Acacia heterophylla 1.4 million years ago. "A. heterophylla" sequences nest within those of the more diverse "A. koa", making the latter species paraphyletic. Both species are descended from an ancestral species in Australia, presumably their sister species, "Acacia melanoxylon"; the means of dispersal to Hawaii and then to Réunion (the latter trip a distance of 18,000 km) is thought to have been seed-carrying by birds such as petrels (the seeds of these species are not adapted for prolonged immersion in seawater). Both species also have very similar ecological niches, which differ from that of "A. melanoxylon". Acacia heterophylla Acacia heterophylla, the highland
Koa wilt to the Hawaiian islands. It ranges in size from 15 to greater than 50 feet with a canopy spread of 20 to 40 feet. It has a showy white flower and blooms sporadically. Mature leaves are sickle shaped. Koa wilt is typically a fatal pathogen for its host. In only a few months, a tree may lose its entire canopy and die. Trees less than fifteen years old are typically the most susceptible. Symptoms include stains in cambium, roots, and sapwood as well as chlorosis (yellowing), canopy dieback, brown and wilted leaves, oozing sap, and bark cankers. The Koa wilt