Megabat evolved early in the lineage of chiropterans. Although echolocation was later lost in family Pteropodidae, bats in the genus "Rousettus" are capable of primitive echolocation through clicking their tongues, and some species have been shown to create clicks similar to those of echolocating bats using their wings. Both echolocation and flight are energetically expensive processes. The nature of the flight and echolocation mechanism of bats allows for creation of echolocation pulses with minimal energy use. Energetic coupling of these two processes is thought to have allowed for both energetically expensive processes to evolve in bats. The loss of echolocation may
Shrew orientation. In contrast to bats, shrews use echolocation only to investigate their habitats rather than additionally to pinpoint food. Except for large and thus strongly reflecting objects, such as a big stone or tree trunk, they probably are not able to disentangle echo scenes, but rather derive information on habitat type from the overall call reverberations. This might be comparable to human hearing whether one calls into a beech forest or into a reverberant wine cellar. The 385 shrew species are in 26 genera, which are grouped into three living subfamilies: Crocidurinae (white-toothed shrews), Myosoricinae (African shrews) and Soricinae (red-toothed