Evolution of cetaceans "Caperea marginata") all have derived characteristics presently unknown in any cetothere and vice versa (such as a sagittal crest). The adaptation of echolocation occurred when toothed whales (Odontoceti) split apart from baleen whales, and distinguishes modern toothed whales from fully aquatic archaeocetes. This happened around 34 million years ago in a second cetacean radiation. Modern toothed whales do not rely on their sense of sight, but rather on their sonar to hunt prey. Echolocation also allowed toothed whales to dive deeper in search of food, with light no longer necessary for navigation, which opened up new food sources. Toothed whales
Captive killer whales they are successful, and withholding it when they are not. Secondary reinforcement—things not essential to life, such as play time, tactile rewards and fun games—can also be used as rewards. The practice of keeping killer whales in captivity is controversial, and organizations such as World Animal Protection and the Whale and Dolphin Conservation campaign against the captivity of killer whales. Orcas in captivity may develop physical pathologies, such as the dorsal fin collapse seen in 80–90% of captive males. The captive environment bears little resemblance to their wild habitat, and the social groups that the killer whales are put into