Nearctic realm represented by the Madrean Region. Although North America and South America are presently joined by the Isthmus of Panama, these continents were separated for about 180 million years, and evolved very different plant and animal lineages. When the ancient supercontinent of Pangaea split into two about 180 million years ago, North America remained joined to Eurasia as part of the supercontinent of Laurasia, while South America was part of the supercontinent of Gondwana. North America later split from Eurasia. North America has been joined by land bridges to both Asia and South America since then, which allowed an exchange of
Gondwana During the Devonian giant club mosses replaced the Baragwanathia Flora, introducing the first trees, and by the Late Devonian this first forest was accompanied by the progymnosperms, including the first large trees "Archaeopteris". The Late Devonian extinction probably also resulted in osteolepiform fishes evolving into the amphibian tetrapods, the earliest land vertebrates, in Greenland and Russia. The only traces of this evolution in Gondwana are amphibian footprints and a single jaw from Australia. The closure of the Rheic Ocean and the formation of Pangaea in the Carboniferous resulted in the rerouting of ocean currents which initiated an Ice House period.