Specimens of Archaeopteryx length. These hindlimb proportions agree roughly with those of the London specimen. The claw arc of the central pedal digit is about 120 degrees, which is close to the average for all "Archaeopteryx" specimens. This arc is also close to the average for that in perching birds and is well above the mean for ground-dwelling birds, despite conclusions against perching habits based on the short hallux and weak flexor tubercles. The remarkably preserved plumage is one of many features that sets the Berlin specimen head-and-coracoids above all other "Archaeopteryx" fossils to date. Clear feather impressions are visible on both wings,
Archaeopteryx be black throughout, with heavier pigmentation in the distal tip. The feather studied was most probably a dorsal covert, which would have partly covered the primary feathers on the wings. The study does not mean that "Archaeopteryx" was entirely black, but suggests that it had some black colouration which included the coverts. Carney pointed out that this is consistent with what we know of modern flight characteristics, in that black melanosomes have structural properties that strengthen feathers for flight. In a 2013 study published in the "Journal of Analytical Atomic Spectrometry", new analyses of "Archaeopteryx"'s feathers revealed that the animal