Macaroni penguin been considered separate species, but the close similarities of their DNA sequences has led some, such as the Australian ornithologists Les Christidis and Walter Boles, to treat the royal as a subspecies of the macaroni. The two species are very similar in appearance; the royal penguin has a white face instead of the usually black face of the macaroni. Interbreeding with the Indo-Pacific subspecies of the southern rockhopper penguin ("E. chrysocome filholi") has been reported at Heard and Marion Islands, with three hybrids recorded there by a 1987–88 Australian National Antarctic Research Expedition. The macaroni penguin is a large, crested
Macaroni (fashion) of "the Macaroni Club, which is composed of all the traveled young men who wear long curls and spying-glasses". The "club" was not a formal one; the expression was particularly used to characterize fops who dressed in high fashion with tall, powdered wigs with a chapeau bras on top that could only be removed on the point of a sword. The shop of engravers and printsellers Mary and Matthew Darly in the fashionable West End of London sold their sets of satirical "macaroni" caricature prints, published between 1771 and 1773. The new Darly shop became known as "the Macaroni Print-Shop".