Petroleum microbiology oil while A. borkumensis simutaneously increases synthesis of anionic glucoproteins, which are used to emulsify hydrocarbons in the environment and increase their bioavailability. The presence of crude oil along with appropriate levels of nitrogen and phosphor catalyzes the removal of petroleum either by mechanisms that enhance the efficiency of substrate uptake or by direct biodegradation of aliphatic chains. Two well-known oil spills exemplify large scale marine bioremediation applications: In 1989, the Exxon Valdez ran aground, spilling 41.6 million liters of crude oil, and launching one of the first major bioremediation efforts for an oil spill. Cleanup of Alaskan shorelines relied
Exxon Valdez oil spill official position was that punitive damages greater than $25 million were not justified because the spill resulted from an accident, and because Exxon spent an estimated $2 billion cleaning up the spill and a further $1 billion to settle related civil and criminal charges. Attorneys for the plaintiffs contended that Exxon bore responsibility for the accident because the company "put a drunk in charge of a tanker in Prince William Sound." Exxon recovered a significant portion of clean-up and legal expenses through insurance claims associated with the grounding of the "Exxon Valdez". Also, in 1991, Exxon made a quiet, separate