Megabat long, thus smaller than some microbats, for example, the Mauritian tomb bat. The largest attain a wingspan of , and weigh as much as . Most fruit bats have large eyes, allowing them to orient themselves visually in twilight and inside caves and forests. Their sense of smell is excellent. In contrast to the microbats, the fruit bats do not use echolocation (with one exception, the Egyptian fruit bat "Rousettus egyptiacus", which uses high-pitched tongue clicks to navigate in caves). Megabats make up the only family (Pteropodidae) in order Chiroptera that is not capable of laryngeal echolocation. Echolocation and flight
Human echolocation later participated in experiments in facial vision (White, et al. 1970). About 1998, he visited the Auditory Neuroethology Laboratory at the University of Maryland and was interviewed about his experience with facial vision. The researchers in the lab study bat echolocation and were aware of the "Wiederorientierung" phenomenon described by Griffin (1959), where bats, despite continuing to emit echolocation calls, use path integration in familiar acoustic space. Scadden indicated that he found echolocation required extra effort, and would not use it to navigate in familiar areas unless he was alert for obstacles, thus providing insight into the bat behavior. The