Pennaceous feather Pennaceous feather The pennaceous feather is a type of feather present in most modern birds and in some other species of maniraptoriform dinosaurs. A pennaceous feather has a stalk or quill. Its basal part, called a "calamus", is embedded in the skin. The calamus is hollow and has pith formed from the dry remains of the feather pulp, and the calamus opens below by an "inferior umbilicus" and above by a "superior umbilicus". The stalk above the calamus is a solid "rachis" having an umbilical groove on its underside. Pennaceous feathers have a rachis with "vanes" or "vaxillum " spreading
Calamus, Iowa Journal, associated himself with Mr. Barter and the paper was continued under the name of Barter and Sunderlin. It did not have a very long existence and for various reasons it suspended publication. Whip and Cook printed a newspaper here in the 1880s. It also had a very brief existence. The Calamus Record was the latest newspaper venture for Calamus. It was established in 1901 by A.J. Gault and enjoyed a longer existence than the other newspapers. Calamus is located at (41.826669, -90.759793). Olive Township, Clinton County, Iowa According to the United States Census Bureau, the city has a total