Cochlea a protein motor called prestin on their outer membranes; it generates additional movement that couples back to the fluid–membrane wave. This "active amplifier" is essential in the ear's ability to amplify weak sounds. The active amplifier also leads to the phenomenon of soundwave vibrations being emitted from the cochlea back into the ear canal through the middle ear (otoacoustic emissions). Otoacoustic emissions are due to a wave exiting the cochlea via the oval window, and propagating back through the middle ear to the eardrum, and out the ear canal, where it can be picked up by a microphone. Otoacoustic emissions
Evolution of the cochlea at the same time as the water-to-land transition of vertebrates, approximately 380 million years ago. The actual coiling or spiral nature of the cochlea occurred to save space inside the skull. The longer the cochlea, the higher is the potential resolution of sound frequencies given the same hearing range. The oldest of the truly coiled mammalian cochleae were approximately 4 mm in length. The earliest evidence available for primates depicts a short cochlea with prominent laminae, suggesting that they had good high-frequency sensitivity as opposed to low-frequency sensitivity. After this, over a period of around 60 million years, evidence suggests