Macaroni penguin female are similar in appearance; males tend to be slightly larger. Males also bear relatively larger bills, which average around compared to in females; this feature has been used to tell the sexes apart. Immature birds are distinguished by their smaller size, smaller, duller-brown bill, dark grey chin and throat, and absent or underdeveloped head plumes, often just a scattering of yellow feathers. The crest is fully developed in birds aged three to four years, a year or two before breeding age. Macaroni penguins moult once a year, a process in which they replace all of their old feathers. They
What Bird is That? Edition What Bird is That?" was published by Australia's Heritage Publishing in 2011, the latest release of Cayley’s “big bird book” complete with Lindsey’s revisions. It is 832 pages, features 769 birds and includes all 460 of Cayley’s full-colour paintings, many showing groups of related birds. New to this edition was an accompanying e-book "What Bird Call is That?", which identifies and illustrates 101 birds from "What Bird is That?", as well as providing sound files of each featured bird’s distinctive call. The sound files were provided by David Stewart. "What Bird is That?" plays a central role in Australian