Sphagnum ones ("Rigida" and "Squarrosa") has been clarified using molecular phylogenetics. All but two species normally identified as "Sphagnum" reside in one clade; two other species have recently been separated into new families within the Sphagnales reflecting an ancestral relationship with the Tasmanian endemic "Ambuchanania" and long phylogenetic distance to the rest of "Sphagnum". Within main clade of "Sphagnum", phylogenetic distance is relatively short, and molecular dating methods suggest nearly all current "Sphagnum" species are descended from a radiation that occurred just 14 million years ago. "Sphagnum" mosses occur mainly in the Northern Hemisphere in peat bogs, conifer forests, and moist
Sphagnum short-lived, and consists almost entirely of a shiny green, spherical spore capsule that becomes black with spores. Sporophytes are raised on stalks to facilitate spore dispersal, but unlike other mosses, "Sphagnum" stalks are produced by the maternal gametophyte. Tetrahedral haploid spores are produced in the sporophyte by meiosis, which are then dispersed when the capsule explosively discharges its cap, called an operculum, and shoots the spores some distance. The spores germinate to produce minute protonemae, which start as filaments, can become thalloid, and can produce a few rhizoids. Soon afterwards, the protonema develops buds and these differentiate into its characteristic,