Swallowtail butterfly tiger swallowtail), Y-linkage determines whether the females are either wild-type (yellow and black) or melanic (dark melanin replaces the yellow background). This genetic difference stems from the fact that melanism is controlled by a single gene, which controls the level of dopamine in the organism. The enzyme BAS, which assists dopamine in producing the yellow pigmentation, normally found on the wings' background, is suppressed. Without the pigmentation, the butterfly appears mostly black (the melanic form) and is a Batesian mimic of "Battus philenor", the pipevine swallowtail. There are also "Papilio glaucus" that are not wholly black; several possess an intermediate
Syntonic comma just 6:5. This result was obtained by narrowing each fifth by a quarter of a syntonic comma, an amount which was considered negligible, and permitted the full development of music with complex texture, such as polyphonic music, or melody with instrumental accompaniment. Since then, other tuning systems were developed, and the syntonic comma was used as a reference value to temper the perfect fifths in an entire family of them. Namely, in the family belonging to the syntonic temperament continuum, including meantone temperaments. The syntonic comma arises in "comma pump" (comma drift) sequences such as C G D A E