Origins of Hutu, Tutsi and Twa phenotype, genotype, cultural memory of inhabitants of Rwanda, archeology, and linguistics. Most Tutsis and Hutus carry the E1b1a paternal haplogroup, which is common among Bantu populations. More recent studies have de-emphasized physical appearance, such as height and nose width, in favor of examining blood factors, the presence of the sickle cell trait, lactose intolerance in adults, and other genotype expressions. Excoffier et al. (1987) found that the Tutsi and Hima, despite being surrounded by Bantu populations, are "closer genetically to Cushites and Ethiosemites". Another study concluded that, while the sickle cell trait among the Rwandan Hutu was comparable to that
History of Rwanda dominated by the majority Hutu under President Grégoire Kayibanda. Unsettled ethnic and political tensions were worsened when Juvénal Habyarimana, who was also Hutu, seized power in 1973. In 1990, the Rwandan Patriotic Front (RPF), a rebel group composed of 10,000 Tutsi refugees from previous decades of unrest, invaded the country, starting the Rwandan Civil War. The war ground on, worsening ethnic tensions, as the Hutu feared losing their gains. The assassination of Habyarimana was the catalyst for the eruption of the 1994 genocide, in which hundreds of thousands of Tutsis and some moderate Hutus were killed. The Tutsi RPF conquered