Heredity particulate inheritance of genes can be attributed to the Moravian monk Gregor Mendel who published his work on pea plants in 1865. However, his work was not widely known and was rediscovered in 1901. It was initially assumed that Mendelian inheritance only accounted for large (qualitative) differences, such as those seen by Mendel in his pea plants—and the idea of additive effect of (quantitative) genes was not realised until R. A. Fisher's (1918) paper, "The Correlation Between Relatives on the Supposition of Mendelian Inheritance" Mendel's overall contribution gave scientists a useful overview that traits were inheritable. His pea plant demonstration
Catholic Church and evolution development of evolutionary theory included those of the Jesuit-educated French scientist Jean-Baptiste Lamarck (1744-1829) and of the Augustinian monk Gregor Mendel (1822-1884). Lamarck developed Lamarckism, the first coherent theory of evolution, proposing in "Philosophie Zoologique" (1809) and other works his theory of the transmutation of species. He constructed a genealogical tree to show the genetic connection of organisms. Mendel entered the Brno Augustinian monastery in 1843, but also trained as a scientist at the Olmutz Philosophical Institute and at the University of Vienna. The Brno monastery was a centre of scholarship, with an extensive library and tradition of scientific research.