Swallowtail butterfly tiger swallowtail), Y-linkage determines whether the females are either wild-type (yellow and black) or melanic (dark melanin replaces the yellow background). This genetic difference stems from the fact that melanism is controlled by a single gene, which controls the level of dopamine in the organism. The enzyme BAS, which assists dopamine in producing the yellow pigmentation, normally found on the wings' background, is suppressed. Without the pigmentation, the butterfly appears mostly black (the melanic form) and is a Batesian mimic of "Battus philenor", the pipevine swallowtail. There are also "Papilio glaucus" that are not wholly black; several possess an intermediate
Holdrian comma fact that the Ancient Greeks (or at least Boethius) believed that in the Pythagorean tuning the tone could be divided in nine commas, four of which forming the diatonic semitone and five the chromatic semitone. If all these commas are exactly of the same size, there results an octave of 5 tones + 2 diatonic semitones, 5 × 9 + 2 × 4 = 53 equal commas. Holder attributes the division of the octave in 53 equal parts to Nicolas Mercator, who would have named the 1/53 part of the octave the "Artificial Comma". Mercator's comma is a name often