Craniate with well-defined heads, thus excluding members of the chordate subphyla Tunicata (tunicates) and Cephalochordata (lancelets), but including Myxini, which have cartilaginous skulls and tooth-like structures composed of keratin. Craniata also includes all lampreys and armoured jawless fishes, armoured fish, sharks, skates, and rays, and teleostomians: spiny sharks, bony fish, lissamphibians, temnospondyls and protoreptiles, sauropsids and mammals. The craniate head consists of a brain, sense organs, including eyes, and a skull. In addition to distinct crania (sing. "cranium"), craniates possess many derived characteristics, which have allowed for more complexity to follow. Molecular-genetic analysis of craniates reveals that, compared to less complex
Emotion in animals during complex social interactions. However, Hof and Van Der Gucht clarify that they do not know the nature of such feelings in these animals and that we cannot just apply what we see in great apes or ourselves to whales. They believe that more work is needed to know whether emotions are the same for humans and whales. From at least the time of Darwin, it has been known that chimpanzees and other great apes also perform a laugh-like vocalization. Research with rats has revealed that under particular conditions, they emit 50-kHz ultrasonic vocalisations (USV) which have been postulated to