The Selfish Gene Another criticism of the book is its treatment of morality, and more particularly altruism, as existing only as a form of selfishness: The philosopher Mary Midgley has suggested this position is a variant of Hobbes' explanation of altruism as enlightened self-interest, and that Dawkins goes a step further to suggest that our genetic programming can be overcome by what amounts to an extreme version of free will. Part of Mary Midgley's concern is that Richard Dawkins' account of "The Selfish Gene" serves as a moral and ideological justification for selfishness to be adopted by modern human societies as simply following
The Selfish Gene to behave selflessly with each other. A lineage is expected to evolve to maximise its inclusive fitness—the number of copies of its genes passed on globally (rather than by a particular individual). As a result, populations will tend towards an evolutionarily stable strategy. The book also coins the term "meme" for a unit of human cultural evolution analogous to the gene, suggesting that such "selfish" replication may also model human culture, in a different sense. Memetics has become the subject of many studies since the publication of the book. In raising awareness of Hamilton's ideas, as well as making its