Toothed whale porpoise as a fish. Toothed whales, as well as baleen whales, are descendants of land-dwelling mammals of the artiodactyl order (even-toed ungulates). They are closely related to the hippopotamus, sharing a common ancestor that lived around 54 million years ago (mya). The primitive cetaceans, or archaeocetes, first took to the sea approximately 49 mya and became fully aquatic by 5–10 million years later. The adaptation of echolocation and enhanced fat synthesis in blubber occurred when toothed whales split apart from baleen whales, and distinguishes modern toothed whales from fully aquatic archaeocetes. This happened around 34 mya. Unlike toothed whales, baleen
Toothed whale and Ziphiidae) that have the highest quantities of wax esters and BCFAs in their blubber are also the species that dive the deepest and for the longest amount of time. Toothed whales have a two-chambered stomach similar in structure to terrestrial carnivores. They have fundic and pyloric chambers. Cetaceans have two flippers on the front, and a tail fin. These flippers contain four digits. Although toothed whales do not possess fully developed hind limbs, some, such as the sperm whale, possess discrete rudimentary appendages, which may contain feet and digits. Toothed whales are fast swimmers in comparison to seals, which