Origins of Hutu, Tutsi and Twa adults has a high capacity for lactose digestion, a surprisingly high number for an agrarian people, which Mamdani suggests may be the result of centuries of intermarriage with Tutsi. Bethwell Ogot in the 1988 UNESCO "General History" further notes that the number of pastoralists in Rwanda increased sharply around the fifteenth century. Although Luis et al. (2004) in a more general study on bi-allelic markers in many African countries found a statistically significant genetic difference between Tutsi and Hutu, the overall difference were not large. Modern-day genetic studies of the Y-chromosome generally indicate that the Tutsi, like the Hutu, are
Kangura 1993 issue stated that a Hutu soldier angry over the Arusha agreement would soon assassinate the president. The January 1994 issue predicted that Habyarimana would be killed in March. "Kangura" had stopped publishing by 6 April 1994, when the plane carrying Presidents Habyarimana and Cyprien Ntaryamira of Burundi was shot down over Kigali. This sparked the genocide. Over 800,000 people were killed, mostly Tutsis but including Hutu moderates and Hutus who attempted to protect Tutsi from the militias. Hassan Ngeze, the founder, editor and accountant for "Kangura" fled the country as the RPF took control. He was arrested in 1997