Pleistocene The lower boundary of the Pleistocene Series is formally defined magnetostratigraphically as the base of the Matuyama (C2r) chronozone, isotopic stage 103. Above this point there are notable extinctions of the calcareous nanofossils: "Discoaster pentaradiatus" and "Discoaster surculus". The Pleistocene covers the recent period of repeated glaciations. The name Plio-Pleistocene has, in the past, been used to mean the last ice age. The revised definition of the Quaternary, by pushing back the start date of the Pleistocene to 2.58 Ma, results in the inclusion of all the recent repeated glaciations within the Pleistocene. The modern continents were essentially at their
Pleistocene "Homo erectus" for much of the Pleistocene. Acheulean lithics appear along with "Homo erectus", some 1.8 million years ago, replacing the more primitive Oldowan industry used by "A. garhi" and by the earliest species of "Homo". The Middle Paleolithic saw more varied speciation within "Homo", including the appearance of "Homo sapiens" about 200,000 years ago. According to mitochondrial timing techniques, modern humans migrated from Africa after the Riss glaciation in the Middle Palaeolithic during the Eemian Stage, spreading all over the ice-free world during the late Pleistocene. A 2005 study posits that humans in this migration interbred with archaic human