Brenner Pass 1,371 m). Completion of the railway enabled the Austrians to move their troops more efficiently; the Austrians had hoped to secure their territories of Venetia and Lombardy (south of the Alps), but lost them to Italy following the Second Italian War of Independence in 1859 and Austro-Prussian War in 1866. At the end of World War I in 1918, the control of Brenner Pass became shared between Italy and Austria under the Treaty of Saint-Germain-en-Laye (1919). The Treaty of London (1915) secretly awarded Italy the territories south of Brenner Pass for supporting the Entente Powers. Welschtirol/Trentino, along with the southern
Sydney Brenner concept of a messenger RNA, based on correctly interpreting the work of Elliot "Ken" Volkin and Larry Astrachan. Then, with Francis Crick, Leslie Barnett and Richard J. Watts-Tobin, Brenner genetically demonstrated the triplet nature of the code of protein translation through the Crick, Brenner, Barnett, Watts-Tobin et al. experiment of 1961, which discovered frameshift mutations. This insight provided early elucidation of the nature of the genetic code. Leslie Barnett helped set up Sydney Brenner's laboratory in Singapore, many years later. Brenner, with George Pieczenik, created the first computer matrix analysis of nucleic acids using TRAC, which Brenner continues to use.