The Evolution of Cooperation insects, where the workers – which are sterile, and therefore incapable of passing on their genes – benefit the queen, who is essentially passing on copies of "their" genes. This is further elaborated in the "selfish gene" theory of Richard Dawkins, that the unit of evolution is not the individual organism, but the gene. (As stated by Wilson: "the organism is only DNA's way of making more DNA.") However, kinship selection works only where the individuals involved are closely related; it fails to explain the presence of altruism and cooperation between unrelated individuals, particularly across species. In a 1971 paper
On the Tendency of Species to form Varieties; and on the Perpetuation of Varieties and Species by Natural Means of Selection Letter" from Darwin to Asa Gray. This was the first announcement of the Darwin – Wallace theory of evolution by natural selection; the papers appeared in print on 20 August 1858. The presentation of the papers spurred Darwin to write a condensed "abstract" of his "big book" on "Natural Selection". This was published in November 1859 as "On the Origin of Species". On the voyage of the "Beagle" the young Charles Darwin took a break between graduating and starting his career as a clergyman to study the natural history of South America, an interest he had developed at the University