Drusilla (Buffy the Vampire Slayer) Drusilla (Buffy the Vampire Slayer) Drusilla, or Dru, is a fictional character created by Joss Whedon and David Greenwalt for the American television series, "Buffy the Vampire Slayer" and "Angel". The character is portrayed by American actress Juliet Landau. Drusilla is introduced alongside her lover Spike (James Marsters) in the second season of "Buffy the Vampire Slayer" to serve as new antagonists to the series' heroine, vampire Slayer Buffy Summers (Sarah Michelle Gellar) and alongside Darla (Julie Benz) and Wolfram & Hart as the main antagonist of the second season of Angel. In contrast to the series' previous central villain,
Buffy the Vampire Slayer (season 4) Buffy the Vampire Slayer (season 4) The fourth season of the television series Buffy the Vampire Slayer premiered on October 5, 1999, on The WB and concluded its 22-episode season on May 23, 2000. It maintained its previous timeslot, airing Tuesdays at 8:00 pm ET. Beginning with this season, the character of Angel was given his own "series", which aired on The WB following "Buffy". Various "Buffy" characters made appearances in "Angel", including Buffy herself; Cordelia Chase, formerly a regular in "Buffy", and Wesley Wyndam-Pryce, who appeared in "Buffy" season three. Season four sees Buffy and Willow enroll at UC