Modern synthesis (20th century) a gene-centred view of evolution following Hamilton's concepts, disputing the idea of evolutionary progress, and attacking the then widespread theory of group selection. Williams argued that natural selection worked by changing the frequency of alleles, and could not work at the level of groups. Gene-centred evolution was popularised by Richard Dawkins in his 1976 book "The Selfish Gene" and developed in his more technical writings. In 1975, E. O. Wilson published his controversial book "", the subtitle alluding to the modern synthesis as he attempted to bring the study of animal society into the evolutionary fold. This appeared radically new,
Natural selection 1900. With the early 20th century integration of evolution with Mendel's laws of inheritance, the so-called modern synthesis, scientists generally came to accept natural selection. The synthesis grew from advances in different fields. Ronald Fisher developed the required mathematical language and wrote "The Genetical Theory of Natural Selection" (1930). J. B. S. Haldane introduced the concept of the "cost" of natural selection. Sewall Wright elucidated the nature of selection and adaptation. In his book "Genetics and the Origin of Species" (1937), Theodosius Dobzhansky established the idea that mutation, once seen as a rival to selection, actually supplied the raw material