Atriplex are C-plants, with a characteristic leaf anatomy (kranz anatomy). The genus "Atriplex" is distributed nearly worldwide from subtropical to temperate and to subarctic regions. Most species-rich are Australia, North America, South America and Eurasia. Many species are halophytes and are adapted to dry environments with salty soils. The genus evolved in Middle Miocene, the C-photosynthesis pathway developed about 14.1-10.9 million years ago, when the climate became increasingly dry. The genus diversified rapidly and spread over the continents. The C"Atriplex" colonized North America probably from Eurasia during the Middle/Late Miocene, about 9.8-8.8 million years ago, and later spread to South America.
Biosalinity 20th century. Research in biosalinity includes studies of the biochemical and physiological mechanisms of salt tolerance in plants, breeding and selection for salt tolerance (halotolerance), discovery of periods in a crop plant's life cycle when it may be less sensitive to salt, use of saline irrigation water to increase desirable traits (such as sugar concentration in a fruit) or to control the ripening process, study of the interaction between salinity and soil properties, and development of naturally salt-tolerant plant species (halophytes) into useful agricultural crops. See also halophile bacteria, which thrive under conditions of high salinity. When properly applied (watering