Atlas the Barbary lion versus the Bengal tiger of Simla its subspecies, was "the largest or most formidable of all African felidae", and therefore, that it was bigger than the Asiatic lion. However, it is not unanimously held that the Barbary and Cape lions were the largest lions (See "Physical comparison of tigers and lions"). Nevertheless, an English Colonel called 'Maitland', who supported the tiger, described the lion as a 'monster'. Both were hungry, as their diets had been reduced from their regular intake of food, beforehand. "Gettysburg Compiler" (1899) said "If the tiger was the personification of graceful strength and supple energy, the lion was the embodiment of massive
Apex predator expected to produce an increase in the numbers of cubs observed. Dozens of other species, including eagles, ravens, magpies, coyotes and black bears have also been documented as scavenging from wolf kills within the park. Ecologists have debated whether humans are apex predators. Sylvain Bonhommeau and colleagues argued in 2013 that across the global food web, a fractional human trophic level (HTL) can be calculated as the mean trophic level of every species in the human diet, weighted by the proportion which that species forms in the diet. This analysis gives an average HTL of 2.21, varying between 2.04 (for