Greater koa finch and maximum growth of a single species of koa finch. The greater koa finch was a granivore, preferring the pods and seeds of koa ("Acacia koa"). It was also observed eating caterpillars and aalii ("Dodonaea viscosa") seeds. The last confirmed sighting was in 1896, although there were sporadic later reports. Like its close relative the lesser koa finch, this bird lived in small stretches of mesic forest on the Big Island of Hawaii. These forests were dominated by its preferred food source, koa ("Acacia koa"), and were logged and replaced with pasture beginning in 1850. Grazing by cattle inhibited the
Acacia koa confirmed this close relationship; the estimated time of divergence is about 1.4 million years ago. "A. heterophylla" sequences nest within those of the more diverse "A. koa", making the latter paraphyletic. Both species are thought to be descended from an ancestral species in Australia, presumably their sister species, "Acacia melanoxylon". Dispersals most likely occurred via seed-carrying by birds such as petrels. Both species have very similar ecological niches, which differ from that of "A. melanoxylon". A closely related species, koaia or koaie ("A. koaia"), is found in dry areas. It is most easily distinguished by having smaller seeds that are