Acacia heterophylla 1.4 million years ago. "A. heterophylla" sequences nest within those of the more diverse "A. koa", making the latter species paraphyletic. Both species are descended from an ancestral species in Australia, presumably their sister species, "Acacia melanoxylon"; the means of dispersal to Hawaii and then to Réunion (the latter trip a distance of 18,000 km) is thought to have been seed-carrying by birds such as petrels (the seeds of these species are not adapted for prolonged immersion in seawater). Both species also have very similar ecological niches, which differ from that of "A. melanoxylon". Acacia heterophylla Acacia heterophylla, the highland
ʻAnianiau Its call is a pair of notes, "tew-weet", while its song is a trill of "wee-see, wee-see, wee-see". Anianiau are found in mesic and wet forests at elevations above . The highest densities occur above . Dominant tree species in its habitat include koa ("Acacia koa"), ōhia lehua ("Metrosideros polymorpha"), ōlapa ("Cheirodendron trigynum"), and lapalapa ("C. platyphyllum"). The anianiau mainly feeds on nectar from the flowers of plants such as ōhia lehua ("Metrosideros polymorpha"), ōhelo ("Vaccinium" spp.), and alani ("Melicope" spp.). It will also take arthropods from trees, shrubs, or vines. The anianiau breeding season ranges from February to June.