Silkie the "SHH" gene, called the ZPA Regulatory Sequence (ZRS). This causes ectopic "SHH" expression in the anterior of the developing limb bud, leading to increased tissue growth and digits. While the feet of the Silkie display polydactyly, the wings have the standard tridactyly (three digit) arrangement. The Japanese Silkie initially develops additional digits in the wing as an embryo, but these are lost prior to hatching. The genetic cause behind Silkie polydactyly differs from those that cause polydactyly in the Dorking chicken breed, which is due to ectopic "FGF4" expression in the AER, with ectopic "SHH" a secondary effect. The
What Is a Nation? and continues to be invoked. Political historian Karl Deutsch, in a quote sometimes mistakenly attributed to Renan, said that a nation is "a group of people united by a mistaken view about the past and a hatred of their neighbours". Benedict Anderson's 1983 work Imagined Communities, which states that a nation is an "imagined political community", argues that Renan contradicts himself when he says French people must have forgotten the St. Bartholomew's Day massacre, yet does not explain what it is. In other words, Renan assumes that all his readers will remember the very massacre he says they have forgotten.