Origins of Hutu, Tutsi and Twa adults has a high capacity for lactose digestion, a surprisingly high number for an agrarian people, which Mamdani suggests may be the result of centuries of intermarriage with Tutsi. Bethwell Ogot in the 1988 UNESCO "General History" further notes that the number of pastoralists in Rwanda increased sharply around the fifteenth century. Although Luis et al. (2004) in a more general study on bi-allelic markers in many African countries found a statistically significant genetic difference between Tutsi and Hutu, the overall difference were not large. Modern-day genetic studies of the Y-chromosome generally indicate that the Tutsi, like the Hutu, are
History of Rwanda RPF control was occupied by France in Operation Turquoise. While the French operation did prevent mass killings it has been alleged that the deployment of French troops was intended to allow the Hutu militias to escape, and that the slaughter of Tutsis continued in the French controlled area. Between July and August 1994, Kagame's Tutsi-led RPF troops first entered Kigali and soon thereafter captured the rest of the country. The Tutsi rebels defeated the Hutu regime and ended the genocide, but approximately two million Hutu refugees - some who participated in the genocide and fearing Tutsi retribution - fled to