Rwandan Revolution posed by the refugees until the 1990s, when a civil war initiated by the Tutsi-refugee "Front Patriotique Rwandais" (FPR) forced the Hutu government into negotiations. This led to a rise in Hutu extremism and the 1994 genocide, in which hundreds of thousands of Tutsi were killed before the FPR took control. The earliest inhabitants of what is now Rwanda were the Twa, a group of aboriginal pygmy hunter-gatherers who settled the area between 8000 and 3000 BC and remain in the country today. Between 700 BC and 1500 AD, a number of Bantu groups migrated into Rwanda and began clearing
Origins of Hutu, Tutsi and Twa largely of Bantu extraction (60% E1b1a, 20% B, 4% E3). Paternal genetic influences associated with the Horn of Africa and North Africa are few (under 3% E1b1b), and are ascribed to much earlier inhabitants who were assimilated. However, the Tutsi have considerably more haplogroup B paternal lineages (14.9% B) than do the Hutu (4.3% B). Trombetta et al. (2015) found 22.2% of E1b1b in a small sample of Tutsis from Burundi, but no bearers of the haplogroup among the local Hutu and Twa populations. The subclade was of the M293 variety, which suggests that the ancestors of Tutsis in this