White-eared titi the males. There is a strong bond between the adult mating pair, they stay close and carry out activities together. Either member of the pair may follow the other and leadership changes through the day. Evidence of the strength of the pair bond is shown by grooming, huddling together with their tails twined, nuzzling, and gentle grasping. Titi monkeys are highly territorial and when confronted with another family group, both will respond with threatening behaviour, males showing increased agitation towards intruding males. When not close together, the pair show a significant amount of distress and agitation. Titi monkeys are well
Collared titi least two species in this complex. But whether the Lucifer titi, "Cheracebus lucifer", or the Colombian black-handed titi, "Cheracebus medemi", are good species from this complex is in doubt. They are probably subspecies of "Cheracebus torquatus". Nevertheless, in this treatment "C. torquatus" is used in the sense of Hershkovitz (1990) until the systematics of this species complex becomes clearer. Five adults weighed an average of 1462 g (range 1410–1722 g) with a head-body length of around 290–390 mm and a tail length of about 350–400 mm. The face has very little hair, being limited to sparse short white hairs over