The Selfish Gene was dropped. The book contains no illustrations. The book has been translated into at least 23 languages including Arabic, Thai and Turkish. In 2006, a 30th-anniversary edition was published with the Trivers foreword and a new introduction by the author. It was accompanied by a "festschrift" entitled "" (2006). In March 2006, a special event entitled "The Selfish Gene": Thirty Years On was held at the London School of Economics. In March 2011, Audible Inc published an audiobook edition narrated by Richard Dawkins and Lalla Ward. In 2016, Oxford University Press published a 40th anniversary edition with a new epilogue,
The Selfish Gene and Natural Selection" (1966), which argued that altruism is not based upon group benefit "per se", but is a result of selection that occurs "at the level of the gene mediated by the phenotype" and any selection at the group level occurred only under rare circumstances. This approach was developed further during the 1960s by W. D. Hamilton and others who opposed group selection and selection aimed directly at benefit to the individual organism: Despite the principle of 'survival of the fittest' the ultimate criterion which determines whether [a gene] "G" will spread is not whether the behavior is to