Animal echolocation in the external ear. There are two hypotheses about the evolution of echolocation in bats. The first suggests that laryngeal echolocation evolved twice in Chiroptera, once in the Yangochiroptera and once in the horseshoe bats (Rhinolophidae). The second proposes that laryngeal echolocation had a single origin in Chiroptera, was subsequently lost in the family Pteropodidae (all megabats), and later evolved as a system of tongue-clicking in the genus "Rousettus". Individual bat species echolocate within specific frequency ranges that suit their environment and prey types. This has sometimes been used by researchers to identify bats flying in an area simply by
Barbastelle this world, participated in an evolutionary arms race. The foe for the bat order Chiroptera is its prey, the moth order Lepidoptera. To minimize the risk of the bats' echolocation, moths [and in rarest examples, butterflies] have evolved to detect the echolocation calls of hunting bats, and evoke evasive flight manoeuvres, or reply with their own ultrasonic clicks to confuse the bat's echolocation. The Arctiidae subfamily of Noctuid moths uniquely respond to bat echolocation in three prevailing hypotheses: startle, sonar jamming, and acoustic aposematic defence. All these differences depend on specific environmental settings and the type of echolocation call; however,