Origins of Hutu, Tutsi and Twa phenotype, genotype, cultural memory of inhabitants of Rwanda, archeology, and linguistics. Most Tutsis and Hutus carry the E1b1a paternal haplogroup, which is common among Bantu populations. More recent studies have de-emphasized physical appearance, such as height and nose width, in favor of examining blood factors, the presence of the sickle cell trait, lactose intolerance in adults, and other genotype expressions. Excoffier et al. (1987) found that the Tutsi and Hima, despite being surrounded by Bantu populations, are "closer genetically to Cushites and Ethiosemites". Another study concluded that, while the sickle cell trait among the Rwandan Hutu was comparable to that
Hutu of Rwandan national unity, but has been criticized as an attempt at historical revisionism. Still others suggest that the two groups are related but not identical, and that differences between them were exacerbated by Europeans, or by a gradual, natural split, as those who owned cattle became known as Tutsi and those who did not became Hutu. Mahmood Mamdani states that the Belgian colonial power designated people as Tutsi or Hutu on the basis of cattle ownership, physical measurements and church records. There is an ongoing debate as to whether the Hutu and Tutsi are really separate groups or not;