Acacia heterophylla Acacia heterophylla Acacia heterophylla, the highland tamarind, is a tree (or shrub in its higher places) endemic to Réunion island where it is commonly named tamarin des hauts The tree has a juvenile stage where its leaves have a pinnate arrangement, but in the adult stage the leaves diminish and the phyllode becomes the dominant photosynthetic structure. It has been introduced into Madagascar where it grows in a subhumid climate at an altitude of about 500–1000 m above sea level. Genetic sequence analysis has shown its closest relative is "Acacia koa" of Hawaii; the estimated time of divergence is about
Acacia koa confirmed this close relationship; the estimated time of divergence is about 1.4 million years ago. "A. heterophylla" sequences nest within those of the more diverse "A. koa", making the latter paraphyletic. Both species are thought to be descended from an ancestral species in Australia, presumably their sister species, "Acacia melanoxylon". Dispersals most likely occurred via seed-carrying by birds such as petrels. Both species have very similar ecological niches, which differ from that of "A. melanoxylon". A closely related species, koaia or koaie ("A. koaia"), is found in dry areas. It is most easily distinguished by having smaller seeds that are