Ladoga seal Ladoga seal The Ladoga ringed seal (; Pusa hispida ladogensis), is a freshwater subspecies of the ringed seal ("Pusa hispida") which are found entirely in Lake Ladoga in northwestern Russia. The subspecies evolved during the last ice age, about 11,000 years ago. As the glaciers retreated and water levels changed, the Baltic ringed seal (including Ladoga seals) was trapped in freshwater lakes and separated from the Arctic ringed seal. It is related to the even smaller population of Saimaa ringed seals in Lake Saimaa, a lake that flows into Ladoga through the Vuoksi River. The adult Ladoga seal grows to
Lake Ladoga of the 1960s Ladoga has become considerably eutrophicated. Nizhnesvirsky Natural Reserve is situated along the shore of Lake Ladoga immediately to the north of the mouth of the River Svir. In the Middle Ages, the lake formed a vital part of the trade route from the Varangians to the Eastern Roman Empire, with the Norse emporium at Staraya Ladoga defending the mouth of the Volkhov since the 8th century. In the course of the Swedish–Novgorodian Wars, the area was disputed between the Novgorod Republic and Sweden. In the early 14th century, the fortresses of Korela (Kexholm) and Oreshek (Nöteborg) were