Crozier (mycology) Crozier (mycology) A crozier is an anatomical feature of many fungi in the phylum Ascomycota that form at the base of asci and look like hook-topped shepherd’s staffs or stylized religious crosiers. During the ascus initial formation the crozier helps to maintain a dikaryotic state in the ascus initial and its side branch that will continue the spreading growth of the ascogenous hyphae in Ascomycota fruitbodies. The tips of developing asci on these ascogenous hyphae curl over. One haploid nucleus migrates into the curved tip while the other compatible haploid nucleus remains in the penultimate space below the hook. The
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