Camouflage the peacock flounder, squid and octopus actively change their skin patterns and colours using special chromatophore cells to resemble their current background, or, as in most chameleons, for signalling. However, Smith's dwarf chameleon does use active colour change for camouflage. Each chromatophore contains pigment of only one colour. In fish and frogs, colour change is mediated by the type of chromatophores known as melanophores that contain dark pigment. A melanophore is star-shaped; it contains many small pigmented organelles which can be dispersed throughout the cell, or aggregated near its centre. When the pigmented organelles are dispersed, the cell makes a
Military camouflage smaller shapes with dithering; this softens the edges of the large scale pattern, making the underlying objects harder to discern. Pixellated shapes pre-date computer aided design by many years, already being used in Soviet Union experiments with camouflage patterns, such as "TTsMKK" developed in 1944 or 1945. In the 1970s, US Army officer Timothy R. O'Neill suggested that patterns consisting of square blocks of colour would provide effective camouflage. By 2000, O'Neill's idea was combined with patterns like the German "Flecktarn" to create pixellated patterns such as CADPAT and MARPAT. Battledress in digital camouflage patterns was first designed by the