Atlas the Barbary lion versus the Bengal tiger of Simla its subspecies, was "the largest or most formidable of all African felidae", and therefore, that it was bigger than the Asiatic lion. However, it is not unanimously held that the Barbary and Cape lions were the largest lions (See "Physical comparison of tigers and lions"). Nevertheless, an English Colonel called 'Maitland', who supported the tiger, described the lion as a 'monster'. Both were hungry, as their diets had been reduced from their regular intake of food, beforehand. "Gettysburg Compiler" (1899) said "If the tiger was the personification of graceful strength and supple energy, the lion was the embodiment of massive
Apex predator Burundi, with a 96.7% plant-based diet) and 2.57 (for Iceland, with 50% meat and fish, 50% plants). These values are comparable to those of non-apex predators like anchovy or pig. Peter D. Roopnarine criticised Bonhommeau's approach, arguing that humans are apex predators, and that the HTL was based on terrestrial farming where indeed humans have a low trophic level, mainly eating producers (crop plants at level 1) or primary consumers (herbivores at level 2), which as expected places humans at a level slightly above 2. Roopnarine instead calculated the position of humans in two marine ecosystems, a Caribbean coral reef