Papal tiara Papal tiara The papal tiara is a crown that was worn by popes of the Catholic Church from as early as the 8th century to the mid-20th. It was last used by Pope Paul VI in 1963 and only at the beginning of his reign. The name "tiara" refers to the entire headpiece, no matter how many crowns, circlets or diadems have adorned it through the ages, while the three-tiered form that it took in the 14th century is also called the triregnum, triple tiara, or triple crown. From 1143 to 1963, the papal tiara was solemnly placed on the
Plant–fungus horizontal gene transfer ancient, they have been difficult to detect and remain relatively unknown. Plant–fungus interactions could play a part in a multi-horizontal gene transfer pathway among many other organisms. Fungus–plant-mediated horizontal gene transfer can occur via phagotrophic mechanisms (mediated by phagotrophic eukaryotes) and nonphagotropic mechanisms. Nonphagotrophic mechanisms have been seen in the transmission of transposable elements, plastid-derived endosymbiotic gene transfer, prokaryote-derived gene transfer, "Agrobacterium tumefaciens"-mediated DNA transfer, cross-species hybridization events, and gene transfer between mitochondrial genes. Horizontal gene transfer could bypass eukaryotic barrier features like linear chromatin-based chromosomes, intron–exon gene structures, and the nuclear envelope. Horizontal gene transfer occurs between microorganisms sharing