Nutrient cycling in the Columbia River Basin eutrophication in part due to the use of the river and its tributaries to irrigate over seven million acres of farmland, an extraction that comprises an average of 93 percent of daily water use in the basin. Natural run-off caused by precipitation and snowpack melt across agricultural lands further transports more of these nutrients into the river. In the late 1990s, the nitrogen load in the Columbia river was over twice the concentration historically observed. Forests are important terrestrial sinks for nutrients due to the ability of vegetation to sequester excess nutrients into their biomass. Additionally, plant roots stabilize soils
Plant to plant communication via mycorrhizal networks help plants in mycorrhizal networks defend themselves against attack by pathogens or herbivores, and transferred nutrients can affect growth and nutrition. Results of studies which demonstrate these modes of communication have led the authors to hypothesize mechanisms by which the transfer of these nutrients can affect the fitness of the connected plants. Plants and fungi form mutualistic symbiotic relationships called mycorrhizae, which take several forms, such as arbuscular mycorrhizae (AM) and ectomycorrhizae (ECM), and are widespread in nature. Host plants provide photosynthetically derived carbohydrates to the mycorrhizal fungi, which use them in metabolism, either for energy or to increase the