Starfish first appeared in the fossil record in the Cambrian. The first known asterozoans were the Somasteroidea, which exhibit characteristics of both groups. Starfish are infrequently found as fossils, possibly because their hard skeletal components separate as the animal decays. Despite this, there are a few places where accumulations of complete skeletal structures occur, fossilized in place in Lagerstätten — so-called "starfish beds". By the late Paleozoic, the crinoids and blastoids were the predominant echinoderms, and some limestones from this period are made almost entirely from fragments from these groups. In the two major extinction events that occurred during the late
Starfish Invasive Species Specialist Group's list of the world's 100 worst invasive species. Starfish may be preyed on by conspecifics, other starfish species, tritons, crabs, fish, gulls and sea otters. Their first lines of defence are the saponins present in their body walls, which have unpleasant flavours. Some starfish such as "Astropecten polyacanthus" also include powerful toxins such as tetrodotoxin among their chemical armoury, and the slime star can ooze out large quantities of repellent mucus. They also have body armour in the form of hard plates and spines. The crown-of-thorns starfish is particularly unattractive to potential predators, being heavily defended