Unit of selection and Confirmation of Evolutionary Theory" provides a basic "philosophical" introduction to the debate. Below, cases of selection at the genic, cellular, individual and group level from within the multi-level selection perspective are presented and discussed. George C. Williams in his influential book "Adaptation and Natural Selection" was one of the first to present a gene-centered view of evolution with the gene as the unit of selection, arguing that a unit of selection should exhibit a high degree of permanence. Richard Dawkins has written several books popularizing and expanding the idea. According to Dawkins, genes cause phenotypes and a gene is
The Selfish Gene to behave selflessly with each other. A lineage is expected to evolve to maximise its inclusive fitness—the number of copies of its genes passed on globally (rather than by a particular individual). As a result, populations will tend towards an evolutionarily stable strategy. The book also coins the term "meme" for a unit of human cultural evolution analogous to the gene, suggesting that such "selfish" replication may also model human culture, in a different sense. Memetics has become the subject of many studies since the publication of the book. In raising awareness of Hamilton's ideas, as well as making its