Greater koa finch below. It had a thick black bill which allowed it to break open seed pods that were found in the trees. In historical times, its range was largely confined to the Kona District of the island of Hawaii, although it was observed in the Kīlauea area in 1895. It was much more widespread prior to the arrival of humans in Hawaii, and related species lived on other islands. In Kona, it co-occurred with the closely related lesser koa finch ("Rhodocanthis flaviceps"), and the Kona grosbeak. The lesser and greater koa finches were once thought by scientists to be the minimum
Hawaiian tropical rainforests milo ("Thespesia populnea") for the past 1,000–2,000 years. The Polynesian-introduced noni ("Morinda citrifolia"), pia ("Tacca leontopetaloides"), and kī ("Cordyline fruticosa") are also common in this zone. Other native species include pololei ("Ophioglossum concinnum"), ākia ("Wikstroemia" spp.), loulu fan palms ("Pritchardia" spp.), ōhia lehua ("Metrosideros polymorpha"), and lama ("Diospyros sandwicensis"). Mixed mesic forests, at on the windward slopes of the large islands in addition to the summit of Mount Lānaihale on Lānai, receive of rainfall annually and thus may not be true rainforests. The forest canopy, dominated by koa ("Acacia koa") and ōhia lehua ("Metrosideros polymorpha"), is somewhat open, but tree