Origins of Hutu, Tutsi and Twa adults has a high capacity for lactose digestion, a surprisingly high number for an agrarian people, which Mamdani suggests may be the result of centuries of intermarriage with Tutsi. Bethwell Ogot in the 1988 UNESCO "General History" further notes that the number of pastoralists in Rwanda increased sharply around the fifteenth century. Although Luis et al. (2004) in a more general study on bi-allelic markers in many African countries found a statistically significant genetic difference between Tutsi and Hutu, the overall difference were not large. Modern-day genetic studies of the Y-chromosome generally indicate that the Tutsi, like the Hutu, are
1996 Burundian coup d'état first democratic presidential election on 27 June 1993, which was won by Hutu Melchior Ndadaye. On 21 October, Ndadaye was assassinated, sparking the Burundi Civil War between Hutu and Tutsi groups. Moderate Hutu Cyprien Ntaryamira became president in February 1994, but both he and Rwandan president Juvénal Habyarimana were assassinated in April that year, an event that sparked the Rwandan Genocide. Ntaryamira was succeeded by another Hutu leader, Sylvestre Ntibantunganya. On 21 July 1996, Hutu rebels attacked a refugee camp in the country and killed more than 300 people. On 23 July, President Ntibantunganya went into hiding at the home