Acacia koa for up to 25 years. Scarification is needed before "A. koa" seeds will germinate. Koa is endemic to the islands of Hawaii, Molokai, Maui, Lānai, Oahu, and Kauai, where it grows at elevations of . It requires of annual rainfall. Acidic to neutral soils (pH of 4-7.4) that are either an Inceptisol derived from volcanic ash or a well-drained histosol are preferred. Its ability to fix nitrogen allows it to grow in very young volcanic soils. Koa and ōhia lehua ("Metrosideros polymorpha") dominate the canopy of mixed mesic forests. It is also common in wet forests. The koa's trunk was
Acacia koa confirmed this close relationship; the estimated time of divergence is about 1.4 million years ago. "A. heterophylla" sequences nest within those of the more diverse "A. koa", making the latter paraphyletic. Both species are thought to be descended from an ancestral species in Australia, presumably their sister species, "Acacia melanoxylon". Dispersals most likely occurred via seed-carrying by birds such as petrels. Both species have very similar ecological niches, which differ from that of "A. melanoxylon". A closely related species, koaia or koaie ("A. koaia"), is found in dry areas. It is most easily distinguished by having smaller seeds that are