ABC model of flower development well as in the determination of the floral meristem. Therefore, the "AG" mutants are devoid of androecium and gynoecium and they have petals and sepals in their place. In addition, the growth in the centre of the flower is undifferentiated, therefore the petals and sepals grow in repetitive verticils. The "PLENA" ("PLE") gene is present in "A. majus", in place of the "AG" gene, although it is not an ortholog. However, the "FARINELLI" ("FAR") gene is an ortholog, which is specific to the development of the anthers and the maturation of pollen. In "Petunia", "Antirrhinum" and in maize the C
Pollen (microsporocyte, pollen mother cell or meiocyte), after meiotic division. After the formation of the four microspores, which are contained by callose walls, the development of the pollen grain walls begins. The callose wall is broken down by an enzyme called callase and the freed pollen grains grow in size and develop their characteristic shape and form a resistant outer wall called the exine and an inner wall called the intine. The exine is what is preserved in the fossil record. Two basic types of microsporogenesis are recognised, simultaneous and successive. In simultaneous microsporogenesis meiotic steps I and II are completed