Gene-centered view of evolution Richard Dawkins and George C. Williams, other biologists and philosophers have expanded and refined the selfish-gene theory, such as John Maynard Smith, George R. Price, Robert Trivers, David Haig, Helena Cronin, David Hull, Philip Kitcher, and Daniel C. Dennett. The gene-centric view has been opposed by Ernst Mayr, Stephen Jay Gould, David Sloan Wilson, and philosopher Elliott Sober. An alternative, multilevel selection (MLS), has been advocated by E. O. Wilson, David Sloan Wilson, Sober, Richard E. Michod, and Samir Okasha. Writing in the "New York Review of Books", Gould has characterized the gene-centered perspective as confusing book-keeping with causality. Gould
Alternatives to evolution by natural selection 1969 paper by Jack Lester King and Thomas H. Jukes, provocatively but misleadingly titled "Non-Darwinian Evolution". It provided a wide variety of evidence including protein sequence comparisons, studies of the Treffers mutator gene in "E. coli", analysis of the genetic code, and comparative immunology, to argue that most protein evolution is due to neutral mutations and genetic drift. According to Kimura, the theory applies only for evolution at the molecular level, while phenotypic evolution is controlled by natural selection, so the neutral theory does not constitute a true alternative. The various alternatives to Darwinian evolution by natural selection were not