Imre Festetics Festetics’s observations highlighted important correlations between variability, adaptation, and development. He also noted the consequences of selection and its role in heredity, believing that variability and his postulated laws of genetics were connected, acting together in breeding as well as in the natural processes controlling populations of different animals,including humans. Festetics, however, was ultimately hindered by the complex nature of his study traits, aspects of wool quality that we now know to be polygenic. When Gregor Mendel turned his attention to inheritance in peas he the latest in a line of Moravian researchers and agriculturalists who had been thinking about
Plant genetics put forth by Mendel. W.E. Castle discovered that while individual traits may segregate and change over time with selection, that when selection is stopped and environmental effects are taken into account, the genetic ratio stops changing and reach a sort of stasis, the foundation of Population Genetics. This was independently discovered by G.H. Hardy and W. Weinberg, which ultimately gave rise to the concept of Hardy-Weinberg equilibrium published in 1908. For a more thorough exploration of the history of population genetics, see History of Population Genetics by Bob Allard. Around this same time, genetic and plant breeding experiments in maize