Silkie the "SHH" gene, called the ZPA Regulatory Sequence (ZRS). This causes ectopic "SHH" expression in the anterior of the developing limb bud, leading to increased tissue growth and digits. While the feet of the Silkie display polydactyly, the wings have the standard tridactyly (three digit) arrangement. The Japanese Silkie initially develops additional digits in the wing as an embryo, but these are lost prior to hatching. The genetic cause behind Silkie polydactyly differs from those that cause polydactyly in the Dorking chicken breed, which is due to ectopic "FGF4" expression in the AER, with ectopic "SHH" a secondary effect. The
What Bird is That? all-time best seller in Australian natural history and is still in print today. While sales and stature of Cayley’s original collection continued to grow, the prolific artist and ornothologist Cayley was working on a much more ambitious project – his so-called “big bird book”. From about 1918, he was creating illustrations of subspecies, plumage stages and eggs (now lost) of all known Australian birds. The project was incomplete at the time of his death in 1950, although the illustrations were all but finished. The ornithologist Terence Lindsey revived Cayley’s plans in the 1980s, creating the last and most significant revision