Modern synthesis (20th century) a gene-centred view of evolution following Hamilton's concepts, disputing the idea of evolutionary progress, and attacking the then widespread theory of group selection. Williams argued that natural selection worked by changing the frequency of alleles, and could not work at the level of groups. Gene-centred evolution was popularised by Richard Dawkins in his 1976 book "The Selfish Gene" and developed in his more technical writings. In 1975, E. O. Wilson published his controversial book "", the subtitle alluding to the modern synthesis as he attempted to bring the study of animal society into the evolutionary fold. This appeared radically new,
Gene-centered view of evolution organisms containing the same allele (kin altruism and green-beard effects), or even its own propagation relative to the other genes within the same organism (selfish genes and intragenomic conflict). The gene-centered perspective can be traced to the philosopher Henri Bergson who wrote, in his book, ‘Creative Evolution’ (1907): ‘Life is like a current passing from germ to germ through the medium of a developed organism. It is as if the organism itself were only an excrescence, a bud caused to sprout by the former endeavouring to continue itself in a new germ’. The gene-centered view of evolution is a model