Hutu as was the person constitutionally entitled to succeed him. This sparked a genocide in Burundi between Hutu political structures and the Tutsi military, in which an estimated 500,000 Burundians died. There were many mass killings of Tutsis and moderate Hutus; these events were deemed genocide by the United Nations International Commission of Inquiry for Burundi. While Tutsi remained in control of Burundi, the conflict resulted in genocide in Rwanda as well. A Tutsi rebel group, the Rwandan Patriotic Front, invaded Rwanda from Uganda, which started a civil war against Rwanda's Hutu government in 1990. A peace agreement was signed, but
Origins of Hutu, Tutsi and Twa communities (17.7% Afro-Asiatic genes found in the mixed Hutu/Tutsi population). While most supporters of the migration theory are also supporters of the "Hamitic theory", namely that the Tutsi came from the Horn of Africa, a later theory proposed that the Tutsi had instead migrated from nearby interior East Africa, and that the physical differences were the result of natural selection in a dry arid climate over millennia. Among the most detailed theories was one put forward by Jean Hiernaux, based on studies of blood factors and archeology. Noting the fossil record of a tall people with narrow facial features several