Animal locomotion many of these same principles may be applied to climbing without trees, such as on rock piles or mountains. The earliest known tetrapod with specializations that adapted it for climbing trees, was "Suminia", a synapsid of the late Permian, about 260 million years ago. Some invertebrate animals are exclusively arboreal in habitat, for example, the tree snail. "Brachiation" (from "brachium", Latin for "arm"), is a form of arboreal locomotion in which primates swing from tree limb to tree limb using only their arms. During brachiation, the body is alternately supported under each forelimb. This is the primary means of locomotion
Arboreal locomotion and chamois. Their adaptations may include a soft rubbery pad between their hooves for grip, hooves with sharp keratin rims for lodging in small footholds, and prominent dew claws. The snow leopard, being a predator of such mountain caprids, also has spectacular balance and leaping abilities; being able to leap up to ≈17m (~50 ft). Other balancers and leapers include the mountain zebra, mountain tapir, and hyraxes. Brachiation is a specialized form of arboreal locomotion, used by primates to move very rapidly while hanging beneath branches. Arguably the epitome of arboreal locomotion, it involves swinging with the arms from one