Feathered dinosaur is possible that feathers first developed in an even earlier group, in light of the discovery of pycnofibers of pterosaurs. Crocodilians also possess beta keratin similar to those of birds, which suggests that they evolved from common ancestral genes. Shortly after the 1859 publication of Charles Darwin's "On the Origin of Species", British biologist Thomas Henry Huxley proposed that birds were descendants of dinosaurs. He compared the skeletal structure of "Compsognathus", a small theropod dinosaur, and the 'first bird' "Archaeopteryx lithographica" (both of which were found in the Upper Jurassic Bavarian limestone of Solnhofen). He showed that, apart from its
Specimens of Archaeopteryx on the main slab. Both arms are preserved close to their natural articulation with the shoulder girdle, though displaced. The ulna is more robust and curved than the slender radius. A confusing jumble of metacarpals has led some researchers to conclude the beginning of fusion between carpals and metacarpals; this has been rejected by later researchers. Most manual bones of the right wing are missing, and the left wing preserves two metacarpals, the semilunate carpal, and two long, slender phalanges with large, strongly-curved claws. The missing third finger led Owen to conclude a two-fingered hand for "Archaeopteryx" in his original