North American beaver large, webbed hind feet. The unwebbed front paws are smaller, with claws. The eyes are covered by a nictitating membrane which allows the beaver to see under water. The nostrils and ears are sealed while submerged. A thick layer of fat under its skin insulates the beaver from its coldwater environment. The beaver's fur consists of long, coarse outer hairs and short, fine inner hairs (see Double coat). The fur has a range of colors, but usually is dark brown. Scent glands near the genitals secrete an oily substance known as castoreum, which the beaver uses to waterproof its fur.
Extracellular polymeric substance Extracellular polymeric substance Extracellular polymeric substances (EPSs) are natural polymers of high molecular weight secreted by microorganisms into their environment. EPSs establish the functional and structural integrity of biofilms, and are considered the fundamental component that determines the physiochemical properties of a biofilm. EPSs are mostly composed of polysaccharides (exopolysaccharides) and proteins, but include other macro-molecules such as DNA, lipids and humic substances. EPSs are the construction material of bacterial settlements and either remain attached to the cell's outer surface, or are secreted into its growth medium. These compounds are important in biofilm formation and cells attachment to surfaces. EPSs