Human hair color account for all possible shades of brown, blond, or red (for example, platinum blond versus dark blond/light brown), nor does it explain why hair color sometimes darkens as a person ages. Several gene pairs control the light versus dark hair color in a cumulative effect. A person's genotype for a multifactorial trait can interact with the environment to produce varying phenotypes (see quantitative trait locus). Natural hair color can be brown, blond, black, red, or white; all natural hair colors including grey, white and lightest blond, are shades of brown. The Fischer–Saller scale, named after Eugen Fischer and , is
George R. Fischer and after his retirement. He died in Tallahassee, Florida, on May 29, 2016. He is survived by his wife and son. Fischer attended Stanford University, performing his Undergraduate work from 1955 to 1960 and earning his Bachelor of Arts in Anthropology, with minor concentrations in English and Geology. He continued with graduate school at Stanford from 1960–62 and has completed all requirements for his Master of Arts in Anthropology except thesis. Fischer dabbled part-time as a special student at Florida State University from 1972 to 1973 while employed at the Southeast Archaeological Center. George Fischer was employed with the National