Gondwana Massif have been dated at 320–330 million years old, indicating that the subduction process initiated in the early Carboniferous.This was relatively short lived (lasting about 20 million years), and initial contact of the two landmasses occurred in the mid-Carboniferous, with broader collision during the early Permian. In the Devonian an island arc named Chaitenia accreted to Patagonia in what is now south-central Chile. Gondwana and Laurasia formed the Pangaea supercontinent during the Carboniferous. Pangaea began to break up in the Mid-Jurassic when the Central Atlantic opened. In the western end of Pangaea, the collision between Gondwana and Laurasia closed the
Gondwana (57% of marine families and 83% of genera went extinct) which came to dominate in the Late Permian and from whom true conifers evolved. Tall lycopods and horsetails dominated the wetlands of Gondwana in the Early Permian. Insects co-evolved with glossopterids across Gondwana and diversified with more than 200 species in 21 orders by the Late Permian, many known from South Africa and Australia. Beetles and cockroaches remained minor elements in this fauna. Tetrapod fossils from the Early Permian have only been found in Laurasia but they became common in Gondwana later during the Permian. The arrival of the therapsids