Acacia heterophylla 1.4 million years ago. "A. heterophylla" sequences nest within those of the more diverse "A. koa", making the latter species paraphyletic. Both species are descended from an ancestral species in Australia, presumably their sister species, "Acacia melanoxylon"; the means of dispersal to Hawaii and then to Réunion (the latter trip a distance of 18,000 km) is thought to have been seed-carrying by birds such as petrels (the seeds of these species are not adapted for prolonged immersion in seawater). Both species also have very similar ecological niches, which differ from that of "A. melanoxylon". Acacia heterophylla Acacia heterophylla, the highland
Hakalau Forest National Wildlife Refuge numerous deep gulches. Rainfall decreases to about at elevations above , where majestic koa ("Acacia koa") and red-blossomed ōhia lehua ("Metrosideros polymorpha") trees form a closed-canopy forest. Further upslope, above , rainfall decreases to or less and native forest merges into abandoned pastureland where alien grasses and weeds, introduced as forage for cattle, are the dominant vegetation. Since 1989, over 400,000 koa, ōhia, and other native plants have been planted in this area as part of the refuge's reforestation program. The Pua Akala Cabin, located within the refuge, was nominated to the National Register of Historic Places in August 2008.