Equus (genus) 54 million years ago (Mya). These animals had three toes on the hind feet and four on the front feet with small hooves in place of claws, but also had soft pads.<ref name="doi10.1126/science.1105458"></ref> Equids developed into larger, three-toed animals (e.g. "Mesohippus") during the Oligocene and Miocene. From there, the side toes became progressively smaller through the Pleistocene until the emergence of the single-toed "Equus". The genus "Equus", which includes all extant equines, is believed to have evolved from "Dinohippus", via the intermediate form "Plesippus". One of the oldest species is "Equus simplicidens", described as zebra-like with a donkey-like head shape.
Proconsul africanus has a talus in which the trochlear surface is highly curved and deeply grooved. The foot of this species possessed a transverse arch. "Proconsul africanus" had a brachial index of 96 which is comparable to the extant genus "Pan". Overall the skeleton of this species can be described as being robust. This species had an average body mass of around . Based upon the dental morphology, it is conjectured that "Proconsul africanus" was a frugivorous species. A study by Aiello in 1981 concluded it was a "below-branch feeder." Based on postcranial pieces, "Proconsul africanus" was likely an arboreal quadruped. Proconsul