Hutu as was the person constitutionally entitled to succeed him. This sparked a genocide in Burundi between Hutu political structures and the Tutsi military, in which an estimated 500,000 Burundians died. There were many mass killings of Tutsis and moderate Hutus; these events were deemed genocide by the United Nations International Commission of Inquiry for Burundi. While Tutsi remained in control of Burundi, the conflict resulted in genocide in Rwanda as well. A Tutsi rebel group, the Rwandan Patriotic Front, invaded Rwanda from Uganda, which started a civil war against Rwanda's Hutu government in 1990. A peace agreement was signed, but
Origins of Hutu, Tutsi and Twa largely of Bantu extraction (60% E1b1a, 20% B, 4% E3). Paternal genetic influences associated with the Horn of Africa and North Africa are few (under 3% E1b1b), and are ascribed to much earlier inhabitants who were assimilated. However, the Tutsi have considerably more haplogroup B paternal lineages (14.9% B) than do the Hutu (4.3% B). Trombetta et al. (2015) found 22.2% of E1b1b in a small sample of Tutsis from Burundi, but no bearers of the haplogroup among the local Hutu and Twa populations. The subclade was of the M293 variety, which suggests that the ancestors of Tutsis in this