The Evolution of Cooperation insects, where the workers – which are sterile, and therefore incapable of passing on their genes – benefit the queen, who is essentially passing on copies of "their" genes. This is further elaborated in the "selfish gene" theory of Richard Dawkins, that the unit of evolution is not the individual organism, but the gene. (As stated by Wilson: "the organism is only DNA's way of making more DNA.") However, kinship selection works only where the individuals involved are closely related; it fails to explain the presence of altruism and cooperation between unrelated individuals, particularly across species. In a 1971 paper
Gene-centered view of evolution for the evolution of social characteristics such as selfishness and altruism. The formulation of the central dogma of molecular biology was summarized by Maynard Smith: The rejection of the inheritance of acquired characters, combined with Ronald Fisher the statistician, giving the subject a mathematical footing, and showing how Mendelian genetics was compatible with natural selection in his 1930 book "The Genetical Theory of Natural Selection". J. B. S. Haldane, and Sewall Wright, paved the way to the formulation of the selfish-gene theory. For cases where environment can influence heredity, see epigenetics. The view of the gene as the unit of