Origins of Hutu, Tutsi and Twa adults has a high capacity for lactose digestion, a surprisingly high number for an agrarian people, which Mamdani suggests may be the result of centuries of intermarriage with Tutsi. Bethwell Ogot in the 1988 UNESCO "General History" further notes that the number of pastoralists in Rwanda increased sharply around the fifteenth century. Although Luis et al. (2004) in a more general study on bi-allelic markers in many African countries found a statistically significant genetic difference between Tutsi and Hutu, the overall difference were not large. Modern-day genetic studies of the Y-chromosome generally indicate that the Tutsi, like the Hutu, are
Hotel Rwanda President Juvénal Habyarimana, a Hutu, was shot down. Following that incident, the genocide began. Hutu extremists belonging to the Interahamwe militia launched plans to destroy the entire Tutsi civilian population. Tutsi and people suspected of being Tutsi were killed in their homes and as they tried to flee the country. It is estimated that some 200,000 people participated in the perpetration of the Rwandan genocide. Hotelier Paul Rusesabagina of the Belgian owned luxury Hôtel des Mille Collines, used his power and influence to personally save both Tutsi and Hutu refugees. Rusesabagina regularly bribed Rwandan Hutu soldiers and kept militias outside