Drusilla (Buffy the Vampire Slayer) Drusilla (Buffy the Vampire Slayer) Drusilla, or Dru, is a fictional character created by Joss Whedon and David Greenwalt for the American television series, "Buffy the Vampire Slayer" and "Angel". The character is portrayed by American actress Juliet Landau. Drusilla is introduced alongside her lover Spike (James Marsters) in the second season of "Buffy the Vampire Slayer" to serve as new antagonists to the series' heroine, vampire Slayer Buffy Summers (Sarah Michelle Gellar) and alongside Darla (Julie Benz) and Wolfram & Hart as the main antagonist of the second season of Angel. In contrast to the series' previous central villain,
Darla (Buffy the Vampire Slayer) Darla (Buffy the Vampire Slayer) Darla is a recurring fictional character created by Joss Whedon and played by Julie Benz in the first, second, and fifth seasons of the American supernatural television series "Buffy the Vampire Slayer". The character later appeared in the "Buffy" spin-off series "Angel", making at least one appearance in every season. She made her last television appearance in 2004, appearing as a special guest star in the fifth and final season of "Angel". Darla is introduced in "Welcome to the Hellmouth", the first episode of "Buffy the Vampire Slayer", in 1997. It is revealed early on