Swallowtail butterfly tiger swallowtail), Y-linkage determines whether the females are either wild-type (yellow and black) or melanic (dark melanin replaces the yellow background). This genetic difference stems from the fact that melanism is controlled by a single gene, which controls the level of dopamine in the organism. The enzyme BAS, which assists dopamine in producing the yellow pigmentation, normally found on the wings' background, is suppressed. Without the pigmentation, the butterfly appears mostly black (the melanic form) and is a Batesian mimic of "Battus philenor", the pipevine swallowtail. There are also "Papilio glaucus" that are not wholly black; several possess an intermediate
Syntonic comma Johnston uses a "−" as an accidental to indicate a note is lowered by a syntonic comma, or a "+" to indicate a note is raised by a syntonic comma. Thus a Pythagorean scale is C D E+ F G A+ B+, while the 5-limit Ptolemaic scale is C D E F G A B. Syntonic comma In music theory, the syntonic comma, also known as the chromatic diesis, the comma of Didymus, the Ptolemaic comma, or the diatonic comma is a small comma type interval between two musical notes, equal to the frequency ratio 81:80 (= 1.0125) (around 21.51