Asplenium rhizophyllum one collection on sandstone in the Shawnee Hills of Illinois. A triploid hybrid between walking fern and Tutwiler's spleenwort was accidentally produced in culture. A similar plant collected from limestone in Shepherdstown, West Virginia could have originated from the same parents, from an unreduced (diploid) gametophyte of Scott's spleenwort crossed with walking fern, or from an unreduced walking fern gametophyte crossed with ebony spleenwort. In 1813, Henry Muhlenberg listed lobed spleenwort as "Asplenium rhizophyllum" var. "pinnatifidum", although he did not provide a description distinguishing the variety from the typical species. It was described as the species "A. pinnatifidum" by Thomas
American Type Founders that company to be even less profitable than was thought, Gillick was forced to resign. Thomas Roy Jones, a businessman with no experience in type founding, replaced Gillick. By 1933 the situation was desperate. Sales of type were less than 30% of 1926 levels while purchases of Kelly presses had plummeted to a mere 6.8% of what they had been. In October 1933 Jones filed a voluntary petition for bankruptcy. ATF was placed under the control of its creditors (chiefly consisting of several banks) and drastic measures were taken. Operations were consolidated, the Jersey City plant was closed and the