Origins of Hutu, Tutsi and Twa phenotype, genotype, cultural memory of inhabitants of Rwanda, archeology, and linguistics. Most Tutsis and Hutus carry the E1b1a paternal haplogroup, which is common among Bantu populations. More recent studies have de-emphasized physical appearance, such as height and nose width, in favor of examining blood factors, the presence of the sickle cell trait, lactose intolerance in adults, and other genotype expressions. Excoffier et al. (1987) found that the Tutsi and Hima, despite being surrounded by Bantu populations, are "closer genetically to Cushites and Ethiosemites". Another study concluded that, while the sickle cell trait among the Rwandan Hutu was comparable to that
Kangura 1993 issue stated that a Hutu soldier angry over the Arusha agreement would soon assassinate the president. The January 1994 issue predicted that Habyarimana would be killed in March. "Kangura" had stopped publishing by 6 April 1994, when the plane carrying Presidents Habyarimana and Cyprien Ntaryamira of Burundi was shot down over Kigali. This sparked the genocide. Over 800,000 people were killed, mostly Tutsis but including Hutu moderates and Hutus who attempted to protect Tutsi from the militias. Hassan Ngeze, the founder, editor and accountant for "Kangura" fled the country as the RPF took control. He was arrested in 1997