Nutrient cycling in the Columbia River Basin eutrophication in part due to the use of the river and its tributaries to irrigate over seven million acres of farmland, an extraction that comprises an average of 93 percent of daily water use in the basin. Natural run-off caused by precipitation and snowpack melt across agricultural lands further transports more of these nutrients into the river. In the late 1990s, the nitrogen load in the Columbia river was over twice the concentration historically observed. Forests are important terrestrial sinks for nutrients due to the ability of vegetation to sequester excess nutrients into their biomass. Additionally, plant roots stabilize soils
Plant to plant communication via mycorrhizal networks have also initiated nutrient transfer by causing the movement of carbon from the roots of the stressed plant to the roots of a conspecific plant over a mycorrhizal network. Thus, nutrients transferred through mychorrhizal networks act as signals and cues to change the behavior of the connected plants. It is hypothesized that fitness is improved by the transfer of infochemicals through common mycorrhizal networks, as these signals and cues can induce responses which can help the receiver survive in its environment. Plants and fungus have evolved heritable genetic traits which influence their interactions with each other, and experiments, such as