Nutrient cycling in the Columbia River Basin eutrophication in part due to the use of the river and its tributaries to irrigate over seven million acres of farmland, an extraction that comprises an average of 93 percent of daily water use in the basin. Natural run-off caused by precipitation and snowpack melt across agricultural lands further transports more of these nutrients into the river. In the late 1990s, the nitrogen load in the Columbia river was over twice the concentration historically observed. Forests are important terrestrial sinks for nutrients due to the ability of vegetation to sequester excess nutrients into their biomass. Additionally, plant roots stabilize soils
Carnivorous plant more or less than this optimum will take up less carbon dioxide than an optimal plant, and hence growing less well. These plants will be at a selective disadvantage. At zero investment the growth is zero, because a non-carnivorous plant cannot survive in a habitat with absolutely no soil-borne nutrients. Such habitats do not exist, so for example, "Sphagnum" absorbs the tiny amounts of nitrates and phosphates in rain very efficiently and also forms symbioses with diazotrophic cyanobacteria. In a habitat with abundant soil nutrients but little light (as shown above), the gross photosynthesis curve will be lower and flatter,