Scotorythra paludicola Scotorythra paludicola Scotorythra paludicola, the koa looper moth, is a moth of the family Geometridae. It was first described by Arthur Gardiner Butler in 1879. It is endemic to the Hawaiian islands of Kauai, Oahu, Maui and Hawaii. The larvae mainly feed on "Acacia koa", but have also been recorded from "Falcataria moluccana", "Grevillea" and "Tristania" species. On occasion, this species develops enormous populations and defoliates entire areas with koa trees. During a recent outbreak of "S. paludicola" in 2013 on Hawaii Island, larvae were found to feed on isolated populations of the invasive tree "Falcataria moluccana" near Akaka Falls
Acacia koa confirmed this close relationship; the estimated time of divergence is about 1.4 million years ago. "A. heterophylla" sequences nest within those of the more diverse "A. koa", making the latter paraphyletic. Both species are thought to be descended from an ancestral species in Australia, presumably their sister species, "Acacia melanoxylon". Dispersals most likely occurred via seed-carrying by birds such as petrels. Both species have very similar ecological niches, which differ from that of "A. melanoxylon". A closely related species, koaia or koaie ("A. koaia"), is found in dry areas. It is most easily distinguished by having smaller seeds that are