Chameleon turquoise, and purple. Chameleon skin has a superficial layer which contains pigments, and under the layer are cells with guanine crystals. Chameleons change colour by changing the space between the guanine crystals, which changes the wavelength of light reflected off the crystals which changes the colour of the skin. Colour change in chameleons has functions in camouflage, but most commonly in social signaling and in reactions to temperature and other conditions. The relative importance of these functions varies with the circumstances, as well as the species. Colour change signals a chameleon's physiological condition and intentions to other chameleons. Chameleons tend
Camouflage zoologist Hugh Cott's 1940 book "Adaptive Coloration in Animals" corrected Thayer's errors, sometimes sharply: "Thus we find Thayer straining the theory to a fantastic extreme in an endeavour to make it cover almost every type of coloration in the animal kingdom." Cott built on Thayer's discoveries, developing a comprehensive view of camouflage based on "maximum disruptive contrast", countershading and hundreds of examples. The book explained how disruptive camouflage worked, using streaks of boldly contrasting colour, paradoxically making objects less visible by breaking up their outlines. While Cott was more systematic and balanced in his view than Thayer, and did include