Comparison of the imperial and US customary measurement systems adopted a system based on the 231-cubic-inch wine gallon for all fluid purposes. This became the US fluid gallon. Both the imperial and US fluid gallon are divided into 4 quarts, 8 pints or 32 gills. However, whereas the US gill is divided into 4 US fluid ounces, the imperial gill is divided into 5 imperial fluid ounces. So whilst the imperial gallon, quart, pint and gill are about 20% larger than their US fluid measure counterparts, the fluid ounce is about 4% smaller. Note that one avoirdupois ounce of water has an approximate volume of one imperial fluid ounce
External gills salamander species, such as most members of the Proteidae family (the olm and mudpuppies) and the Sirenidae family, which naturally never metamorphose into an air-breathing form. The embryos of frogs and caecilians also develop external gills at some point in their development, though these are either resorbed before or disappear shortly after hatching. Fossils of the distant relatives of modern amphibians, such as "Branchiosaurus" and "Apateon", also show evidence of external gills. The external gills commonly consist of a single stalk (rami) protruding from a gill arch behind the head of the animal, above an associated gill slit. The stalk