Megabat long, thus smaller than some microbats, for example, the Mauritian tomb bat. The largest attain a wingspan of , and weigh as much as . Most fruit bats have large eyes, allowing them to orient themselves visually in twilight and inside caves and forests. Their sense of smell is excellent. In contrast to the microbats, the fruit bats do not use echolocation (with one exception, the Egyptian fruit bat "Rousettus egyptiacus", which uses high-pitched tongue clicks to navigate in caves). Megabats make up the only family (Pteropodidae) in order Chiroptera that is not capable of laryngeal echolocation. Echolocation and flight
Evolutionary arms race on the amount of evolutionary time and whether or not the moth species has developed secondary uses for hearing. Some bats are known to use clicks at frequencies above or below moths' hearing ranges. This is known as the allotonic frequency hypothesis. It argues that the auditory systems in moths have driven their bat predators to use higher or lower frequency echolocation to circumvent the moth hearing.Barbastelle bats have evolved to use a quieter mode of echolocation, calling at a reduced volume and further reducing the volume of their clicks as they close in on prey moths. The lower volume