Pollen Pollen Pollen is a fine to coarse powdery substance comprising pollen grains which are male microgametophytes of seed plants, which produce male gametes (sperm cells). Pollen grains have a hard coat made of sporopollenin that protects the gametophytes during the process of their movement from the stamens to the pistil of flowering plants, or from the male cone to the female cone of coniferous plants. If pollen lands on a compatible pistil or female cone, it germinates, producing a pollen tube that transfers the sperm to the ovule containing the female gametophyte. Individual pollen grains are small enough to require
Pollen form two sperm cells. The sperm cells are carried to their destination in the tip of the pollen tube. Double-strand breaks in DNA that arise during pollen tube growth appear to be efficiently repaired in the generative cell that carries the male genomic information to be passed on to the next plant generation. However, the vegetative cell that is responsible for tube elongation appears to lack this DNA repair capability. Pollen's sporopollenin outer sheath affords it some resistance to the rigours of the fossilisation process that destroy weaker objects; it is also produced in huge quantities. There is an extensive