Sphagnum ones ("Rigida" and "Squarrosa") has been clarified using molecular phylogenetics. All but two species normally identified as "Sphagnum" reside in one clade; two other species have recently been separated into new families within the Sphagnales reflecting an ancestral relationship with the Tasmanian endemic "Ambuchanania" and long phylogenetic distance to the rest of "Sphagnum". Within main clade of "Sphagnum", phylogenetic distance is relatively short, and molecular dating methods suggest nearly all current "Sphagnum" species are descended from a radiation that occurred just 14 million years ago. "Sphagnum" mosses occur mainly in the Northern Hemisphere in peat bogs, conifer forests, and moist
Plant cell quite elongated, and may divide transversely to give a septate appearance. The role of this cell type is to support the plant in axes still growing in length, and to confer flexibility and tensile strength on tissues. The primary wall lacks lignin that would make it tough and rigid, so this cell type provides what could be called plastic support – support that can hold a young stem or petiole into the air, but in cells that can be stretched as the cells around them elongate. Stretchable support (without elastic snap-back) is a good way to describe what collenchyma does.