid strengthen a close relationship of archaeopterygids to deinonychosaurian theropods. Well-preserved imprints of wing and tail plumage show details of the barbs, but not barbules, and the shafts of 11 primaries can be counted on the right wing.