Animal echolocation in the external ear. There are two hypotheses about the evolution of echolocation in bats. The first suggests that laryngeal echolocation evolved twice in Chiroptera, once in the Yangochiroptera and once in the horseshoe bats (Rhinolophidae). The second proposes that laryngeal echolocation had a single origin in Chiroptera, was subsequently lost in the family Pteropodidae (all megabats), and later evolved as a system of tongue-clicking in the genus "Rousettus". Individual bat species echolocate within specific frequency ranges that suit their environment and prey types. This has sometimes been used by researchers to identify bats flying in an area simply by
Human echolocation since at least the 1950s. In earlier times, human echolocation was sometimes described as "facial vision" or "obstacle sense," as it was believed that the proximity of nearby objects caused pressure changes on the skin. Only in the 1940s did a series of experiments performed in the Cornell Psychological Laboratory show that sound and hearing, rather than pressure changes on the skin, were the mechanisms driving this ability. The field of human and animal echolocation was surveyed in book form as early as 1959. See also White, et al. (1970) Many blind individuals passively use natural environmental echoes to sense