Galanthus (convolute). He merged "Nivalis" and "Plicati" into series "Galanthus", and divided "Latifolii" into two subseries, "Glaucaefolii" and "Viridifolii" . Early molecular phylogenetic studies confirmed the genus was monophyletic and suggested four clades, which were labelled as series, and showed that Davis' subseries were not monophyletic. An expanded study in 2013 demonstrated seven major clades corresponding to biogeographical distribution. This study used nuclear encoded nrITS (Nuclear ribosomal internal transcribed spacer), and plastid encoded "matK" (Maturase K), "trnL-F", "ndhF", and "psbK–psbI", and examined all species recognised at the time, and also two naturally occurring putative hybrids. The morphological characteristic of vernation that
Chlamydomonas nivalis such that it is covered by the carotenoid globules full of astaxanthin that will provide protection against UV radiation. The majority (91%) of astaxanthin derivatives are stored in its monoester form within dormant "C. nivalis" red cysts. Astaxanthin is the pigment that makes the cell appear deep red. Other pigments that can also be found in "C. nivalis" include violaxanthin and fadonirubin. Visible algal blooms could be a crucial determinant of surface albedo. It has been suggested that algal blooms partially composed of "C. nivalis" may contribute to lowering ice and snow albedo. The red coloured pigments produced by the