Origins of Hutu, Tutsi and Twa on economic criteria. Formal and discrete social divisions were consequently imposed upon ambiguous biological distinctions. To some extent, the permeability of these categories in the intervening decades helped to reify the biological distinctions, generating a taller elite and a shorter underclass, but with little relation to the gene pools that had existed a few centuries ago. The social categories are thus real, but there is little if any detectable genetic differentiation between Hutu and Tutsi. Tishkoff et al. (2009) found their mixed Hutu and Tutsi samples from Rwanda to be predominately of Bantu origin, with minor gene flow from Afro-Asiatic
Rape during the Rwandan genocide during the genocide. Although Tutsi women were the main targets, moderate Hutu women were also raped during the genocide. Along with the Hutu moderates, Hutu women who were married to Tutsis and Hutu women who hid Tutsis were targeted. In a testimonial, Maria Louise Niyobuhungiro recalls seeing local people and Hutu men watched her get raped up to five times a day, and that when she was kept under watch by a woman, she received neither sympathy nor help and was also forced to farm between rapes. Tutsi women were also targeted with the intent of destroying their reproductive capabilities.