Origins of Hutu, Tutsi and Twa phenotype, genotype, cultural memory of inhabitants of Rwanda, archeology, and linguistics. Most Tutsis and Hutus carry the E1b1a paternal haplogroup, which is common among Bantu populations. More recent studies have de-emphasized physical appearance, such as height and nose width, in favor of examining blood factors, the presence of the sickle cell trait, lactose intolerance in adults, and other genotype expressions. Excoffier et al. (1987) found that the Tutsi and Hima, despite being surrounded by Bantu populations, are "closer genetically to Cushites and Ethiosemites". Another study concluded that, while the sickle cell trait among the Rwandan Hutu was comparable to that
Rwandan genocide the capital and the country. An estimated 2,000,000 Rwandans, mostly Hutu, were displaced and became refugees. The genocide was planned by members of the core Hutu political elite, many of whom occupied positions at top levels of the national government. Perpetrators came from the Rwandan army, the Gendarmerie, and government-backed militias including the "Interahamwe" and "Impuzamugambi". The genocide took place in the context of the Rwandan Civil War, a conflict beginning in 1990 between the Hutu-led government and the Rwandan Patriotic Front (RPF). The latter was made up largely of Tutsi refugees whose families had fled to Uganda after the