Animal echolocation in the external ear. There are two hypotheses about the evolution of echolocation in bats. The first suggests that laryngeal echolocation evolved twice in Chiroptera, once in the Yangochiroptera and once in the horseshoe bats (Rhinolophidae). The second proposes that laryngeal echolocation had a single origin in Chiroptera, was subsequently lost in the family Pteropodidae (all megabats), and later evolved as a system of tongue-clicking in the genus "Rousettus". Individual bat species echolocate within specific frequency ranges that suit their environment and prey types. This has sometimes been used by researchers to identify bats flying in an area simply by
Animal echolocation mammals other than bats known to echolocate include two genera ("Sorex" and "Blarina") of shrews and the tenrecs of Madagascar. These include the wandering shrew ("Sorex vagrans"), the common or Eurasian shrew ("Sorex araneus"), and the short-tailed shrew ("Blarina brevicauda"). The nature of shrew sounds unlike those of bats are low amplitude, broadband, multi-harmonic and frequency modulated. They contain no ‘echolocation clicks’ with reverberations and would seem to be used for simple, close range spatial orientation. In contrast to bats, shrews use echolocation only to investigate their habitat rather than additionally to pinpoint food. There is evidence that blinded laboratory