Origins of Hutu, Tutsi and Twa adults has a high capacity for lactose digestion, a surprisingly high number for an agrarian people, which Mamdani suggests may be the result of centuries of intermarriage with Tutsi. Bethwell Ogot in the 1988 UNESCO "General History" further notes that the number of pastoralists in Rwanda increased sharply around the fifteenth century. Although Luis et al. (2004) in a more general study on bi-allelic markers in many African countries found a statistically significant genetic difference between Tutsi and Hutu, the overall difference were not large. Modern-day genetic studies of the Y-chromosome generally indicate that the Tutsi, like the Hutu, are
Rwandan genocide or simply having a "Tutsi appearance". The RPF was making slow but steady gains in the north and east of the country, ending the killings in each area occupied. The genocide was effectively ended during April in areas of Ruhengeri, Byumba, Kibungo and Kigali provinces. The killings ceased during April in the "akazu" heartlands of western Ruhengeri and Gisenyi, as almost every Tutsi had been eliminated. Large numbers of Hutu in the RPF conquered areas fled, fearing retribution for the genocide; 500,000 Kibungo residents walked over the bridge at Rusumo Falls, into Tanzania, in a few days at the end