Origin of birds mitochondrial uncoupling protein, UCP1, which is thermogenic. In mammals, UCP1 functions within brown adipose tissue to protect newborns against hypothermia. In modern birds, skeletal muscle serves a similar function and is presumed to have done so in their ancestors. In this view, bipedality and other avian skeletal alterations were side effects of muscle hyperplasia, with further evolutionary modifications of the forelimbs, including adaptations for flight or swimming, and vestigiality, being secondary consequences of two-leggedness. "Archaeopteryx" has historically been considered the first bird, or "Urvogel". Although newer fossil discoveries filled the gap between theropods and "Archaeopteryx", as well as the gap
Specimens of Archaeopteryx that "Archaeopteryx" could have raised its short second toe in a similar manner to its dromaeosaur cousin "Deinonychus", whose second toe bore an enlarged "sickle claw" for probable use in predation. It remains unclear whether "Archaeopteryx" also used its hyperextended toe-claw in a similar fashion or for some other purpose. Taken together, many of the unique features of the skull, feet and coracoid strengthen a close relationship of archaeopterygids to deinonychosaurian theropods. Well-preserved imprints of wing and tail plumage show details of the barbs, but not barbules, and the shafts of 11 primaries can be counted on the right wing.