Acacia heterophylla 1.4 million years ago. "A. heterophylla" sequences nest within those of the more diverse "A. koa", making the latter species paraphyletic. Both species are descended from an ancestral species in Australia, presumably their sister species, "Acacia melanoxylon"; the means of dispersal to Hawaii and then to Réunion (the latter trip a distance of 18,000 km) is thought to have been seed-carrying by birds such as petrels (the seeds of these species are not adapted for prolonged immersion in seawater). Both species also have very similar ecological niches, which differ from that of "A. melanoxylon". Acacia heterophylla Acacia heterophylla, the highland
Acacia koa used by ancient Hawaiians to build "waa" (dugout outrigger canoes) and "papa hee nalu" (surfboards). Only "paipo" (bodyboards), "kikoo", and "alaia" surfboards were made from koa, however; "olo", the longest surfboards, were made from the lighter and more buoyant "wiliwili" ("Erythrina sandwicensis"). The reddish wood is very similar in strength and weight to that of Black Walnut ("Juglans nigra"), with a specific gravity of 0.55, and is sought for use in wood carving and furniture. Koa is also a tonewood, often used in the construction of ukuleles, acoustic guitars, and Weissenborn-style Hawaiian steel guitars. B.C. Rich used koa on some