Drusilla (Buffy the Vampire Slayer) Drusilla (Buffy the Vampire Slayer) Drusilla, or Dru, is a fictional character created by Joss Whedon and David Greenwalt for the American television series, "Buffy the Vampire Slayer" and "Angel". The character is portrayed by American actress Juliet Landau. Drusilla is introduced alongside her lover Spike (James Marsters) in the second season of "Buffy the Vampire Slayer" to serve as new antagonists to the series' heroine, vampire Slayer Buffy Summers (Sarah Michelle Gellar) and alongside Darla (Julie Benz) and Wolfram & Hart as the main antagonist of the second season of Angel. In contrast to the series' previous central villain,
Buffy the Vampire Slayer comics and Joss Whedon (or his office), and the books were therefore later published as official Buffy merchandise. The stories in the Buffy comics take place in between episodes of the "Buffy the Vampire Slayer" TV series, and continue after the series ended. Issues 1 to 63 take place during the period that the series was still on air, and none are written by Joss Whedon himself. The earlier issues are not very easy to place in the series other than the season they are set in. The stories get more specific later on, however; for example, the "Oz" Buffy comics