History of the domestic sheep exact line of descent between domestic sheep and their wild ancestors is unclear. The most common hypothesis states that "Ovis aries" is descended from the Asiatic ("O. orientalis") species of mouflon. A few breeds of sheep, such as the Castlemilk Moorit from Scotland, were formed through crossbreeding with wild European mouflon. The urial ("O. vignei") was once thought to have been a forebear of domestic sheep, as they occasionally interbreed with mouflon in the Iranian part of their range. However, the urial, argali ("O. ammon"), and snow sheep ("O. nivicola") have a different number of chromosomes than other Ovis species,
Dmitry Belyayev (zoologist) also been of great interest to Darwin. It was recognized that domesticated animals differ in several ways from their wild counterparts, but it was not known what principle of selection had guided the Neolithic farmers who had first domesticated these species thousands of years ago. Belyayev's hypothesis was that "all domesticated species had been selected for a single criterion: tameness." Belyayev further theorized that this attribute "had dragged along with it most of the other features that distinguish domestic animals from their wild forebears, like droopy ears, patches of white in the fur and changes in skull shape." In other