Hutu and 14% of residents in Rwanda and Burundi, respectively. The Twa pygmies, the smallest of the two countries' principal populations, also share language and culture with the Hutu and Tutsi. However, they are distinguished by a considerably shorter stature. The Hutu are believed to have first emigrated to the Great Lake region from West Africa in the great Bantu expansion. Various theories have emerged to explain the purported physical differences between them and their fellow Bantu-speaking neighbors, the Tutsi. One such thesis, largely based on oral tradition, posits that the Tutsi experienced some admixture with or were partially descended from
Origins of Hutu, Tutsi and Twa largely of Bantu extraction (60% E1b1a, 20% B, 4% E3). Paternal genetic influences associated with the Horn of Africa and North Africa are few (under 3% E1b1b), and are ascribed to much earlier inhabitants who were assimilated. However, the Tutsi have considerably more haplogroup B paternal lineages (14.9% B) than do the Hutu (4.3% B). Trombetta et al. (2015) found 22.2% of E1b1b in a small sample of Tutsis from Burundi, but no bearers of the haplogroup among the local Hutu and Twa populations. The subclade was of the M293 variety, which suggests that the ancestors of Tutsis in this