Equus (genus) 54 million years ago (Mya). These animals had three toes on the hind feet and four on the front feet with small hooves in place of claws, but also had soft pads.<ref name="doi10.1126/science.1105458"></ref> Equids developed into larger, three-toed animals (e.g. "Mesohippus") during the Oligocene and Miocene. From there, the side toes became progressively smaller through the Pleistocene until the emergence of the single-toed "Equus". The genus "Equus", which includes all extant equines, is believed to have evolved from "Dinohippus", via the intermediate form "Plesippus". One of the oldest species is "Equus simplicidens", described as zebra-like with a donkey-like head shape.
Equus (genus) each other both visually and vocally. Human activities have threatened wild equine populations. The word "equus" is Latin for "horse", and is cognate with the Greek ἵππος ("hippos"), "horse", and Mycenaean Greek "i-qo" /ikkʷos/, the earliest attested variant of the Greek word, written in Linear B syllabic script.; compare the alternative development of the Proto-Greek labiovelar in Ionic ἴκκος ("ikkos"). The genus "Equus" was first described by Carl Linnaeus in 1758. It is the only recognized extant genus in the family Equidae. The first equids were small, dog-sized mammals (e.g. "Eohippus") adapted for browsing on shrubs during the Eocene, around