Onychonycteris known species, hence "Onychonycteris" meaning "clawed bat". The specific epithet is a tribute to the fossil prospector and preparator who discovered it, Bonnie Finney. "Onychonycteris finneyi" was the strongest evidence so far in the debate on whether bats developed echolocation before or after they evolved the ability to fly. "O. finneyi" had well-developed wings, and could clearly fly, but lacked the enlarged cochlea of all extant echolocating bats, closer resembling the old world fruit bats which do not echolocate. This indicates that early bats could fly before they could echolocate. However, an independent evaluation of the "Onychonycteris" reference fossil in
Animal echolocation naturalist Louis Jurine repeated Spallanzani's experiments (using different species of bat), and concluded that when bats hunt at night, they rely on hearing. In 1908, Walter Louis Hahn confirmed Spallanzani's and Jurine's findings. In 1912, the inventor Hiram Maxim independently proposed that bats used sound below the human auditory range to avoid obstacles. In 1920, the English physiologist Hamilton Hartridge correctly proposed instead that bats used frequencies above the range of human hearing. Echolocation in odontocetes (toothed whales) was not properly described until two decades after Griffin and Galambos' work, by Schevill and McBride in 1956. However, in 1953, Jacques