Toothed whale brain size with the expected brain size based on such allometric analysis provides an encephalisation quotient that can be used as another indication of animal intelligence. Sperm whales have the largest brain mass of any animal on earth, averaging and in mature males, in comparison to the average human brain which averages in mature males. The brain to body mass ratio in some odontocetes, such as belugas and narwhals, is second only to humans. Dolphins are known to engage in complex play behaviour, which includes such things as producing stable underwater toroidal air-core vortex rings or "bubble rings". Two main
Whale shark feeder, targeting concentrations of plankton or fish. It is able to ram filter feed or can gulp in a stationary position. This is in contrast to the passive feeding basking shark, which does not pump water. Instead, it swims to force water across its gills. A juvenile whale shark is estimated to eat 21 kg (46 pounds) of plankton per day. The BBC program "Planet Earth" filmed a whale shark feeding on a school of small fish. The same documentary showed footage of a whale shark timing its arrival to coincide with the mass spawning of fish shoals and feeding