Galanthus (convolute). He merged "Nivalis" and "Plicati" into series "Galanthus", and divided "Latifolii" into two subseries, "Glaucaefolii" and "Viridifolii" . Early molecular phylogenetic studies confirmed the genus was monophyletic and suggested four clades, which were labelled as series, and showed that Davis' subseries were not monophyletic. An expanded study in 2013 demonstrated seven major clades corresponding to biogeographical distribution. This study used nuclear encoded nrITS (Nuclear ribosomal internal transcribed spacer), and plastid encoded "matK" (Maturase K), "trnL-F", "ndhF", and "psbK–psbI", and examined all species recognised at the time, and also two naturally occurring putative hybrids. The morphological characteristic of vernation that
Chlamydomonas nivalis "Uredo nivalis". Over the next century, many researchers disputed over whether these organisms were lichen, plants, alga, or animal. It was not until the early 20th century when researchers finally began to agree on the algal nature of the organism and gave its currently known name, "Chlamydomonas nivalis". In 1968 "C. nivalis" was officially recognized as a collective taxon. Unfortunately, due to the lack of sequencing techniques, reliance on visually examining similarly looking snow alga, and complicated life cycle of this species, errors continued to be made in classifying this and other species of snow algae. Today, "C. nivalis" has