Pangaea and along the adjacent margins of east Africa, Antarctica and Madagascar, new rifts were forming that would lead to the formation of the southwestern Indian Ocean that would open up in the Cretaceous. The second major phase in the break-up of Pangaea began in the Early Cretaceous (150–140 Ma), when the minor supercontinent of Gondwana separated into multiple continents (Africa, South America, India, Antarctica, and Australia). The subduction at Tethyan Trench probably caused Africa, India and Australia to move northward, causing the opening of a "South Indian Ocean". In the Early Cretaceous, Atlantica, today's South America and Africa, finally separated
Gondwana (57% of marine families and 83% of genera went extinct) which came to dominate in the Late Permian and from whom true conifers evolved. Tall lycopods and horsetails dominated the wetlands of Gondwana in the Early Permian. Insects co-evolved with glossopterids across Gondwana and diversified with more than 200 species in 21 orders by the Late Permian, many known from South Africa and Australia. Beetles and cockroaches remained minor elements in this fauna. Tetrapod fossils from the Early Permian have only been found in Laurasia but they became common in Gondwana later during the Permian. The arrival of the therapsids