Pleiotropy correlations and responses to selection most often exemplify pleiotropy. Pleiotropic traits had been previously recognized in the scientific community but had not been experimented on until Gregor Mendel's 1866 pea plant experiment. Mendel recognized that certain pea plant traits (seed coat color, flower color, and axial spots) seemed to be inherited together; however, their correlation to a single gene has never been proven. The term "pleiotropie" was first coined by Ludwig Plate in his Festschrift, which was published in 1910. He originally defined pleiotropy as occurring when "several characteristics are dependent upon ... [inheritance]; these characteristics will then always appear
Catholic Church and evolution development of evolutionary theory included those of the Jesuit-educated French scientist Jean-Baptiste Lamarck (1744-1829) and of the Augustinian monk Gregor Mendel (1822-1884). Lamarck developed Lamarckism, the first coherent theory of evolution, proposing in "Philosophie Zoologique" (1809) and other works his theory of the transmutation of species. He constructed a genealogical tree to show the genetic connection of organisms. Mendel entered the Brno Augustinian monastery in 1843, but also trained as a scientist at the Olmutz Philosophical Institute and at the University of Vienna. The Brno monastery was a centre of scholarship, with an extensive library and tradition of scientific research.