Forward genetics a bare minimum for the appearance of specific phenotypes. However, such initial screens were either incomplete as they were missing redundant loci and epigenetic effects, and such screens were difficult to undertake for certain phenotypes that lack directly measurable phenotypes. Additionally a classical genetics approach takes significantly longer. Gregor Mendel experimented with pea plant phenotypes and published his conclusions about genes and inheritance in 1865. Around the early 1900s Thomas Hunt Morgan was mutating Drosophila using radium and attempting to find heritable mutations Alfred Sturtevant later began mapping genes of Drosophila with mutations they had been following. In the 1990s
Plant genetics an English translation. Mendel died in 1884. The significance of Mendel's work was not recognized until the turn of the 20th century. Its rediscovery prompted the foundation of modern genetics. His discoveries, deduction of segregation ratios, and subsequent laws have not only been used in research to gain a better understanding of plant genetics, but also play a large role in plant breeding. Mendel's works along with the works of Charles Darwin and Alfred Wallace on selection provided the basis for much of genetics as a discipline. In the early 1900s, botanists and statisticians began to examine the segregation ratios