Plant rapidly, maturing within 110 days, whereas others, in the same environmental conditions, grow more slowly and mature within 155 days. Growth is also determined by environmental factors, such as temperature, available water, available light, carbon dioxide and available nutrients in the soil. Any change in the availability of these external conditions will be reflected in the plant's growth and the timing of its development. Biotic factors also affect plant growth. Plants can be so crowded that no single individual produces normal growth, causing etiolation and chlorosis. Optimal plant growth can be hampered by grazing animals, suboptimal soil composition, lack of
Eutrophication Humankind has increased the rate of phosphorus cycling on Earth by four times, mainly due to agricultural fertilizer production and application. Between 1950 and 1995, an estimated 600,000,000 tonnes of phosphorus was applied to Earth's surface, primarily on croplands. Policy changes to control point sources of phosphorus have resulted in rapid control of eutrophication. Although eutrophication is commonly caused by human activities, it can also be a natural process, particularly in lakes. Eutrophy occurs in many lakes in temperate grasslands, for instance. Paleolimnologists now recognise that climate change, geology, and other external influences are critical in regulating the natural productivity