Galanthus earlier authors had mainly relied on was shown to be highly homoplasious. A number of species, such as "G. nivalis" and "G. elwesii" demonstrated intraspecific biogeographical clades, indicating problems with speciation and there may be a need for recircumscription. These clades were assigned names, partly according to Davis' previous groupings. In this model clade Platyphyllus is sister to the rest of the genus. By contrast another study performed at the same time, using both nuclear and chloroplast DNA but limited to the 14 species found in Turkey, largely confirmed Davis' series and subseries, and with biogeographical correlation. Series "Galanthus" in
Chlamydomonas nivalis have been found on the surface of "C. nivalis" cysts. The unknown bacteria were not detected in control samples that did not contain "C. nivalis" which strongly suggests that it must be associated with the algae. Another bacterium, "Mesorhizobium loti", was found as contamination in a "C. nivalis" culture, but further testing suggested that this bacteria may be synthesizing vitamin B12 for the algae. In cryoconite holes "C. nivalis" can be found among bacteria, virus-like particles, ciliates, and Chlorophyte species. Ice worms have also been found to live preferentially under "C. nivalis" in glaciers, possibly using the algae as a