Extraterrestrial atmosphere star undergoing runaway greenhouse effect. For planets in the habitable zone, it was thought that water photolysis would be strongly limited by cold-trapping of water vapour in the lower atmosphere. However, the extent of HO cold-trapping depends strongly on the amount of non-condensible gases in the atmosphere such as nitrogen N and argon. In the absence of such gases, the likelihood of build-up of oxygen also depends in complex ways on the planet's accretion history, internal chemistry, atmospheric dynamics, and orbital state. Therefore, oxygen, on its own, cannot be considered a robust biosignature. The ratio of nitrogen and argon to
Photorespiration times more abundant, and in solution is 25 times more abundant than . The ability of RuBisCO to specify between the two gases is known as its selectivity factor (or Srel), and it varies between species, with angiosperms more efficient than other plants, but with little variation among the vascular plants. A suggested explanation into RuBisCO's inability to discriminate completely between and is that it is an evolutionary relic: The early atmosphere in which primitive plants originated contained very little oxygen, the early evolution of RuBisCO was not influenced by its ability to discriminate between and . Photorespiration rates are