Group selection accustomed to group selection thinking that he did not realize his views "contravened orthodox Darwinian theory". The ethologist Niko Tinbergen praised Lorenz for his interest in the survival value of behavior, and naturalists enjoyed Lorenz's writings for the same reason. In 1962, group selection was used as a popular explanation for adaptation by the zoologist V. C. Wynne-Edwards. In 1976, Richard Dawkins wrote a well-known book on the importance of evolution at the level of the gene or the individual, "The Selfish Gene". From the mid 1960s, evolutionary biologists argued that natural selection acted primarily at the level of the
The Genetical Theory of Natural Selection chapter, Fisher considers the nature of inheritance, rejecting blending inheritance, because it would eliminate genetic variance, in favour of particulate inheritance. The second chapter introduces Fisher's fundamental theorem of natural selection. The third considers the evolution of dominance, which Fisher believed was strongly influenced by modifiers. Other chapters discuss parental investment, Fisher's geometric model, concerning how spontaneous mutations affect biological fitness, Fisher's principle which explains why the sex ratio between males and females is almost always 1:1, reproductive value, examining the demography of having girl children. Using his knowledge of statistics, the Fisherian runaway, which explores how sexual selection can