Feathered dinosaur the deciduous sheath, forming a tuft of unbranched barbs with a basal calamus. Stage III consists of two developmental novelties, IIIa and IIIb, as either could have occurred first. Stage IIIa involves helical displacement of barb ridges arising within the collar. The barb ridges on the anterior midline of the follicle fuse together, forming the rachis. The creation of a posterior barb locus follows, giving an indeterminate number of barbs. This resulted in a feather with a symmetrical, primarily branched structure with a rachis and unbranched barbs. In stage IIIb, barbules paired within the peripheral barbule plates of the barb
Rachis achieved pachytene and are attached to the walls of the gonadal tube. The rachis is filled with cytoplasm. In plants, a "rachis" is the main axis of a compound structure. It can be the main stem of a compound leaf, such as in "Acacia" or ferns, or the main, flower-bearing portion of an inflorescence above a supporting peduncle. A ripe head of wild-type wheat is easily shattered into dispersal units when touched or blown by the wind. A series of layers forms that divides the rachis into dispersal units consisting of a small group of flowers (a single spikelet) attached