Feathered dinosaur the deciduous sheath, forming a tuft of unbranched barbs with a basal calamus. Stage III consists of two developmental novelties, IIIa and IIIb, as either could have occurred first. Stage IIIa involves helical displacement of barb ridges arising within the collar. The barb ridges on the anterior midline of the follicle fuse together, forming the rachis. The creation of a posterior barb locus follows, giving an indeterminate number of barbs. This resulted in a feather with a symmetrical, primarily branched structure with a rachis and unbranched barbs. In stage IIIb, barbules paired within the peripheral barbule plates of the barb
Calamus, Iowa a more favorable location in what is now the town of Calamus. The present day location of Calamus, a small village on the Union Pacific Railway, was platted in 1860 by R. S. Dickinson, who owned the land on the north side of the railroad, and Col. Milo Smith, who owned that on the south. James Keith opened the first store here containing a small stock of groceries and accessories, located about where the present day Faith Lutheran Church stands. In 1861, R.S. Dickinson and his son A.L. built the first general store, and opened a large and complete stock