John Hanning Speke a new expedition and took a boat along the entire shore of Lake Victoria; he established that Lake Tanganyika and the Nile were not connected in any way, and he explored the headwaters of Lake Edward. It was now proven that Speke had been right all along, and that the Nile flowed from Lake Victoria via Ripon Falls and Murchison Falls to Lake Albert and from there to Gondokoro. It was in later years, however, that other explorers found that Lake Victoria is not the southernmost source of the Nile, rather it is the Kagera River that flows from the
John Hanning Speke "race" comprised many tribes, including the Watusi (Tutsi). Speke described their physical appearances as having retaineddespite the hair-curling and skin-darkening effects of intermarriage"a high stamp of Asiatic feature, of which a marked characteristic is a bridged instead of bridgeless nose". Two species of African reptiles are named in his honor: Speke's hinge-back tortoise, "Kinixys spekii"; and Speke's sand lizard, "Heliobolus spekii". Two species of antelope are named in his honor; "Tragelaphus spekii" and "Gazella spekei". The film "Mountains of the Moon" (1990), starring Scottish actor Iain Glen as Speke, related the story of the Burton-Speke controversy, portrayed as having been