Group selection accustomed to group selection thinking that he did not realize his views "contravened orthodox Darwinian theory". The ethologist Niko Tinbergen praised Lorenz for his interest in the survival value of behavior, and naturalists enjoyed Lorenz's writings for the same reason. In 1962, group selection was used as a popular explanation for adaptation by the zoologist V. C. Wynne-Edwards. In 1976, Richard Dawkins wrote a well-known book on the importance of evolution at the level of the gene or the individual, "The Selfish Gene". From the mid 1960s, evolutionary biologists argued that natural selection acted primarily at the level of the
The Selfish Gene to behave selflessly with each other. A lineage is expected to evolve to maximise its inclusive fitness—the number of copies of its genes passed on globally (rather than by a particular individual). As a result, populations will tend towards an evolutionarily stable strategy. The book also coins the term "meme" for a unit of human cultural evolution analogous to the gene, suggesting that such "selfish" replication may also model human culture, in a different sense. Memetics has become the subject of many studies since the publication of the book. In raising awareness of Hamilton's ideas, as well as making its