Modern synthesis (20th century) a gene-centred view of evolution following Hamilton's concepts, disputing the idea of evolutionary progress, and attacking the then widespread theory of group selection. Williams argued that natural selection worked by changing the frequency of alleles, and could not work at the level of groups. Gene-centred evolution was popularised by Richard Dawkins in his 1976 book "The Selfish Gene" and developed in his more technical writings. In 1975, E. O. Wilson published his controversial book "", the subtitle alluding to the modern synthesis as he attempted to bring the study of animal society into the evolutionary fold. This appeared radically new,
On the Tendency of Species to form Varieties; and on the Perpetuation of Varieties and Species by Natural Means of Selection Darwin apparently mistook Wallace's meaning, writing "nothing very new... Uses my simile of tree, [but] it seems all creation with him". However, he spelt out the details of Natural Selection to Lyell, who found the idea hard to accept but urged Darwin to publish to establish priority. On 14 May 1856 Darwin began what became his draft for a book titled "Natural Selection". Wallace collected specimens and corresponded with Darwin from Borneo. In December 1857, he wrote to ask if Darwin's book would delve into human origins, to which Darwin responded that "I think I shall avoid the whole subject,