Craniate with well-defined heads, thus excluding members of the chordate subphyla Tunicata (tunicates) and Cephalochordata (lancelets), but including Myxini, which have cartilaginous skulls and tooth-like structures composed of keratin. Craniata also includes all lampreys and armoured jawless fishes, armoured fish, sharks, skates, and rays, and teleostomians: spiny sharks, bony fish, lissamphibians, temnospondyls and protoreptiles, sauropsids and mammals. The craniate head consists of a brain, sense organs, including eyes, and a skull. In addition to distinct crania (sing. "cranium"), craniates possess many derived characteristics, which have allowed for more complexity to follow. Molecular-genetic analysis of craniates reveals that, compared to less complex
Tool use by animals Tool use by animals Tool use by animals is a phenomenon in which an animal uses any kind of tool in order to achieve a goal such as acquiring food and water, grooming, defense, recreation or construction. Originally thought to be a skill only possessed by humans, some tool use requires a sophisticated level of cognition. There is considerable discussion about the definition of what constitutes a tool and therefore which behaviours can be considered as true examples of tool use. A wide range of animals are considered to use tools including mammals, birds, fish, cephalopods and insects. Primates are