Biological pigment general, contain six ubiquitous carotenoids: neoxanthin, violaxanthin, antheraxanthin, zeaxanthin, lutein and β-carotene. Lutein is a yellow pigment found in fruits and vegetables and is the most abundant carotenoid in plants. Lycopene is the red pigment responsible for the color of tomatoes. Other less common carotenoids in plants include lutein epoxide (in many woody species), lactucaxanthin (found in lettuce), and alpha carotene (found in carrots). In cyanobacteria, many other carotenoids exist such as canthaxanthin, myxoxanthophyll, synechoxanthin, and echinenone. Algal phototrophs such as dinoflagellates use peridinin as a light harvesting pigment. While carotenoids can be found complexed within chlorophyll-binding proteins such as
Pigment Constantinople. The pigment was expensive and complex to produce, and items colored with it became associated with power and wealth. Greek historian Theopompus, writing in the 4th century BCE, reported that "purple for dyes fetched its weight in silver at Colophon [in Asia Minor]." Mineral pigments were also traded over long distances. The only way to achieve a deep rich blue was by using a semi-precious stone, lapis lazuli, to produce a pigment known as ultramarine, and the best sources of lapis were remote. Flemish painter Jan van Eyck, working in the 15th century, did not ordinarily include blue in