Origins of Hutu, Tutsi and Twa adults has a high capacity for lactose digestion, a surprisingly high number for an agrarian people, which Mamdani suggests may be the result of centuries of intermarriage with Tutsi. Bethwell Ogot in the 1988 UNESCO "General History" further notes that the number of pastoralists in Rwanda increased sharply around the fifteenth century. Although Luis et al. (2004) in a more general study on bi-allelic markers in many African countries found a statistically significant genetic difference between Tutsi and Hutu, the overall difference were not large. Modern-day genetic studies of the Y-chromosome generally indicate that the Tutsi, like the Hutu, are
Hutu of Rwandan national unity, but has been criticized as an attempt at historical revisionism. Still others suggest that the two groups are related but not identical, and that differences between them were exacerbated by Europeans, or by a gradual, natural split, as those who owned cattle became known as Tutsi and those who did not became Hutu. Mahmood Mamdani states that the Belgian colonial power designated people as Tutsi or Hutu on the basis of cattle ownership, physical measurements and church records. There is an ongoing debate as to whether the Hutu and Tutsi are really separate groups or not;