Pollen cell. In flowering plants the vegetative tube cell produces the pollen tube, and the generative cell divides to form the two sperm cells. Pollen is produced in the microsporangia in the male cone of a conifer or other gymnosperm or in the anthers of an angiosperm flower. Pollen grains come in a wide variety of shapes, sizes, and surface markings characteristic of the species (see electron micrograph, right). Pollen grains of pines, firs, and spruces are winged. The smallest pollen grain, that of the forget-me-not ("Myosotis" spp.), is around 6 µm (0.006 mm) in diameter. Wind-borne pollen grains can be
Pollen tube grains are deposited in the micropyle of the ovule of a female cone or megastrobilus, where they mature for up to a year. In conifers and Gnetophytes the pollen germinate to produce a pollen tube that penetrates the megasporangium or nucellus carrying with it sperm nuclei that are transferred to the egg cell in the developing archegonia of the female plant. The female sporophyte must recognize the pollen stuck to the stigma. Often, only pollen of the same species can successfully grow and outcrossed pollen is more successful in growing. With self-incompatibility systems, outcrossed pollen grows and outcompetes self pollen.