Megabat evolved early in the lineage of chiropterans. Although echolocation was later lost in family Pteropodidae, bats in the genus "Rousettus" are capable of primitive echolocation through clicking their tongues, and some species have been shown to create clicks similar to those of echolocating bats using their wings. Both echolocation and flight are energetically expensive processes. The nature of the flight and echolocation mechanism of bats allows for creation of echolocation pulses with minimal energy use. Energetic coupling of these two processes is thought to have allowed for both energetically expensive processes to evolve in bats. The loss of echolocation may
Lesser long-eared bat sight is mostly restricted to aerial prey while they a technique that the species uses to take advantage of seasonal booms of tettigoniid crickets as well as general ground and aerial prey, known as passive listing. This method of hunting does not rely on sight or echolocation. Some moths species, like noctuids, have evolved to develop their own form of "ears" that are able to pick up the frequency of bat calls, however they are unable to sense long-eared bats as the bats use a minute whispering technique different from echolocation which bounces off the moths and is picked up