Feather These barbules have minute hooks called barbicels for cross-attachment. Down feathers are fluffy because they lack barbicels, so the barbules float free of each other, allowing the down to trap air and provide excellent thermal insulation. At the base of the feather, the rachis expands to form the hollow tubular "calamus" (or quill) which inserts into a follicle in the skin. The basal part of the calamus is without vanes. This part is embedded within the skin follicle and has an opening at the base (proximal umbilicus) and a small opening on the side (distal umbilicus). Hatchling birds of some
Acorus calamus asexually. The tetraploid variety is usually known as "Acorus calamus var. angustatus" Besser. A number of synonyms are known, but a number are contested as to which variety they belong. It is morphologically diverse, with some forms having very broad and some narrow leaves. It is furthermore also cytotypically diverse, with an array of different karyotypes. "A. calamus" has been an item of trade in many cultures for thousands of years. It has been used medicinally for a wide variety of ailments, and its aroma makes calamus essential oil valued in the perfume industry. The essence from the rhizome is