Megabat long, thus smaller than some microbats, for example, the Mauritian tomb bat. The largest attain a wingspan of , and weigh as much as . Most fruit bats have large eyes, allowing them to orient themselves visually in twilight and inside caves and forests. Their sense of smell is excellent. In contrast to the microbats, the fruit bats do not use echolocation (with one exception, the Egyptian fruit bat "Rousettus egyptiacus", which uses high-pitched tongue clicks to navigate in caves). Megabats make up the only family (Pteropodidae) in order Chiroptera that is not capable of laryngeal echolocation. Echolocation and flight
Animal echolocation two cave dwelling bird groups, the so-called cave swiftlets in the genus "Aerodramus" (formerly "Collocalia") and the unrelated Oilbird "Steatornis caripensis". The term "echolocation" was coined by the American zoologist Donald Griffin, whose work with Robert Galambos was the first to convincingly demonstrate its existence in bats in 1938. As Griffin described in his book, the 18th century Italian scientist Lazzaro Spallanzani had, by means of a series of elaborate experiments, concluded that when bats fly at night, they rely on some sense besides vision, but he did not discover that the other sense was hearing. The Swiss physician and