Megabat long, thus smaller than some microbats, for example, the Mauritian tomb bat. The largest attain a wingspan of , and weigh as much as . Most fruit bats have large eyes, allowing them to orient themselves visually in twilight and inside caves and forests. Their sense of smell is excellent. In contrast to the microbats, the fruit bats do not use echolocation (with one exception, the Egyptian fruit bat "Rousettus egyptiacus", which uses high-pitched tongue clicks to navigate in caves). Megabats make up the only family (Pteropodidae) in order Chiroptera that is not capable of laryngeal echolocation. Echolocation and flight
Waterhouse's leaf-nosed bat happen in that last trimester. Like many bats, this species uses echolocation as a means to locate their prey. In particular, "Macrotus waterhousii" uses a low intensity, broadband-like call to aide in their search for food. The maximum frequency of a call is 73.65 kHz with a minimum of 46.19 kHz, creating a bandwidth of 27.46 kHz. The call ranges from about 1 second to 3 seconds. In addition, to echolocation, "M. waterhousii" uses the sound made by the prey itself to locate it. This type of foraging behavior leads to a preference for a cluttered habitat. As "M. waterhousii"