Sphagnum short-lived, and consists almost entirely of a shiny green, spherical spore capsule that becomes black with spores. Sporophytes are raised on stalks to facilitate spore dispersal, but unlike other mosses, "Sphagnum" stalks are produced by the maternal gametophyte. Tetrahedral haploid spores are produced in the sporophyte by meiosis, which are then dispersed when the capsule explosively discharges its cap, called an operculum, and shoots the spores some distance. The spores germinate to produce minute protonemae, which start as filaments, can become thalloid, and can produce a few rhizoids. Soon afterwards, the protonema develops buds and these differentiate into its characteristic,
Plant functional type key parameters are defined, such as fecundity, competitiveness, resorption (rate at which plant decays and returns nutrients to the soil after death), etc.; the value of each parameter is determined or inferred from observable characteristics such as plant height, leaf area, etc. Climatologists and ecologists struggle to determine which minimal set of plant characteristics best model the actual responses of the biosphere in response to climate changes. Plant functional type Plant functional types (PFTs) is a system used by climatologists to classify plants according to their physical, phylogenetic and phenological characteristics as part of an overall effort to develop a