Sphagnum ones ("Rigida" and "Squarrosa") has been clarified using molecular phylogenetics. All but two species normally identified as "Sphagnum" reside in one clade; two other species have recently been separated into new families within the Sphagnales reflecting an ancestral relationship with the Tasmanian endemic "Ambuchanania" and long phylogenetic distance to the rest of "Sphagnum". Within main clade of "Sphagnum", phylogenetic distance is relatively short, and molecular dating methods suggest nearly all current "Sphagnum" species are descended from a radiation that occurred just 14 million years ago. "Sphagnum" mosses occur mainly in the Northern Hemisphere in peat bogs, conifer forests, and moist
Sphagnum all other land plants, has an alternation of generations; like other bryophytes, the haploid gametophyte generation is dominant and persistent. Unlike other mosses, the long-lived gametophytes do not rely upon rhizoids to assist in water uptake. "Sphagnum" species can be unisexual (male or female, dioecious) or bisexual (male and female gametes produced from the same plant; monoecious); In North America, 80% of "Sphagnum" species are unisexual. Gametophytes have substantial asexual reproduction by fragmentation, producing much of the living material in sphagnum peatlands. Swimming sperm fertilize eggs contained in archegonia that remain attached to the female gametophyte. The sporophyte is relatively