Animal locomotion many of these same principles may be applied to climbing without trees, such as on rock piles or mountains. The earliest known tetrapod with specializations that adapted it for climbing trees, was "Suminia", a synapsid of the late Permian, about 260 million years ago. Some invertebrate animals are exclusively arboreal in habitat, for example, the tree snail. "Brachiation" (from "brachium", Latin for "arm"), is a form of arboreal locomotion in which primates swing from tree limb to tree limb using only their arms. During brachiation, the body is alternately supported under each forelimb. This is the primary means of locomotion
Sexual dimorphism in non-human primates to the likelihood of higher vulnerability to predators. Another hypothesis suggests that arboreal primates have limitations on their upper body size, given that larger body size could disrupt their usage of terminal branches for locomotion. However, among some species of guenons ("Cercopithecus"), arboreal blue monkeys ("C. mitis") appear to be more sexually dimorphic than terrestrial vervet monkeys ("C. aethiops"). It has been hypothesized that niche divergence between the sexes attributes to the evolution of size dimorphism in primates. Males and females are known to have different preferences for ecological habitat due to different reproductive activities, which could possibly lead to