Specimens of Archaeopteryx feather's earlier discovery, von Meyer made the London fossil the type specimen of the genus ("Archaeopteryx") and species ("lithographica"), establishing it as the exemplar of the species' features. Its discovery and subsequent descriptions, a mere handful of years after Darwin's "On the Origin of Species", fulfilled the hope of a truly transitional form in its then-unique amalgamation of bird and reptilian characters. The specimen is preserved on two counter slabs broken originally into four pieces. The main, overlaying slab contains most of the skeletal elements, while only impressions and a few bone fragments are visible on the counter slab. This
Archaeopteryx be related particularly closely to "Anchiornis" and "Xiaotingia", which were recovered as basal troodontids instead. Agnolín and Novas (2013) found "Archaeopteryx" and (possibly synonymous) "Wellnhoferia" to be the basalmost avialans (Avialae being defined by the authors as including "Archaeopteryx lithographica" and "Passer", their most recent common ancestor and all of its descendants), with Microraptoria, Unenlagiinae, and the clade containing "Anchiornis" and "Xiaotingia" being successively closer outgroups to the Avialae. Another phylogenetic study by Godefroit, "et al.", using a more inclusive matrix than the one from the analysis in the description of "Eosinopteryx brevipenna", also found "Archaeopteryx" to be a member