Origins of Hutu, Tutsi and Twa of neighboring people, it was almost non-existent among Rwandan Tutsi. Presence of the sickle cell trait is evidence of survival in the presence of malaria over many centuries, suggesting differing origins. Regional studies of the ability to digest lactose are also supportive. The ability to digest lactose among adults is widespread only among desert-dwelling nomadic groups that have depended upon milk for millennia. Three quarters of the adult Tutsi of Rwanda and Burundi have a high ability to digest lactose, while only 5% of the adults of the neighboring Shi people of eastern Congo can. Among Hutu, one in three
Tutsi after mid-1600. Beginning in about 1880, Roman Catholic missionaries arrived in the Great Lakes region. Later, when German forces occupied the area during World War I, the conflict and efforts for Catholic conversion became more pronounced. As the Tutsi resisted conversion, missionaries found success only among the Hutu. In an effort to reward conversion, the colonial government confiscated traditionally Tutsi land and reassigned it to Hutu tribes. In Burundi, meanwhile, Tutsi domination was even more entrenched. A ruling faction, the Ganwa, soon emerged from amongst the Tutsi and assumed effective control of the country's administration. The area was ruled as