Origins of Hutu, Tutsi and Twa phenotype, genotype, cultural memory of inhabitants of Rwanda, archeology, and linguistics. Most Tutsis and Hutus carry the E1b1a paternal haplogroup, which is common among Bantu populations. More recent studies have de-emphasized physical appearance, such as height and nose width, in favor of examining blood factors, the presence of the sickle cell trait, lactose intolerance in adults, and other genotype expressions. Excoffier et al. (1987) found that the Tutsi and Hima, despite being surrounded by Bantu populations, are "closer genetically to Cushites and Ethiosemites". Another study concluded that, while the sickle cell trait among the Rwandan Hutu was comparable to that
Genocides in central Africa Ituri conflict and the mass murder of Pygmies known as "Effacer le tableau". In Uganda, 300 thousand people were murdered during the regime of Idi Amin and 500 thousand during the rule of his successor, Milton Obote. Amin's genocides targeted the Acholi and Lango peoples; these two groups went on to murder other groups (mainly the Baganda) under Obote's regime. In the early 1970s, over 150 thousand Hutu people were killed by Tutsi people in Burundi by order of General Michel Micombero. Twenty years later, one million Tutsi people were murdered by Hutu people during the 1994 Rwandan genocide. In