Amazon biome and cerrado (savannah) to the south, and includes lands to the northeast extending to the Atlantic ocean with similar vegetation to the Amazon basin. J. J. Morrone (2006) defines the Amazonian subregion in this broader sense, divided into the biogeographical provinces of Guyana, Humid Guyana, Napo, Imeri, Roraima, Amapá, Várzea, Ucayali, Madeira, Tapajós-Xingu, Pará, Yungas and Pantanal. The World Wildlife Fund takes a similar scope, where the Amazon biome includes the Guiana Shield rain forests in the north and the Chiquitano dry forests of Bolivia. The biome covers parts of Brazil, Bolivia, Peru, Ecuador, Colombia, Venezuela, Guyana, Suriname and French
Iwokrama Forest degrees west longitude. The Iwokrama Forest is bordered to the west by the Pakaraima Mountains and to the east by the isolated highlands scattered through central-east Guyana, Suriname and French Guiana. It is also bordered by savannahs in the southwest and northeast of Guyana and southwestern Suriname. The Essequibo River forms the eastern boundary. The northern boundary is the Siparuni River. The Burro-Burro River runs through the centre, and most of its watershed is within the Forest. The area is covered with lowland tropical forest, and dominated by tall tropical trees with a dense canopy 20 to 30 metres (70