Megabat evolved early in the lineage of chiropterans. Although echolocation was later lost in family Pteropodidae, bats in the genus "Rousettus" are capable of primitive echolocation through clicking their tongues, and some species have been shown to create clicks similar to those of echolocating bats using their wings. Both echolocation and flight are energetically expensive processes. The nature of the flight and echolocation mechanism of bats allows for creation of echolocation pulses with minimal energy use. Energetic coupling of these two processes is thought to have allowed for both energetically expensive processes to evolve in bats. The loss of echolocation may
Barbastelle this world, participated in an evolutionary arms race. The foe for the bat order Chiroptera is its prey, the moth order Lepidoptera. To minimize the risk of the bats' echolocation, moths [and in rarest examples, butterflies] have evolved to detect the echolocation calls of hunting bats, and evoke evasive flight manoeuvres, or reply with their own ultrasonic clicks to confuse the bat's echolocation. The Arctiidae subfamily of Noctuid moths uniquely respond to bat echolocation in three prevailing hypotheses: startle, sonar jamming, and acoustic aposematic defence. All these differences depend on specific environmental settings and the type of echolocation call; however,