Silkie the "SHH" gene, called the ZPA Regulatory Sequence (ZRS). This causes ectopic "SHH" expression in the anterior of the developing limb bud, leading to increased tissue growth and digits. While the feet of the Silkie display polydactyly, the wings have the standard tridactyly (three digit) arrangement. The Japanese Silkie initially develops additional digits in the wing as an embryo, but these are lost prior to hatching. The genetic cause behind Silkie polydactyly differs from those that cause polydactyly in the Dorking chicken breed, which is due to ectopic "FGF4" expression in the AER, with ectopic "SHH" a secondary effect. The
What Bird is That? according to their usual habitat. The accompanying text included brief notes on distribution, breeding and behaviour. While initial sales were poor, history tells us that Cayley had accurately discerned a market need and successfully responded to it. During WWII sales accelerated and Cayley’s first collection, reprinted time and time again, eventually became a household name among birdwatchers and the book was recognised as an Australian classic. The original sponsorship of "What Bird is That?" proved a great investment for the Gould League, who benefited from the sale of its numerous reprints and revised editions. By 1960, it was rated the