Apatosaurus "Apatosaurus" may have been more solitary than other Morrison Formation dinosaurs. Fossils of the genus have only been found in the upper levels of the formation. Those of "Apatosaurus ajax" are known exclusively from the upper Brushy Basin Member, about 152–151 mya. "A.louisae" fossils are rare, known only from one site in the upper Brushy Basin Member; they date to the late Kimmeridgian stage, about 151mya. Additional "Apatosaurus" remains are known from similarly aged or slightly younger rocks, but they have not been identified as any particular species, and thus may instead belong to "Brontosaurus". The Morrison Formation records a
Apatosaurus have a close counterpart"Brachiosaurus" and "Lusotitan", "Camptosaurus" and "Draconyx", and "Apatosaurus" and "Dinheirosaurus". Other vertebrates that are known to have shared this paleo-environment include ray-finned fishes, frogs, salamanders, turtles, sphenodonts, lizards, terrestrial and aquatic crocodylomorphans, and several species of pterosaur. Shells of bivalves and aquatic snails are also common. The flora of the period has been evidenced in fossils of green algae, fungi, mosses, horsetails, cycads, ginkgoes, and several families of conifers. Vegetation varied from river-lining forests of tree ferns with fern understory (gallery forests), to fern savannas with occasional trees such as the "Araucaria"-like conifer "Brachyphyllum". Apatosaurus Apatosaurus (;