Sphagnum ones ("Rigida" and "Squarrosa") has been clarified using molecular phylogenetics. All but two species normally identified as "Sphagnum" reside in one clade; two other species have recently been separated into new families within the Sphagnales reflecting an ancestral relationship with the Tasmanian endemic "Ambuchanania" and long phylogenetic distance to the rest of "Sphagnum". Within main clade of "Sphagnum", phylogenetic distance is relatively short, and molecular dating methods suggest nearly all current "Sphagnum" species are descended from a radiation that occurred just 14 million years ago. "Sphagnum" mosses occur mainly in the Northern Hemisphere in peat bogs, conifer forests, and moist
Portlethen Moss lower elevations in every direction. Due to the high winds, moderate precipitation and cool temperatures that generally prevail, conditions are favourable for formation of an acid bog, since water stagnates, but eventually evaporates with ensuing acidity enhancement of decaying organic matter. There was virtually no drainage outlet in prehistoric times, and little drainage even in modern times. Many coastal mosses were initiated by the process of glaciation, which sheared rock formations to a generally level terrain, while also gouging moderate sized craters that would pond. This description fits the fundamental situation of Portlethen Moss, where sphagnum would have flourished over