Megabat evolved early in the lineage of chiropterans. Although echolocation was later lost in family Pteropodidae, bats in the genus "Rousettus" are capable of primitive echolocation through clicking their tongues, and some species have been shown to create clicks similar to those of echolocating bats using their wings. Both echolocation and flight are energetically expensive processes. The nature of the flight and echolocation mechanism of bats allows for creation of echolocation pulses with minimal energy use. Energetic coupling of these two processes is thought to have allowed for both energetically expensive processes to evolve in bats. The loss of echolocation may
Waterhouse's leaf-nosed bat happen in that last trimester. Like many bats, this species uses echolocation as a means to locate their prey. In particular, "Macrotus waterhousii" uses a low intensity, broadband-like call to aide in their search for food. The maximum frequency of a call is 73.65 kHz with a minimum of 46.19 kHz, creating a bandwidth of 27.46 kHz. The call ranges from about 1 second to 3 seconds. In addition, to echolocation, "M. waterhousii" uses the sound made by the prey itself to locate it. This type of foraging behavior leads to a preference for a cluttered habitat. As "M. waterhousii"