Pennaceous feather Pennaceous feather The pennaceous feather is a type of feather present in most modern birds and in some other species of maniraptoriform dinosaurs. A pennaceous feather has a stalk or quill. Its basal part, called a "calamus", is embedded in the skin. The calamus is hollow and has pith formed from the dry remains of the feather pulp, and the calamus opens below by an "inferior umbilicus" and above by a "superior umbilicus". The stalk above the calamus is a solid "rachis" having an umbilical groove on its underside. Pennaceous feathers have a rachis with "vanes" or "vaxillum " spreading
Calamus (DTP) for its support for automatic kerning even where adjacent characters are set in different fonts or at different sizes. Its high modularity offers features for almost every purpose in desktop publishing. Calamus also was one of the first DTP apps to support real virtual objects and frames, nondestructive vector masks, and editable PS/PDF import. Its (adjustable) measurement base of 1/10,000mm allows accurate positioning of elements. Calamus was ported to Windows by MGI Software and was released as Calamus 95. The current version of Calamus is Calamus SL 2015, also available as SLC 2015 (complete edition with all additional modules) and