Origin of birds mitochondrial uncoupling protein, UCP1, which is thermogenic. In mammals, UCP1 functions within brown adipose tissue to protect newborns against hypothermia. In modern birds, skeletal muscle serves a similar function and is presumed to have done so in their ancestors. In this view, bipedality and other avian skeletal alterations were side effects of muscle hyperplasia, with further evolutionary modifications of the forelimbs, including adaptations for flight or swimming, and vestigiality, being secondary consequences of two-leggedness. "Archaeopteryx" has historically been considered the first bird, or "Urvogel". Although newer fossil discoveries filled the gap between theropods and "Archaeopteryx", as well as the gap
Specimens of Archaeopteryx skeleton of a feathered animal from the same lithographic shale deposits, which would later be known as the London specimen. Coincidentally, von Meyer proposed the name "Archaeopteryx lithographica" for the feather, but not for the skeleton. Therefore, the official name of the animal was originally linked to the single feather rather than any actual skeleton, and is formally considered the original holotype. Though 1860 is often the year named for the feather's discovery, there is no proof of this date, and some authors consider it more likely to have been found in 1861, as it seems reasonable that von Meyer's