Animal echolocation in the external ear. There are two hypotheses about the evolution of echolocation in bats. The first suggests that laryngeal echolocation evolved twice in Chiroptera, once in the Yangochiroptera and once in the horseshoe bats (Rhinolophidae). The second proposes that laryngeal echolocation had a single origin in Chiroptera, was subsequently lost in the family Pteropodidae (all megabats), and later evolved as a system of tongue-clicking in the genus "Rousettus". Individual bat species echolocate within specific frequency ranges that suit their environment and prey types. This has sometimes been used by researchers to identify bats flying in an area simply by
Animal echolocation as 212 kHz. Insectivorous aerial-hawking bats have a call frequency between 20 kHz and 60 kHz because it is the frequency that gives the best range and image acuity and makes them less conspicuous to insects. However, low frequencies are adaptive for some species with different prey and environments. "Euderma maculatum", a species that feeds on moths, uses a particularly low frequency of 12.7 kHz that cannot be heard by moths. Echolocation calls can be composed of two different types of frequency structure: frequency modulated (FM) sweeps, and constant frequency (CF) tones. A particular call can consist of one, the