Modern synthesis (20th century) a gene-centred view of evolution following Hamilton's concepts, disputing the idea of evolutionary progress, and attacking the then widespread theory of group selection. Williams argued that natural selection worked by changing the frequency of alleles, and could not work at the level of groups. Gene-centred evolution was popularised by Richard Dawkins in his 1976 book "The Selfish Gene" and developed in his more technical writings. In 1975, E. O. Wilson published his controversial book "", the subtitle alluding to the modern synthesis as he attempted to bring the study of animal society into the evolutionary fold. This appeared radically new,
The Selfish Genius evolutionary theory that portrays natural selection as acting at the level of the individual gene to the exclusion of group selection models which state that it could also act at the level of organisms or species. The book also claims Dawkins embraces an outdated and prescriptive conception of evolution that actually restricts debate rather than promoting it. The book has provoked controversy with some online commentary claiming it is another 'flea', a term used by Dawkins to describe people who have published books about him or his ideas. An article by Fern Elsdon-Baker in "New Scientist" of 19 July 2009