John Edward Gray wrote nearly 500 papers, including many descriptions of species new to science. These had been presented to the Museum by collectors from around the world, and included all branches of zoology, although Gray usually left the descriptions of new birds to his younger brother and colleague George. Gray was also active in malacology, the study of molluscs. John Edward Gray was buried at St Mary's Church, Lewisham. Gray was one of the most prolific taxonomists in the history of zoology. He described more than 300 species and subspecies of reptiles, only surpassed by his successors at the British Museum, George
Primatology statistics and modern data collection to conclude her 1978 doctoral thesis regarding orangutan behavior and interactions. Long-term sites of research tend to be best associated with their founders, and this led to some tension between younger primatologists and the veterans in the field. The discipline of Japanese primatology was developed out of animal ecology. It is mainly credited to Kinji Imanishi and Junichiro Itani. Imanishi was an animal ecologist who began studying wild horses before focusing more on primate ecology. He helped found the Primate Research Group in 1950. Junichiro was a renowned anthropologist and a professor at Kyoto University.