Megabat evolved early in the lineage of chiropterans. Although echolocation was later lost in family Pteropodidae, bats in the genus "Rousettus" are capable of primitive echolocation through clicking their tongues, and some species have been shown to create clicks similar to those of echolocating bats using their wings. Both echolocation and flight are energetically expensive processes. The nature of the flight and echolocation mechanism of bats allows for creation of echolocation pulses with minimal energy use. Energetic coupling of these two processes is thought to have allowed for both energetically expensive processes to evolve in bats. The loss of echolocation may
Syconycteris Syconycteris is also capable of echolocation. The Etolo tribe of New Guinea will sometimes include Syconycteris bats in their diet, along with other bats. Syconycteris is unique among Pteropid bats because they do not produce spats. Spats are created by slow feeding bats that chew and swallow the juices of their food while pressing the pulp and seeds into a spat or wadge which is then spit out. This makes them more effective seed dispersers because seeds in spats tend to travel less distance and are more vulnerable to fungi compared to seeds that are defacated like those eaten by