Archaeopteryx for flightless birds. In 2010, Robert L. Nudds and Gareth J. Dyke in the journal "Science" published a paper in which they analysed the rachises of the primary feathers of "Confuciusornis" and "Archaeopteryx". The analysis suggested that the rachises on these two genera were thinner and weaker than those of modern birds relative to body mass. The authors determined that "Archaeopteryx" and "Confuciusornis", were unable to use flapping flight. This study was criticized by Philip J. Currie and Luis Chiappe. Chiappe suggested that it is difficult to measure the rachises of fossilized feathers, and Currie speculated that "Archaeopteryx" and "Confuciusornis"
Specimens of Archaeopteryx — limiting its ability to perch on branches and implying a terrestrial or trunk-climbing lifestyle. This has been interpreted as evidence of theropod ancestry. In 1988, Gregory S. Paul claimed to have found evidence of a hyperextensible second toe, but this was not verified and accepted by other scientists until the Thermopolis specimen was described. "Until now, the feature was thought to belong only to the species' close relatives, the deinonychosaurs." The Thermopolis Specimen was assigned to "Archaeopteryx siemensii" in 2007. The specimen is considered the most complete and well preserved "Archaeopteryx" remains yet. This specimen first came to light