Mouflon vignei" (or "orientalis") "arkal" and "O. v./o. bochariensis", grouped with two different clades of argali ("Ovis ammon"). The ancestral sheep is presumed to have had 60 chromosomes, as in goats ("Capra"). Mouflon and domestic sheep have 54 chromosomes, with three pairs (1+3, 2+8, 5+11) of ancestral acrocentric chromosomes joined to form bi-armed chromosomes. This is in contrast to the argali and urial, which have 56 and 58 chromosomes respectively. If the urial is as closely related to the mouflons as mitochondrial DNA indicates, then two chromosomes would need to have split during its evolution away from the mouflon (sub)species. In
Mouflon domesticated animals, where they have naturalized in the mountainous interiors of these islands over the past few thousand years, giving rise to the subspecies known as European mouflon ("O. orientalis musimon"). On the island of Cyprus, the mouflon or "agrino" became a different and endemic subspecies known as the Cyprus mouflon ("O. o. ophion"). The Cyprus mouflon population contains only about 3,000 animals. They are now rare on the islands, but are classified as feral animals by the IUCN. They were later successfully introduced into continental Europe, including Portugal, Spain, France, Germany, central Italy, Switzerland, Austria, the Netherlands, Belgium, the