Megabat evolved early in the lineage of chiropterans. Although echolocation was later lost in family Pteropodidae, bats in the genus "Rousettus" are capable of primitive echolocation through clicking their tongues, and some species have been shown to create clicks similar to those of echolocating bats using their wings. Both echolocation and flight are energetically expensive processes. The nature of the flight and echolocation mechanism of bats allows for creation of echolocation pulses with minimal energy use. Energetic coupling of these two processes is thought to have allowed for both energetically expensive processes to evolve in bats. The loss of echolocation may
Bulldog bat are long and narrow and their ears are large and funnel shaped. Unusual among bats, they have cheek-pouches for storing food. They also have full lips divided by a fold of skin giving a 'hare lip' look which together with the cheek pouches gives them their bulldog-like appearance. The species of lesser bulldog bats are insectivorous and whilst the greater bulldog bats also eat insects their chief food is fish. They use their echolocation to pinpoint the ripples they make on the surfaces of water. The greater bulldog bat trawls the water with its long, curved talons approximately 2–3 cm