Megabat evolved early in the lineage of chiropterans. Although echolocation was later lost in family Pteropodidae, bats in the genus "Rousettus" are capable of primitive echolocation through clicking their tongues, and some species have been shown to create clicks similar to those of echolocating bats using their wings. Both echolocation and flight are energetically expensive processes. The nature of the flight and echolocation mechanism of bats allows for creation of echolocation pulses with minimal energy use. Energetic coupling of these two processes is thought to have allowed for both energetically expensive processes to evolve in bats. The loss of echolocation may
Long-legged bat family Phyllostomidae that use low-intensity “whispering” echos, the Long-Legged Bat has been shown to produce high-intense echolocation calls to detect its prey. M. macrophyllum will adjust the intensity of its calls depending on the environment that it’s in. When in areas with high acoustic clutter, the bats will decrease their signal intensity, while in more open areas, the bats will increase their signal intensity. By increasing call intensity in a more open environment, this allows these bats to have a broader detection range for food. Average intensity output has been shown increase by 11 dB, corresponding to a 4x amplitude