Pollen as large as about 90–100 µm. In angiosperms, during flower development the anther is composed of a mass of cells that appear undifferentiated, except for a partially differentiated dermis. As the flower develops, four groups of sporogenous cells form within the anther. The fertile sporogenous cells are surrounded by layers of sterile cells that grow into the wall of the pollen sac. Some of the cells grow into nutritive cells that supply nutrition for the microspores that form by meiotic division from the sporogenous cells. In a process called microsporogenesis, four haploid microspores are produced from each diploid sporogenous cell
Pollen great quantities of very lightweight pollen grains, sometimes with air-sacs. Non-flowering seed plants (e.g., pine trees) are characteristically anemophilous. Anemophilous flowering plants generally have inconspicuous flowers. Entomophilous (literally insect-loving) plants produce pollen that is relatively heavy, sticky and protein-rich, for dispersal by insect pollinators attracted to their flowers. Many insects and some mites are specialized to feed on pollen, and are called palynivores. In non-flowering seed plants, pollen germinates in the pollen chamber, located beneath the micropyle, underneath the integuments of the ovule. A pollen tube is produced, which grows into the nucellus to provide nutrients for the developing sperm