Origins of Hutu, Tutsi and Twa phenotype, genotype, cultural memory of inhabitants of Rwanda, archeology, and linguistics. Most Tutsis and Hutus carry the E1b1a paternal haplogroup, which is common among Bantu populations. More recent studies have de-emphasized physical appearance, such as height and nose width, in favor of examining blood factors, the presence of the sickle cell trait, lactose intolerance in adults, and other genotype expressions. Excoffier et al. (1987) found that the Tutsi and Hima, despite being surrounded by Bantu populations, are "closer genetically to Cushites and Ethiosemites". Another study concluded that, while the sickle cell trait among the Rwandan Hutu was comparable to that
Ethnic groups in Burundi declared a presidential republic; however peace did not ensue as a result. Even after 1966, the subsequent dictatorial regimes of Michel Micombero (1966–76), Jean-Baptiste Bagaza (1976–87), and Pierre Buyoya (1987–93) were dominated by Tutsi elites which often consciously marginalized and discriminated against the Hutu majority. The first Hutu to become head of state was Melchior Ndadaye in 1993, although he was assassinated shortly after taking power. Ethnic violence peaked in 1972 when 100,000 people, mainly Hutu, were killed by the Tutsi regime in the Burundian Genocide. Twa have also been targeted, specifically in violence directed against the Tutsi with whom