Macaroni penguin been considered separate species, but the close similarities of their DNA sequences has led some, such as the Australian ornithologists Les Christidis and Walter Boles, to treat the royal as a subspecies of the macaroni. The two species are very similar in appearance; the royal penguin has a white face instead of the usually black face of the macaroni. Interbreeding with the Indo-Pacific subspecies of the southern rockhopper penguin ("E. chrysocome filholi") has been reported at Heard and Marion Islands, with three hybrids recorded there by a 1987–88 Australian National Antarctic Research Expedition. The macaroni penguin is a large, crested
What Bird is That? Edition What Bird is That?" was published by Australia's Heritage Publishing in 2011, the latest release of Cayley’s “big bird book” complete with Lindsey’s revisions. It is 832 pages, features 769 birds and includes all 460 of Cayley’s full-colour paintings, many showing groups of related birds. New to this edition was an accompanying e-book "What Bird Call is That?", which identifies and illustrates 101 birds from "What Bird is That?", as well as providing sound files of each featured bird’s distinctive call. The sound files were provided by David Stewart. "What Bird is That?" plays a central role in Australian