Plant tolerance to herbivory affect plant growth and also tolerance are water, light, carbon dioxide and soil nutrients. Water and light levels are generally assumed to be positively associated with tolerance (Strauss and Agrawal 1999). However, there are exceptions such as evidence of decreased tolerance in "Madia sativa" with increased water availability (Wise and Abrahamson 2007, Gonzales "et al." 2008). Many studies have found CO levels to decrease tolerance in plants (Lau and Tiffin 2009). Increased nutrient levels are also commonly found to be negatively associated with tolerance (Wise and Abrahamson 2007). There are currently three prominent models that predict how resource levels may
Plant tolerance to herbivory Using simulated herbivory allows for the control of environmental variables, but replicating natural herbivory is difficult, causing plants to respond differently from imposed and natural herbivory (Tiffin and Inouye 2000). Growing plants in the control environment of the greenhouse may also affect their response as it is still a novel environment to the plants. Even if the plots are grown in natural settings, the methods of excluding or including herbivores, such as using cages or pesticides, may also affect plant tolerance (Tiffin and Inouye 2000). Lastly, models have predicted that manipulated herbivory may actually result in less precise estimates of