River Tywi these fish to over in weight. In summer and autumn there is also a substantial run of Atlantic salmon ("Salmo salar"). In May the Tywi has a run of the rare and protected twaite shad ("Alosa fallax") and allis shad ("Alosa alosa"). The Tywi also contains brown trout, eels, pike, and a variety of small fish species, and is home to brook lampreys, river lampreys and sea lampreys. The Tywi has the distinction of having accidentally produced by far the biggest fish ever taken on rod and line in fresh water in Britain. This was a sturgeon ("Acipwienser sturio") weighing
Alosa lifespan of "Alosa" species can be up to 10 years, but this is generally uncommon, as many die after spawning. The systematics and distribution of "Alosa" shads are complex. The genus inhabits a wide range of habitats, and many taxa are migratory. A few forms are landlocked, including one from Killarney in Ireland, two from lakes in northern Italy, and two in Greece. Several species are native to the Black Sea and Caspian Sea. "Alosa" species of the Caspian are systemically characterized by the number of rakers on the first gill arch. They are classified as being "multirakered", "medium-rakered", or