Origins of Hutu, Tutsi and Twa phenotype, genotype, cultural memory of inhabitants of Rwanda, archeology, and linguistics. Most Tutsis and Hutus carry the E1b1a paternal haplogroup, which is common among Bantu populations. More recent studies have de-emphasized physical appearance, such as height and nose width, in favor of examining blood factors, the presence of the sickle cell trait, lactose intolerance in adults, and other genotype expressions. Excoffier et al. (1987) found that the Tutsi and Hima, despite being surrounded by Bantu populations, are "closer genetically to Cushites and Ethiosemites". Another study concluded that, while the sickle cell trait among the Rwandan Hutu was comparable to that
Rape during the Rwandan genocide during the genocide. Although Tutsi women were the main targets, moderate Hutu women were also raped during the genocide. Along with the Hutu moderates, Hutu women who were married to Tutsis and Hutu women who hid Tutsis were targeted. In a testimonial, Maria Louise Niyobuhungiro recalls seeing local people and Hutu men watched her get raped up to five times a day, and that when she was kept under watch by a woman, she received neither sympathy nor help and was also forced to farm between rapes. Tutsi women were also targeted with the intent of destroying their reproductive capabilities.