Gene-centered view of evolution Richard Dawkins and George C. Williams, other biologists and philosophers have expanded and refined the selfish-gene theory, such as John Maynard Smith, George R. Price, Robert Trivers, David Haig, Helena Cronin, David Hull, Philip Kitcher, and Daniel C. Dennett. The gene-centric view has been opposed by Ernst Mayr, Stephen Jay Gould, David Sloan Wilson, and philosopher Elliott Sober. An alternative, multilevel selection (MLS), has been advocated by E. O. Wilson, David Sloan Wilson, Sober, Richard E. Michod, and Samir Okasha. Writing in the "New York Review of Books", Gould has characterized the gene-centered perspective as confusing book-keeping with causality. Gould
Natural selection 1900. With the early 20th century integration of evolution with Mendel's laws of inheritance, the so-called modern synthesis, scientists generally came to accept natural selection. The synthesis grew from advances in different fields. Ronald Fisher developed the required mathematical language and wrote "The Genetical Theory of Natural Selection" (1930). J. B. S. Haldane introduced the concept of the "cost" of natural selection. Sewall Wright elucidated the nature of selection and adaptation. In his book "Genetics and the Origin of Species" (1937), Theodosius Dobzhansky established the idea that mutation, once seen as a rival to selection, actually supplied the raw material