Origins of Hutu, Tutsi and Twa on economic criteria. Formal and discrete social divisions were consequently imposed upon ambiguous biological distinctions. To some extent, the permeability of these categories in the intervening decades helped to reify the biological distinctions, generating a taller elite and a shorter underclass, but with little relation to the gene pools that had existed a few centuries ago. The social categories are thus real, but there is little if any detectable genetic differentiation between Hutu and Tutsi. Tishkoff et al. (2009) found their mixed Hutu and Tutsi samples from Rwanda to be predominately of Bantu origin, with minor gene flow from Afro-Asiatic
History of Rwanda RPF control was occupied by France in Operation Turquoise. While the French operation did prevent mass killings it has been alleged that the deployment of French troops was intended to allow the Hutu militias to escape, and that the slaughter of Tutsis continued in the French controlled area. Between July and August 1994, Kagame's Tutsi-led RPF troops first entered Kigali and soon thereafter captured the rest of the country. The Tutsi rebels defeated the Hutu regime and ended the genocide, but approximately two million Hutu refugees - some who participated in the genocide and fearing Tutsi retribution - fled to