Rwandan genocide training to the Banyamulenge, a Tutsi group in the Zairian South Kivu province, helping them to defeat Zairian security forces. Rwandan forces, the Banyamulenge, and other Zairian Tutsi, then attacked the refugee camps, targeting the Hutu militia. These attacks caused hundreds of thousands of refugees to flee; many returned to Rwanda despite the presence of the RPF, while others ventured further west into Zaire. The defeated forces of the former regime continued a cross-border insurgency campaign, supported initially by the predominantly Hutu population of Rwanda's northwestern provinces. By 1999, a programme of propaganda and Hutu integration into the national army,
Origins of Hutu, Tutsi and Twa largely of Bantu extraction (60% E1b1a, 20% B, 4% E3). Paternal genetic influences associated with the Horn of Africa and North Africa are few (under 3% E1b1b), and are ascribed to much earlier inhabitants who were assimilated. However, the Tutsi have considerably more haplogroup B paternal lineages (14.9% B) than do the Hutu (4.3% B). Trombetta et al. (2015) found 22.2% of E1b1b in a small sample of Tutsis from Burundi, but no bearers of the haplogroup among the local Hutu and Twa populations. The subclade was of the M293 variety, which suggests that the ancestors of Tutsis in this