Asplenium rhizophyllum one collection on sandstone in the Shawnee Hills of Illinois. A triploid hybrid between walking fern and Tutwiler's spleenwort was accidentally produced in culture. A similar plant collected from limestone in Shepherdstown, West Virginia could have originated from the same parents, from an unreduced (diploid) gametophyte of Scott's spleenwort crossed with walking fern, or from an unreduced walking fern gametophyte crossed with ebony spleenwort. In 1813, Henry Muhlenberg listed lobed spleenwort as "Asplenium rhizophyllum" var. "pinnatifidum", although he did not provide a description distinguishing the variety from the typical species. It was described as the species "A. pinnatifidum" by Thomas
What Is and What Should Never Be descending riff [of "What Is and What Should Never Be"] is amazing: It's like a bow is being drawn back, and then it releases. The rhythm of the vocals is almost like a rap. It's insane — one of their most psychedelic songs." This was also one of the first songs recorded by the band for which Robert Plant received writing credit. According to rock journalist Stephen Davis, the author of the Led Zeppelin biography "Hammer of the Gods: The Led Zeppelin Saga", the lyrics for this song reflect a romance Plant had with his wife's younger sister. "What Is