Megabat evolved early in the lineage of chiropterans. Although echolocation was later lost in family Pteropodidae, bats in the genus "Rousettus" are capable of primitive echolocation through clicking their tongues, and some species have been shown to create clicks similar to those of echolocating bats using their wings. Both echolocation and flight are energetically expensive processes. The nature of the flight and echolocation mechanism of bats allows for creation of echolocation pulses with minimal energy use. Energetic coupling of these two processes is thought to have allowed for both energetically expensive processes to evolve in bats. The loss of echolocation may
Evolutionary arms race best survival rate. Bats have evolved to use echolocation to detect and catch their prey. Moths have in turn evolved to detect the echolocation calls of hunting bats, and evoke evasive flight maneuvers, or reply with their own ultrasonic clicks to confuse the bat's echolocation. The Arctiidae subfamily of Noctuid moths uniquely respond to bat echolocation in three prevailing hypotheses: startle, sonar jamming, and acoustic aposematic defense. All these differences depend on specific environmental settings and the type of echolocation call; however, these hypotheses are not mutually exclusive and can be used by the same moth for defense. The different