Acids in wine ripened grapes. While concentration varies among grape varieties, it is usually found in higher levels with red wine grapes. The acid is created as a byproduct of the metabolization of nitrogen by yeast cells during fermentation. The combination of succinic acid with one molecule of ethanol will create the ester mono-ethyl succinate responsible for a mild, fruit aroma in wines. Acidity is highest in wine grapes just before the start of "veraison", which ushers in the ripening period of the annual cycle of grape vines. As the grapes ripen, their sugar levels increase and their acidity levels decrease. Through the
Wine Grapes and original work that is much more extensive in coverage, and also incorporates recent findings from ampelographic research, including DNA profiling of grape varieties. "Wine Grapes" include a catalog listing of 1,368 grape varieties from across the globe including 377 Italian, 204 French and 77 Portuguese wine grape varieties. Coverage includes a listing of synonyms as well as the genetic relationship between varieties derived from DNA analysis. According to Jancis Robinson and José Vouillamoz, "Wine Grapes" includes details about almost 300 previously unpublished relationship between different grape varieties including the extensive family tree of the Pinot grape that includes a