Heredity particulate inheritance of genes can be attributed to the Moravian monk Gregor Mendel who published his work on pea plants in 1865. However, his work was not widely known and was rediscovered in 1901. It was initially assumed that Mendelian inheritance only accounted for large (qualitative) differences, such as those seen by Mendel in his pea plants—and the idea of additive effect of (quantitative) genes was not realised until R. A. Fisher's (1918) paper, "The Correlation Between Relatives on the Supposition of Mendelian Inheritance" Mendel's overall contribution gave scientists a useful overview that traits were inheritable. His pea plant demonstration
Johann Karl Nestler stock animals and their offspring Nestler hoped to explain the (genetic history). Examination of the records of the ancestors of the best breeding animals, called by Nestler "developmental history" () was for him the reverse side of the same coin. According to Wood and Orel the search for rules of heredity in sheep created an atmosphere of enquiry about heredity in general, which had influence on later work of Gregor Johann Mendel, himself a student at the Faculty of Philosophy in 1840–43. Nestler was also a very active member of the "Moravian-Silesian Society for Advancement of Agriculture" (, in which