Pennaceous feather Pennaceous feather The pennaceous feather is a type of feather present in most modern birds and in some other species of maniraptoriform dinosaurs. A pennaceous feather has a stalk or quill. Its basal part, called a "calamus", is embedded in the skin. The calamus is hollow and has pith formed from the dry remains of the feather pulp, and the calamus opens below by an "inferior umbilicus" and above by a "superior umbilicus". The stalk above the calamus is a solid "rachis" having an umbilical groove on its underside. Pennaceous feathers have a rachis with "vanes" or "vaxillum " spreading
What Bird is That? author Nick Earls' 1999 young adult novel "48 Shades of Brown". The 48 shades refers to the number of distinct browns used by Cayley to describe bird plumage colours. What Bird is That? What Bird is That? A Guide to the Birds of Australia is a book first published in 1931 by Angus & Robertson in Sydney. Authored and illustrated by Neville William Cayley, it was Australia’s first fully illustrated national field guide to birds, a function it served alone for nearly 40 years. In 1960 it was rated the all-time best seller in Australian natural history. "What Bird Is