Nutrient cycling in the Columbia River Basin eutrophication in part due to the use of the river and its tributaries to irrigate over seven million acres of farmland, an extraction that comprises an average of 93 percent of daily water use in the basin. Natural run-off caused by precipitation and snowpack melt across agricultural lands further transports more of these nutrients into the river. In the late 1990s, the nitrogen load in the Columbia river was over twice the concentration historically observed. Forests are important terrestrial sinks for nutrients due to the ability of vegetation to sequester excess nutrients into their biomass. Additionally, plant roots stabilize soils
Plant to plant communication via mycorrhizal networks be related; therefore, kin selection might improve inclusive fitness and explain why a plant might support a fungus that helps other plants to acquire nutrients. Receipt of defensive signals or cues from an infested plant would be adaptive, as the receiving plant would be able to prime its own defenses in advance of an attack by herbivores. Allelopathic chemicals transferred via CMNs could also affect which plants are selected for survival by limiting the growth of competitors through a reduction of their access to nutrients and light. Therefore, transfer of the different classes of infochemicals might prove adaptive for plants.