Gondwana As Gondwana began to rotate clockwise, Australia shifted south to more temperate latitudes. An ice cap initially covered most of southern Africa and South America but began to spread to eventually cover most of the supercontinent, save for northern-most Africa-South America and eastern Australia. Giant lycopod and horsetail forests continued to evolve in tropical Laurasia together with a diversified assemblage of true insects. In Gondwana, in contrast, ice and, in Australia, volcanism decimated the Devonian flora to a low-diversity seed fern flora – the pteridophytes were increasingly replaced by the gymnosperms which were to dominate until the Mid-Cretaceous. Australia, however,
Gondwana from all continents in the Southern Hemisphere, has a "Gondwanan distribution" and is often described as an archaic, or relict, lineage. The distributions in Proteaceae is, nevertheless, the result of both Gondwanan rafting and later oceanic dispersal. During the Silurian Gondwana extended from the Equator (Australia) to the South Pole (North Africa and South America) whilst Laurasia was located on the Equator opposite to Australia. A short-lived Late Ordovician glaciation was followed by a Silurian Hot House period. The End-Ordovician extinction, which resulted in 27% of marine invertebrate families and 57% of genera going extinct, occurred during this shift from