River Tywi these fish to over in weight. In summer and autumn there is also a substantial run of Atlantic salmon ("Salmo salar"). In May the Tywi has a run of the rare and protected twaite shad ("Alosa fallax") and allis shad ("Alosa alosa"). The Tywi also contains brown trout, eels, pike, and a variety of small fish species, and is home to brook lampreys, river lampreys and sea lampreys. The Tywi has the distinction of having accidentally produced by far the biggest fish ever taken on rod and line in fresh water in Britain. This was a sturgeon ("Acipwienser sturio") weighing
Coastal fish but ascend rivers to spawn; eels are catadromous, living in rivers and streams, but returning to the sea to breed. Besides the species that migrate through estuaries, there are many other fish that use them as "nursery grounds" for spawning or as places young fish can feed and grow before moving elsewhere. For example, herring and plaice are two commercially important species that use the Thames Estuary for this purpose. Mangrove swamps are associated brackish water habitats. Many, though not all, mangrove swamps fringe estuaries and lagoons where the salinity changes with each tide. Among the most specialised residents of