Asiatic lion in size to the Central African lion, and smaller than large African lions. Adult males weigh , while females weigh . Colour and development of manes in male lions varies between regions, among populations and with age of lions. In general, the Asiatic lion differs from the African lion by a less developed mane. The manes of most lions in ancient Greece and Asia Minor were also less developed and did not extend to below the belly, sides or ulnas. Lions with such smaller manes were also known in the Syrian region, Arabian peninsula and Egypt, whereas Barbary and Cape
Apex predator expected to produce an increase in the numbers of cubs observed. Dozens of other species, including eagles, ravens, magpies, coyotes and black bears have also been documented as scavenging from wolf kills within the park. Ecologists have debated whether humans are apex predators. Sylvain Bonhommeau and colleagues argued in 2013 that across the global food web, a fractional human trophic level (HTL) can be calculated as the mean trophic level of every species in the human diet, weighted by the proportion which that species forms in the diet. This analysis gives an average HTL of 2.21, varying between 2.04 (for