Plant tolerance to herbivory affect plant growth and also tolerance are water, light, carbon dioxide and soil nutrients. Water and light levels are generally assumed to be positively associated with tolerance (Strauss and Agrawal 1999). However, there are exceptions such as evidence of decreased tolerance in "Madia sativa" with increased water availability (Wise and Abrahamson 2007, Gonzales "et al." 2008). Many studies have found CO levels to decrease tolerance in plants (Lau and Tiffin 2009). Increased nutrient levels are also commonly found to be negatively associated with tolerance (Wise and Abrahamson 2007). There are currently three prominent models that predict how resource levels may
Plant defense against herbivory change their defenses in response to changes in nutrients. For example, if plants are grown in low-nitrogen conditions, then these plants will implement a defensive strategy composed of constitutive carbon-based defenses. If nutrient levels subsequently increase, by for example the addition of fertilizers, these carbon-based defenses will decrease. The growth rate hypothesis, also known as the "resource availability hypothesis", states that defense strategies are determined by the inherent growth rate of the plant, which is in turn determined by the resources available to the plant. A major assumption is that available resources are the limiting factor in determining the maximum