Toothed whale porpoise as a fish. Toothed whales, as well as baleen whales, are descendants of land-dwelling mammals of the artiodactyl order (even-toed ungulates). They are closely related to the hippopotamus, sharing a common ancestor that lived around 54 million years ago (mya). The primitive cetaceans, or archaeocetes, first took to the sea approximately 49 mya and became fully aquatic by 5–10 million years later. The adaptation of echolocation and enhanced fat synthesis in blubber occurred when toothed whales split apart from baleen whales, and distinguishes modern toothed whales from fully aquatic archaeocetes. This happened around 34 mya. Unlike toothed whales, baleen
Cetacea salinity. A few toothed whales, such as some killer whales, feed on mammals, such as pinnipeds and other whales. Toothed whales have well-developed senses – their eyesight and hearing are adapted for both air and water, and they have advanced sonar capabilities using their melon. Their hearing is so well-adapted for both air and water that some blind specimens can survive. Some species, such as sperm whales, are well adapted for diving to great depths. Several species of toothed whales show sexual dimorphism, in which the males differ from the females, usually for purposes of sexual display or aggression. Cetacean