Feather a dromaeosaurid found in the Yixian formation (124.6 MYA). Previously, a temporal paradox existed in the evolution of feathers—theropods with highly derived bird-like characteristics occurred at a later time than "Archaeopteryx"—suggesting that the descendants of birds arose before the ancestor. However, the discovery of "Anchiornis huxleyi" in the Late Jurassic Tiaojishan Formation (160 MYA) in western Liaoning in 2009 resolved this paradox. By predating "Archaeopteryx", "Anchiornis" proves the existence of a modernly feathered theropod ancestor, providing insight into the dinosaur-bird transition. The specimen shows distribution of large pennaceous feathers on the forelimbs and tail, implying that pennaceous feathers spread to
Specimens of Archaeopteryx — limiting its ability to perch on branches and implying a terrestrial or trunk-climbing lifestyle. This has been interpreted as evidence of theropod ancestry. In 1988, Gregory S. Paul claimed to have found evidence of a hyperextensible second toe, but this was not verified and accepted by other scientists until the Thermopolis specimen was described. "Until now, the feature was thought to belong only to the species' close relatives, the deinonychosaurs." The Thermopolis Specimen was assigned to "Archaeopteryx siemensii" in 2007. The specimen is considered the most complete and well preserved "Archaeopteryx" remains yet. This specimen first came to light