Major explorations after the Age of Discovery to locate the source of the River Nile. Expeditions by Burton and Speke (1857–1858) and Speke and Grant (1863) located Lake Tanganyika and Lake Victoria. It was eventually proved to be the latter from which the Nile flowed. Explorers were also active in other parts of the continent. Southern Morocco, the Sahara and the Sudan were traversed in many directions between 1860 and 1875 by Georg Schweinfurth and Gustav Nachtigal. These travellers not only added considerably to geographical knowledge, but obtained invaluable information concerning the people, languages and natural history of the countries in which they sojourned. Among the discoveries
John Hanning Speke "race" comprised many tribes, including the Watusi (Tutsi). Speke described their physical appearances as having retaineddespite the hair-curling and skin-darkening effects of intermarriage"a high stamp of Asiatic feature, of which a marked characteristic is a bridged instead of bridgeless nose". Two species of African reptiles are named in his honor: Speke's hinge-back tortoise, "Kinixys spekii"; and Speke's sand lizard, "Heliobolus spekii". Two species of antelope are named in his honor; "Tragelaphus spekii" and "Gazella spekei". The film "Mountains of the Moon" (1990), starring Scottish actor Iain Glen as Speke, related the story of the Burton-Speke controversy, portrayed as having been