Asiatic lion known as "Indian lion" and "Persian lion". Following Meyer's first description of an Asiatic lion skin from Persia, other naturalists and zoologists also described lion specimens from other parts of Asia that used to be considered synonyms of "P. l. persica": In 2017, the Asiatic lion was subsumed to "P. l. leo" due to close morphological and molecular genetic similarities with Barbary lion specimens. Fossil remains of "Panthera spelaea" excavated in the Cromer Stage indicate that it represented a genetically isolated and highly distinct lineage. Fossil lion remains were found in Pleistocene deposits in West Bengal. A fossil carnassial excavated
Apex predator expected to produce an increase in the numbers of cubs observed. Dozens of other species, including eagles, ravens, magpies, coyotes and black bears have also been documented as scavenging from wolf kills within the park. Ecologists have debated whether humans are apex predators. Sylvain Bonhommeau and colleagues argued in 2013 that across the global food web, a fractional human trophic level (HTL) can be calculated as the mean trophic level of every species in the human diet, weighted by the proportion which that species forms in the diet. This analysis gives an average HTL of 2.21, varying between 2.04 (for