Acacia heterophylla 1.4 million years ago. "A. heterophylla" sequences nest within those of the more diverse "A. koa", making the latter species paraphyletic. Both species are descended from an ancestral species in Australia, presumably their sister species, "Acacia melanoxylon"; the means of dispersal to Hawaii and then to Réunion (the latter trip a distance of 18,000 km) is thought to have been seed-carrying by birds such as petrels (the seeds of these species are not adapted for prolonged immersion in seawater). Both species also have very similar ecological niches, which differ from that of "A. melanoxylon". Acacia heterophylla Acacia heterophylla, the highland
Koa wilt mortality rate over several years due to Koa wilt. It is also a legume, giving it the ability to form symbiosis with nitrogen fixing bacteria, which has led to efforts to use koa in agroecological systems with crops such as coffee and cacao. Ecologically, koa is an important species because it is one of the few native trees that remains dominant in alien mixed forests on the Hawaiian Islands. Invasive species make up the majority of the State's current plant population. Koa trees also are important because provide a habitat for many native bird species. Historically, there were once two