Archaeopteryx for flightless birds. In 2010, Robert L. Nudds and Gareth J. Dyke in the journal "Science" published a paper in which they analysed the rachises of the primary feathers of "Confuciusornis" and "Archaeopteryx". The analysis suggested that the rachises on these two genera were thinner and weaker than those of modern birds relative to body mass. The authors determined that "Archaeopteryx" and "Confuciusornis", were unable to use flapping flight. This study was criticized by Philip J. Currie and Luis Chiappe. Chiappe suggested that it is difficult to measure the rachises of fossilized feathers, and Currie speculated that "Archaeopteryx" and "Confuciusornis"
Specimens of Archaeopteryx on the main slab. Both arms are preserved close to their natural articulation with the shoulder girdle, though displaced. The ulna is more robust and curved than the slender radius. A confusing jumble of metacarpals has led some researchers to conclude the beginning of fusion between carpals and metacarpals; this has been rejected by later researchers. Most manual bones of the right wing are missing, and the left wing preserves two metacarpals, the semilunate carpal, and two long, slender phalanges with large, strongly-curved claws. The missing third finger led Owen to conclude a two-fingered hand for "Archaeopteryx" in his original